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Aliases for GCN1 Gene
Aliases for GCN1 Gene
- GCN1 Activator Of EIF2AK4 2 3 5
- GCN1 (General Control Of Amino-Acid Synthesis 1, Yeast)-Like 1 2 3
- General Control Of Amino-Acid Synthesis 1-Like Protein 1 3 4
- GCN1, EIF2 Alpha Kinase Activator Homolog 2 3
- GCN1 EIF-2-Alpha Kinase Activator Homolog 3 4
- EIF-2-Alpha Kinase Activator GCN1 3 4
- Translational Activator GCN1 3 4
- GCN1-Like Protein 1 3 4
External Ids for GCN1 Gene
- HGNC: 4199
- Entrez Gene: 10985
- Ensembl: ENSG00000089154
- OMIM: 605614
- UniProtKB: Q92616
Previous HGNC Symbols for GCN1 Gene
- GCN1L1
Summaries for GCN1 Gene
GeneCards Summary for GCN1 Gene
GCN1 (GCN1 Activator Of EIF2AK4) is a Protein Coding gene. Diseases associated with GCN1 include Ogden Syndrome and Myopathy, Centronuclear, 2. An important paralog of this gene is ECPAS.
UniProtKB/Swiss-Prot Summary for GCN1 Gene
-
Acts as a positive activator of the EIF2AK4/GCN2 protein kinase activity in response to amino acid starvation. Forms a complex with EIF2AK4/GCN2 on translating ribosomes; during this process, GCN1 seems to act as a chaperone to facilitate delivery of uncharged tRNAs that enter the A site of ribosomes to the tRNA-binding domain of EIF2AK4/GCN2, and hence stimulating EIF2AK4/GCN2 kinase activity. Participates in the repression of global protein synthesis and in gene-specific mRNA translation activation, such as the transcriptional activator ATF4, by promoting the EIF2AK4/GCN2-mediated phosphorylation of eukaryotic translation initiation factor 2 (eIF-2-alpha/EIF2S1) on 'Ser-52', and hence allowing ATF4-mediated reprogramming of amino acid biosynthetic gene expression to alleviate nutrient depletion.
Additional gene information for GCN1 Gene
- HGNC(4199)
- Entrez Gene(10985)
- Ensembl(ENSG00000089154)
- OMIM(605614)
- UniProtKB(Q92616)
- Open Targets Platform(ENSG00000089154)
- Search for GCN1 at DataMed
- Search for GCN1 at HumanCyc
No data available for Entrez Gene Summary , CIViC Summary , Tocris Summary , PharmGKB "VIP" Summary , Rfam classification and piRNA Summary for GCN1 Gene
Genomics for GCN1 Gene
GeneHancer (GH) Regulatory Elements Pubs
Promoters and enhancers for GCN1 Gene
- Elite GeneHancer and/or Elite GeneHancer-gene association
Download GeneHancer data from 2017 publication |
Request up-to-date GeneHancer data (full dataset)
GeneHancers around GCN1 on the GeneHancer Hub at the UCSC Golden Path
Cistromic (ChIP-Seq) regulation report from SPP (The Signaling Pathways Project) for GCN1
Genomic Locations for GCN1 Gene
Genomic Locations for GCN1 Gene
- chr12:120,127,202-120,194,715
- (GRCh38/hg38)
- Size:
- 67,514 bases
- Orientation:
- Minus strand
- chr12:120,565,007-120,632,513
- (GRCh37/hg19)
- Size:
- 67,507 bases
- Orientation:
- Minus strand
Genomic View for GCN1 Gene
Genes around GCN1 on UCSC Golden Path with GeneCards custom track
- Cytogenetic band:
-
- 12q24.23 by HGNC
- 12q24.23 by Entrez Gene
- 12q24.23 by Ensembl
GCN1 Gene in genomic location: bands according to
Ensembl, locations according to GeneLoc
(and/or Entrez Gene and/or Ensembl if different)
Genomic Neighborhood
• Exon Structure
• Gene Density
RefSeq DNA sequence for GCN1 Gene
Proteins for GCN1 Gene
-
Protein details for GCN1 Gene (UniProtKB/Swiss-Prot)
- Protein Symbol:
- Q92616-GCN1_HUMAN
- Recommended name:
- eIF-2-alpha kinase activator GCN1
- Protein Accession:
- Q92616
- A8KAY1
- O95001
- O95651
- Q6P2S3
- Q86X65
- Q8N5I5
- Q8WU80
- Q99736
- Q9UE60
Protein attributes for GCN1 Gene
- Size:
- 2671 amino acids
- Molecular mass:
- 292758 Da
- Quaternary structure:
-
- Interacts with EIF2AK4/GCN2; this interaction stimulates the EIF2AK4/GCN2 kinase activity and is impaired by IMPACT upon a variety of stress conditions, such as amino acid depletion, UV-C irradiation, proteasome inhibitor treatment and glucose deprivation. Interacts with IMPACT; this prevents the interaction of GCN1 with EIF2AK4/GCN2 and inhibits EIF2AK4/GCN2 kinase activity.
