Aliases for TWSG1 Gene
Aliases for TWSG1 Gene
External Ids for TWSG1 Gene
- HGNC: 12429
- Entrez Gene: 57045
- Ensembl: ENSG00000128791
- OMIM: 605049
- UniProtKB: Q9GZX9
Previous GeneCards Identifiers for TWSG1 Gene
- GC00U990679
- GC18P009391
Summaries for TWSG1 Gene
GeneCards Summary for TWSG1 Gene
TWSG1 (Twisted Gastrulation BMP Signaling Modulator 1) is a Protein Coding gene. Diseases associated with TWSG1 include Holoprosencephaly. Among its related pathways are Mesodermal Commitment Pathway.
UniProtKB/Swiss-Prot for TWSG1 Gene
-
May be involved in dorsoventral axis formation. Seems to antagonize BMP signaling by forming ternary complexes with CHRD and BMPs, thereby preventing BMPs from binding to their receptors. In addition to the anti-BMP function, also has pro-BMP activity, partly mediated by cleavage and degradation of CHRD, which releases BMPs from ternary complexes. May be an important modulator of BMP-regulated cartilage development and chondrocyte differentiation. May play a role in thymocyte development (By similarity).
No data available for Entrez Gene Summary , CIViC summary , Tocris Summary , PharmGKB "VIP" Summary , fRNAdb sequence ontologies and piRNA Summary for TWSG1 Gene
Genomics for TWSG1 Gene
GeneHancer (GH) Regulatory Elements for TWSG1 Gene
Promoters and enhancers for TWSG1 Gene
- Elite GeneHancer and/or Elite GeneHancer-gene association
Download GeneHancer data dump
GeneHancers around TWSG1 on UCSC Golden Path with GeneCards custom track
Regulatory Element Products
Genomic Locations for TWSG1 Gene
Genomic Locations for TWSG1 Gene
- chr18:9,334,767-9,402,420
- (GRCh38/hg38)
- Size:
- 67,654 bases
- Orientation:
- Plus strand
- chr18:9,334,765-9,402,418
- (GRCh37/hg19)
Genomic View for TWSG1 Gene
Genes around TWSG1 on UCSC Golden Path with GeneCards custom track
- Cytogenetic band:
-
- 18p11.22 by Ensembl
- 18p11.22 by Entrez Gene
- 18p11.22 by HGNC
TWSG1 Gene in genomic location: bands according to
Ensembl, locations according to GeneLoc
(and/or Entrez Gene and/or Ensembl if different)
Genomic Neighborhood
• Exon Structure
• Gene Density
RefSeq DNA sequence for TWSG1 Gene
Proteins for TWSG1 Gene
-
Protein details for TWSG1 Gene (UniProtKB/Swiss-Prot)
- Protein Symbol:
- Q9GZX9-TWSG1_HUMAN
- Recommended name:
- Twisted gastrulation protein homolog 1
- Protein Accession:
- Q9GZX9
- B2RE08
- D3DUH9
- Q8NBI7
- Q96K46
Protein attributes for TWSG1 Gene
- Size:
- 223 amino acids
- Molecular mass:
- 25017 Da
- Quaternary structure:
-
- Interacts with CHRD and BMP4. This interaction enhances CHRD/BMP4 complex formation. Interacts with BMP7 (By similarity).
- SequenceCaution:
-
- Sequence=BAB55246.1; Type=Erroneous translation; Note=Wrong choice of CDS.; Evidence={ECO:0000305};
Protein Expression for TWSG1 Gene
Post-translational modifications for TWSG1 Gene
Other Protein References for TWSG1 Gene
- ENSEMBL proteins:
- REFSEQ proteins:
Antibody Products
- R&D Systems Antibodies for TWSG1 (TSG)
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-
Abcam antibodies for TWSG1
-
Cloud-Clone Corp. Antibodies for TWSG1
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Protein Products
- R&D Systems Proteins and Enzymes for TWSG1 (TSG)
-
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-
Cloud-Clone Corp. Proteins for TWSG1
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-
Abcam proteins for TWSG1
Assay Products
-
Cloud-Clone Corp. Assay Kits for TWSG1
- antibodies-online: Search results for 7 available TWSG1 Elisa Kits ranked by validation data
- Compare Top TWSG1 Elisa Kits
-
Quality Products:
No data available for DME Specific Peptides for TWSG1 Gene
Domains & Families for TWSG1 Gene
Gene Families for TWSG1 Gene
- Human Protein Atlas (HPA):
-
- Predicted secreted proteins
Suggested Antigen Peptide Sequences for TWSG1 Gene
- GenScript: Design optimal peptide antigens:
Graphical View of Domain Structure for InterPro Entry
Q9GZX9UniProtKB/Swiss-Prot:
TWSG1_HUMAN :- The N-terminal domain is sufficient to interact with BMP4.
- Belongs to the twisted gastrulation protein family.
- Domain:
-
- The N-terminal domain is sufficient to interact with BMP4.
- Family:
-
- Belongs to the twisted gastrulation protein family.
Function for TWSG1 Gene
Molecular function for TWSG1 Gene
- UniProtKB/Swiss-Prot Function:
- May be involved in dorsoventral axis formation. Seems to antagonize BMP signaling by forming ternary complexes with CHRD and BMPs, thereby preventing BMPs from binding to their receptors. In addition to the anti-BMP function, also has pro-BMP activity, partly mediated by cleavage and degradation of CHRD, which releases BMPs from ternary complexes. May be an important modulator of BMP-regulated cartilage development and chondrocyte differentiation. May play a role in thymocyte development (By similarity).
Phenotypes From GWAS Catalog for TWSG1 Gene
| GO ID | Qualified GO term | Evidence | PubMed IDs |
|---|---|---|---|
| GO:0005515 | protein binding | IPI | 25416956 |
| GO:0050431 | transforming growth factor beta binding | IDA | 17164348 |
Phenotypes for TWSG1 Gene
- MGI mutant phenotypes for TWSG1:
-
inferred from 6 alleles
- nervous system phenotype
- homeostasis/metabolism phenotype
- respiratory system phenotype
- cardiovascular system phenotype
- mortality/aging
- immune system phenotype
- cellular phenotype
- behavior/neurological phenotype
- growth/size/body region phenotype
- digestive/alimentary phenotype
- endocrine/exocrine gland phenotype
- vision/eye phenotype
- hearing/vestibular/ear phenotype
- embryo phenotype
- hematopoietic system phenotype
- skeleton phenotype
- renal/urinary system phenotype
- craniofacial phenotype
- limbs/digits/tail phenotype
- GenomeRNAi human phenotypes for TWSG1:
Animal Models for TWSG1 Gene
Animal Model Products
-
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-
-
ViGene Biosciences lentiviral particle packaged cDNA for TWSG1 gene
-
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- Search ViGene Biosciences for TWSG1
CRISPR Products
-
OriGene CRISPR knockouts for TWSG1
- genomics-online: gRNA clones - Search results for 101 available TWSG1 gene related products
- Overview of 101 available TWSG1 gene related products
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-
Vectors and viruses for KO, Activation, Repression, and more
-
Santa Cruz Biotechnology (SCBT) CRISPR for TWSG1
- GenScript: Design CRISPR guide RNA sequences for TWSG1
miRNA for TWSG1 Gene
- miRTarBase miRNAs that target TWSG1
-
- hsa-mir-124-3p (MIRT022445)
- hsa-mir-577 (MIRT040515)
- hsa-mir-142-3p (MIRT149046)
- hsa-mir-155-5p (MIRT149068)
- hsa-mir-4291 (MIRT649715)
- hsa-mir-24-3p (MIRT649716)
- hsa-mir-4284 (MIRT649717)
- hsa-mir-6778-3p (MIRT649718)
- hsa-mir-149-5p (MIRT649719)
- hsa-mir-106b-3p (MIRT649720)
- hsa-mir-6756-3p (MIRT649721)
- hsa-mir-3127-3p (MIRT649722)
- hsa-mir-545-3p (MIRT649723)
- hsa-mir-892b (MIRT649724)
- hsa-mir-3591-3p (MIRT664754)
- hsa-mir-21-3p (MIRT664755)
miRNA Products
- Search ViGene Biosciences for TWSG1
Inhibitory RNA Products
- Origene RNAi, shrna, and sirna products in human, mouse, rat for TWSG1
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- Overview of 101 available TWSG1 gene related products
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ViGene Biosciences ready-to-package AAV shRNAs for TWSG1 gene
Clone Products
- Sino Biological Human cDNA Clone for TWSG1
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-
VectorBuilder Other custom vectors
- Mammalian expression: PiggyBac
- Mammalian Tet-on expression: plasmid
- Mammalian conditional (Cre-Lox): plasmid and PiggyBac
- Mammalian shRNA knockdown: lentiviral, adenoviral, AAV, and PiggyBac
- CRISPR: plasmid gRNA, lentiviral gRNA, and donor plasmid
- Bacterial expression: pET, pBAD, and pCS
- Yeast expression
- Applied Biological Materials Clones for TWSG1
-
Vectors and viruses for ORF, Lenti, Retro, Adenovirus, AAV, and more
Cell Line Products
-
Horizon Cell Lines for TWSG1
-
ViGene Biosciences adenoviral particle packaged cDNA for TWSG1 gene
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-
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No data available for Enzyme Numbers (IUBMB) , Human Phenotype Ontology , Transcription Factor Targets and HOMER Transcription for TWSG1 Gene
Localization for TWSG1 Gene
Subcellular locations from UniProtKB/Swiss-Prot for TWSG1 Gene
- Secreted.
| Compartment | Confidence |
|---|---|
| extracellular | 5 |
| cytosol | 3 |
| cytoskeleton | 2 |
| plasma membrane | 1 |
| mitochondrion | 1 |
| nucleus | 1 |
| endoplasmic reticulum | 1 |
- Centrosome (3)
- Cytosol (3)
| GO ID | Qualified GO term | Evidence | PubMed IDs |
|---|---|---|---|
| GO:0005576 | extracellular region | IEA | -- |
| GO:0005615 | extracellular space | IEA | -- |
| GO:0070062 | extracellular exosome | HDA,IDA | 23376485 |
Pathways & Interactions for TWSG1 Gene
| SuperPathway | Contained pathways | ||
|---|---|---|---|
| 1 | Mesodermal Commitment Pathway | ||
Pathways by source for TWSG1 Gene
1 BioSystems pathway for TWSG1 Gene
Interacting Proteins for TWSG1 Gene
Selected Interacting proteins:
Q9GZX9-TWSG1_HUMAN
ENSP00000262120
for TWSG1 Gene via
IID
STRING
UniProtKB
| GO ID | Qualified GO term | Evidence | PubMed IDs |
|---|---|---|---|
| GO:0001503 | ossification | IEA | -- |
| GO:0001707 | mesoderm formation | IEA | -- |
| GO:0001818 | negative regulation of cytokine production | IDA | 17164348 |
| GO:0007179 | transforming growth factor beta receptor signaling pathway | IDA | 17164348 |
| GO:0007275 | multicellular organism development | IEA | -- |
No data available for SIGNOR curated interactions for TWSG1 Gene
Transcripts for TWSG1 Gene
mRNA/cDNA for TWSG1 Gene
- (1) REFSEQ mRNAs :
- (7) Additional mRNA sequences :
- (209) Selected AceView cDNA sequences:
- (3) Ensembl transcripts including schematic representations, and UCSC links where relevant :
Unigene Clusters for TWSG1 Gene
CRISPR Products
-
OriGene CRISPR knockouts for TWSG1
- genomics-online: gRNA clones - Search results for 101 available TWSG1 gene related products
- Overview of 101 available TWSG1 gene related products
- Applied Biological Materials CRISPR for TWSG1
-
Vectors and viruses for KO, Activation, Repression, and more
-
Santa Cruz Biotechnology (SCBT) CRISPR for TWSG1
- GenScript: Design CRISPR guide RNA sequences for TWSG1
miRNA Products
- Search ViGene Biosciences for TWSG1
Inhibitory RNA Products
- Origene RNAi, shrna, and sirna products in human, mouse, rat for TWSG1
- Browse OriGene Inhibitory RNA Products For TWSG1
- genomics-online: shRNA clones - Search results for 101 available TWSG1 gene related products
- Overview of 101 available TWSG1 gene related products
-
ViGene Biosciences ready-to-package AAV shRNAs for TWSG1 gene
Clone Products
- Sino Biological Human cDNA Clone for TWSG1
- VectorBuilder custom plasmid, inducible vectors for TWSG1
- VectorBuilder custom lentivirus, adenovirus, AAV vector/virus packaging for TWSG1
-
VectorBuilder Other custom vectors
- Mammalian expression: PiggyBac
- Mammalian Tet-on expression: plasmid
- Mammalian conditional (Cre-Lox): plasmid and PiggyBac
- Mammalian shRNA knockdown: lentiviral, adenoviral, AAV, and PiggyBac
- CRISPR: plasmid gRNA, lentiviral gRNA, and donor plasmid
- Bacterial expression: pET, pBAD, and pCS
- Yeast expression
- Applied Biological Materials Clones for TWSG1
-
Vectors and viruses for ORF, Lenti, Retro, Adenovirus, AAV, and more
| ExUns: | 1a | · | 1b | ^ | 2a | · | 2b | · | 2c | ^ | 3 | ^ | 4a | · | 4b | ^ | 5 | ^ | 6 | ^ | 7 | ^ | 8a | · | 8b | ^ | 9a | · | 9b |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| SP1: | - | - | - | ||||||||||||||||||||||||||
| SP2: | - | - | - | - | - | ||||||||||||||||||||||||
| SP3: | - | - | - | - | |||||||||||||||||||||||||
| SP4: | - | - | - | - | - | - | |||||||||||||||||||||||
| SP5: | |||||||||||||||||||||||||||||
| SP6: | - |
Expression for TWSG1 Gene
mRNA expression in embryonic tissues and stem cells from LifeMap Discovery
-
Skeletal Muscle (Muscoskeletal System)
- Muscle Progenitor Cells Mandibular Arch Muscles
- Muscle Progenitor Cells Hyoid Arch Muscles
- Limb (Muscoskeletal System)
-
Uterus (Reproductive System)
-
Fibroblasts (Uncategorized)
This gene is overexpressed in Urine (38.5), Serum (11.5), and Peripheral blood mononuclear cells (6.4).
Integrated Proteomics: protein expression in normal tissues and cell lines from ProteomicsDB, PaxDb, MaxQB, and MOPED for TWSG1 Gene
NURSA nuclear receptor signaling pathways regulating expression of TWSG1 Gene:
TWSG1SOURCE GeneReport for Unigene cluster for TWSG1 Gene:
Hs.514685Evidence on tissue expression from TISSUES for TWSG1 Gene
- Lung(4.6)
- Nervous system(3.7)
- Kidney(2.1)
- Heart(2)
- Liver(2)
Primer Products
-
OriGene qPCR primer pairs for TWSG1
-
OriGene qPCR primer pairs and template standards for TWSG1
- genomics-online: primer clones - Search results for 101 available TWSG1 gene related products
- Overview of 101 available TWSG1 gene related products
Gene Expression Assay Products
No data available for mRNA differential expression in normal tissues , Protein tissue co-expression partners , mRNA Expression by UniProt/SwissProt and Phenotype-based relationships between genes and organs from Gene ORGANizer for TWSG1 Gene
Orthologs for TWSG1 Gene
This gene was present in the common ancestor of animals.
| Organism | Taxonomy | Gene | Similarity | Type | Details |
|---|---|---|---|---|---|
| chimpanzee (Pan troglodytes) |
Mammalia | TWSG1 33 34 |
|
||
| oppossum (Monodelphis domestica) |
Mammalia | TWSG1 34 |
|
OneToOne | |
| platypus (Ornithorhynchus anatinus) |
Mammalia | TWSG1 34 |
|
OneToOne | |
| dog (Canis familiaris) |
Mammalia | TWSG1 33 34 |
|
||
| cow (Bos Taurus) |
Mammalia | TWSG1 33 34 |
|
||
| rat (Rattus norvegicus) |
Mammalia | Twsg1 33 |
|
||
| mouse (Mus musculus) |
Mammalia | Twsg1 33 16 34 |
|
||
| chicken (Gallus gallus) |
Aves | TWSG1 33 34 |
|
||
| lizard (Anolis carolinensis) |
Reptilia | TWSG1 34 |
|
OneToOne | |
| tropical clawed frog (Silurana tropicalis) |
Amphibia | twsg1 33 |
|
||
| Str.1743 33 |
|
||||
| African clawed frog (Xenopus laevis) |
Amphibia | Xl.778 33 |
|
||
| zebrafish (Danio rerio) |
Actinopterygii | twsg1a 33 34 |
|
||
| twsg1b 34 |
|
OneToMany | |||
| tsga 33 |
|
||||
| rainbow trout (Oncorhynchus mykiss) |
Actinopterygii | Omy.1024 33 |
|
||
| fruit fly (Drosophila melanogaster) |
Insecta | tsg 33 34 |
|
||
| srw 34 |
|
OneToMany | |||
| cv 34 |
|
OneToMany | |||
| sea squirt (Ciona savignyi) |
Ascidiacea | -- 34 |
|
OneToOne |
- Species where no ortholog for TWSG1 was found in the sources mined by GeneCards:
-
- A. gosspyii yeast (Ashbya gossypii)
- Actinobacteria (Mycobacterium tuberculosis)
- African malaria mosquito (Anopheles gambiae)
- Alicante grape (Vitis vinifera)
- alpha proteobacteria (Wolbachia pipientis)
- amoeba (Dictyostelium discoideum)
- Archea (Pyrococcus horikoshii)
- baker's yeast (Saccharomyces cerevisiae)
- barley (Hordeum vulgare)
- beta proteobacteria (Neisseria meningitidis)
- bread mold (Neurospora crassa)
- Chromalveolata (Phytophthora infestans)
- common water flea (Daphnia pulex)
- corn (Zea mays)
- E. coli (Escherichia coli)
- filamentous fungi (Aspergillus nidulans)
- Firmicute bacteria (Streptococcus pneumoniae)
- fission yeast (Schizosaccharomyces pombe)
- green algae (Chlamydomonas reinhardtii)
- honey bee (Apis mellifera)
- K. lactis yeast (Kluyveromyces lactis)
- loblloly pine (Pinus taeda)
- malaria parasite (Plasmodium falciparum)
- medicago trunc (Medicago Truncatula)
- moss (Physcomitrella patens)
- orangutan (Pongo pygmaeus)
- pig (Sus scrofa)
- rice (Oryza sativa)
- rice blast fungus (Magnaporthe grisea)
- schistosome parasite (Schistosoma mansoni)
- sea anemone (Nematostella vectensis)
- sea urchin (Strongylocentrotus purpuratus)
- sorghum (Sorghum bicolor)
- soybean (Glycine max)
- stem rust fungus (Puccinia graminis)
- sugarcane (Saccharum officinarum)
- thale cress (Arabidopsis thaliana)
- tomato (Lycopersicon esculentum)
- toxoplasmosis (Toxoplasma gondii)
- Trichoplax (Trichoplax adhaerens)
- wheat (Triticum aestivum)
- worm (Caenorhabditis elegans)
Paralogs for TWSG1 Gene
No data available for Paralogs for TWSG1 Gene
Variants for TWSG1 Gene
| SNP ID | Clin | Chr 18 pos | Variation | AA Info | Type |
|---|---|---|---|---|---|
| rs1000030508 | -- | 9,364,833(+) | T/C | intron_variant | |
| rs1000073747 | -- | 9,334,164(+) | C/A | upstream_transcript_variant | |
| rs1000098931 | -- | 9,396,531(+) | T/C | coding_sequence_variant, missense_variant | |
| rs1000135743 | -- | 9,375,793(+) | A/G | intron_variant | |
| rs1000155242 | -- | 9,369,793(+) | A/G | intron_variant |
| Variant ID | Type | Subtype | PubMed ID |
|---|---|---|---|
| dgv1546n106 | CNV | deletion | 24896259 |
| dgv55n6 | CNV | deletion | 16902084 |
| esv2716769 | CNV | deletion | 23290073 |
| esv3304810 | CNV | mobile element insertion | 20981092 |
| esv3310271 | CNV | mobile element insertion | 20981092 |
| esv3332314 | CNV | insertion | 20981092 |
| esv3555034 | CNV | deletion | 23714750 |
| esv7261 | CNV | loss | 19470904 |
| nsv1056407 | CNV | gain | 25217958 |
| nsv1069735 | CNV | deletion | 25765185 |
| nsv1133243 | CNV | tandem duplication | 24896259 |
| nsv1151888 | CNV | deletion | 26484159 |
| nsv1160506 | CNV | duplication | 26073780 |
| nsv507859 | OTHER | sequence alteration | 20534489 |
| nsv576447 | CNV | gain | 21841781 |
| nsv833585 | CNV | gain+loss | 17160897 |
| nsv954080 | CNV | deletion | 24416366 |
Residual Variation Intolerance Score:
36.1% of all genes are more intolerant (likely to be disease-causing)
Gene Damage Index Score:
1.26;
25.24% of all genes are more intolerant (likely to be disease-causing)
Additional Variant Information for TWSG1 Gene
SNP Genotyping and Copy Number Assay Products
No data available for Polymorphic Variants from UniProtKB/Swiss-Prot for TWSG1 Gene
Disorders for TWSG1 Gene
| Disorder | Aliases | PubMed IDs |
|---|---|---|
| holoprosencephaly |
|
|
Search TWSG1 in MalaCards
View complete list of genes associated with diseases
Additional Disease Information for TWSG1
- Human Genome Epidemiology Navigator
- (HuGE)
- ATLAS of Genetics and Cytogenetics in Oncology and Haematology
No data available for UniProtKB/Swiss-Prot and Genatlas for TWSG1 Gene
Publications for TWSG1 Gene
- Homologues of Twisted gastrulation are extracellular cofactors in antagonism of BMP signalling. (PMID: 11260715) Scott IC … Greenspan DS (Nature 2001) 2 3 4 58
- Evolutionary conservation, developmental expression, and genomic mapping of mammalian Twisted gastrulation. (PMID: 11420619) Graf D … Merkenschlager M (Mammalian genome : official journal of the International Mammalian Genome Society 2001) 3 4 22 58
- Signal sequence and keyword trap in silico for selection of full-length human cDNAs encoding secretion or membrane proteins from oligo-capped cDNA libraries. (PMID: 16303743) Otsuki T … Isogai T (DNA research : an international journal for rapid publication of reports on genes and genomes 2005) 3 4 58
- Complete sequencing and characterization of 21,243 full-length human cDNAs. (PMID: 14702039) Ota T … Sugano S (Nature genetics 2004) 3 4 58
- Architecture of the human interactome defines protein communities and disease networks. (PMID: 28514442) Huttlin EL … Harper JW (Nature 2017) 3 58
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- Mammalian expression: PiggyBac
- Mammalian Tet-on expression: plasmid
- Mammalian conditional (Cre-Lox): plasmid and PiggyBac
- Mammalian shRNA knockdown: lentiviral, adenoviral, AAV, and PiggyBac
- CRISPR: plasmid gRNA, lentiviral gRNA, and donor plasmid
- Bacterial expression: pET, pBAD, and pCS
- Yeast expression
- antibodies-online: Search results for 42 available TWSG1 Antibodies ranked by validation data
- Compare Top TWSG1 Antibodies
- Quality Products:
- antibodies-online: Search results for 7 available TWSG1 Elisa Kits ranked by validation data
- Compare Top TWSG1 Elisa Kits
- Quality Products:
- antibodies-online: Search results for 14 available TWSG1 Proteins ranked by validation data
- Compare Top TWSG1 Proteins
- GeneTex TWSG1 antibody for TWSG1
- GeneTex Proteins for TWSG1
- ViGene Biosciences adenoviral particle packaged cDNA for TWSG1 gene
- ViGene Biosciences lentiviral particle packaged cDNA for TWSG1 gene
- ViGene Biosciences ready-to-package AAV shRNAs for TWSG1 gene
- Search ViGene Biosciences for TWSG1
- Horizon Cell Lines for TWSG1
- genomics-online: cdna clones - Search results for 101 available TWSG1 gene related products
- Overview of 101 available TWSG1 gene related products
- orf clones - Search results for 101 available TWSG1 gene related products
- Overview of 101 available TWSG1 gene related products
- genomics-online: gRNA clones - Search results for 101 available TWSG1 gene related products
- Overview of 101 available TWSG1 gene related products
- genomics-online: primer clones - Search results for 101 available TWSG1 gene related products
- Overview of 101 available TWSG1 gene related products
- genomics-online: shRNA clones - Search results for 101 available TWSG1 gene related products
- Overview of 101 available TWSG1 gene related products
Sources for TWSG1 Gene
- (1) GeneCards
- (2) HGNC
- (3) EntrezGene
- (4) Swiss-Prot
- (5) Ensembl
- (6) OMIM
- (7) GeneLoc
- (8) Gene Wiki
- (9) UCSC
- (10) PhosphoSitePlus
- (11) GO
- (12) TrEMBL
- (13) InterPro
- (14) ProtoNet
- (15) Blocks
- (16) MGI
- (17) IUBMB
- (18) KEGG
- (19) MINT
- (20) STRING
- (21) IntAct
- (22) Novoseek
- (23) PharmGKB
- (24) DrugBank
- (25) HMDB
- (26) UniGene
- (27) AceView
- (28) ASD
- (29) ECgene
- (30) GeneAnnot
- (31) CGAP SAGE
- (32) SOURCE
- (33) HomoloGene
- (34) PanEnsembl
- (35) euGenes
- (36) SGD
- (37) FlyBase
- (38) WormBase
- (39) Pseudogene
- (40) DGV
- (41) dbSNP
- (42) GenAtlas
- (43) HGMD
- (44) GAD
- (45) BGMUT
- (46) HuGE
- (47) Atlas
- (48) Cell Signaling Technology
- (49) GenBank
- (50) H-invDB
- (51) HORDE
- (52) HUGE
- (53) IMGT
- (54) Leiden
- (55) miRBase
- (56) DME
- (57) OriGene
- (58) PubMed
- (59) R&D Systems
- (60) TGDB
- (61) Tocris
- (62) Abcam
- (63) Novus Biologicals
- (64) ProSpec
- (65) Sino Biological
- (66) GenScript
- (67) Qiagen
- (68) Cloud-Clone Corp.
- (69) OCA
- (70) Proteopedia
- (71) MOPED
- (72) neXtProt
- (73) Reactome
- (74) GeneGo (Thomson Reuters)
- (75) fRNAdb
- (76) DISEASES
- (77) SIMAP
- (78) GenomeRNAi
- (79) LifeMap
- (80) miRTarBase
- (81) MalaCards
- (82) Invitrogen
- (83) BitterDB
- (84) Vector BioLabs
- (85) ESI-BIO
- (86) RefSeq
- (87) BioSystems
- (88) MaxQB
- (89) IUPHAR
- (90) BioGPS
- (91) Illumina
- (92) COMPARTMENTS
- (93) HOMER
- (94) PaxDb
- (95) ApexBio
- (96) Addgene
- (97) antibodies-online
- (98) CYP
- (99) NONCODE
- (100) SwitchGear Genomics
- (101) TreeFam
- (102) PathCards
- (103) GeneReviews
- (104) GeneTex
- (105) Taconic Biosciences
- (106) GTEx
- (107) ProteomicsDB
- (108) SCBT
- (109) DGIdb
- (110) ClinicalTrials
- (111) FDA Approved Drugs
- (112) RVIS
- (113) SIGNOR
- (114) diseasecard
- (115) NIH Rare Diseases
- (116) Orphanet
- (117) UMLS
- (118) GTR
- (119) Disease Ontology
- (120) Genetics Home Reference
- (121) MeSH
- (122) MedlinePlus
- (123) CDC
- (124) NINDS
- (125) NCBI Bookshelf
- (126) ClinVar
- (127) Gene Damage Index
- (128) ViGene Biosciences
- (129) HPO
- (130) UDN
- (131) VISTA
- (132) FANTOM5
- (133) ENCODE
- (134) ProSci
- (135) Horizon
- (136) NURSA
- (137) IID
- (138) Cyagen
- (139) VectorBuilder
- (140) SNPedia
- (141) BRCA Exchange
- (142) St John's Lab
- (143) CIViC
- (144) ProteoGenix
- (145) dbSUPER
- (146) TISSUES
- (147) Gene ORGANizer
- (148) abm
- (149) CrownBio
- (150) Human Protein Atlas
- (151) GWAS Catalog
- (152) Monarch Initiative
- (153) DataMed
- (154) HumanCyc
- (155) genomics-online
- (156) UCNEbase
- (157) EPDnew
- (158) Applied Biosystems