- SequenceCaution:
-
- Sequence=AAC51648.1; Type=Frameshift; Evidence={ECO:0000305}; Sequence=AAC83183.1; Type=Erroneous gene model prediction; Evidence={ECO:0000305}; Sequence=BAA13209.2; Type=Erroneous initiation; Note=Extended N-terminus.; Evidence={ECO:0000305};
Post-translational modifications for GCN1 Gene
Other Protein References for GCN1 Gene
- ENSEMBL proteins:
- REFSEQ proteins:
Antibody Products
- Novus Biologicals Antibodies for GCN1
- Invitrogen Antibodies for GCN1 (AFLGC-GCN1)
-
Biorbyt antibodies for GCN1
-
Boster Bio Antibodies for GCN1
Protein Products
- Search Origene for Purified Proteins, MassSpec and Protein Over-expression Lysates for GCN1
- Origene Custom Protein Services for GCN1
- Novus Biologicals proteins for GCN1
No data available for DME Specific Peptides for GCN1 Gene
Domains & Families for GCN1 Gene
Gene Families for GCN1 Gene
- HGNC:
- Human Protein Atlas (HPA):
-
- Plasma proteins
- Predicted intracellular proteins
Protein Domains for GCN1 Gene
- InterPro:
- Blocks:
-
- Armadillo repeat
- Bacterial regulatory protein LysR, HTH motif
- ProtoNet:
Suggested Antigen Peptide Sequences for GCN1 Gene
- GenScript: Design optimal peptide antigens:
-
- Peroxisome proliferator activated receptor interacting complex protein (E1NZA1_HUMAN)
- GCN1-like protein 1 (GCN1L_HUMAN)
Graphical View of Domain Structure for InterPro Entry
Q92616- Family:
-
- Belongs to the GCN1 family.
Function for GCN1 Gene
Molecular function for GCN1 Gene
- UniProtKB/Swiss-Prot Function:
- Acts as a positive activator of the EIF2AK4/GCN2 protein kinase activity in response to amino acid starvation. Forms a complex with EIF2AK4/GCN2 on translating ribosomes; during this process, GCN1 seems to act as a chaperone to facilitate delivery of uncharged tRNAs that enter the A site of ribosomes to the tRNA-binding domain of EIF2AK4/GCN2, and hence stimulating EIF2AK4/GCN2 kinase activity. Participates in the repression of global protein synthesis and in gene-specific mRNA translation activation, such as the transcriptional activator ATF4, by promoting the EIF2AK4/GCN2-mediated phosphorylation of eukaryotic translation initiation factor 2 (eIF-2-alpha/EIF2S1) on 'Ser-52', and hence allowing ATF4-mediated reprogramming of amino acid biosynthetic gene expression to alleviate nutrient depletion.
Phenotypes From GWAS Catalog for GCN1 Gene
| GO ID | Qualified GO term | Evidence | PubMed IDs |
|---|---|---|---|
| GO:0003723 | RNA binding | HDA | 22681889 |
| GO:0008135 | translation factor activity, RNA binding | NAS | 9234705 |
| GO:0019887 | protein kinase regulator activity | IEA | -- |
| GO:0019901 | protein kinase binding | IEA | -- |
| GO:0043022 | ribosome binding | IEA | -- |
Phenotypes for GCN1 Gene
Animal Model Products
- Taconic Biosciences: Generate A Custom CRISPR Mouse Model For Your Study
CRISPR Products
- Applied Biological Materials (abm): CRISPR Clones for GCN1 - Select products 50% OFF >
- * GCN1 CRISPR as ready-to-use vector or virus: ORF | Lenti- | Adeno- | AAV- | Protein Vector - Browse All
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-
OriGene CRISPR knockouts for GCN1
- Search GeneCopoeia for CRISPR/Cas9 clones, lentivirus and AAV products for GCN1
miRNA for GCN1 Gene
- miRTarBase miRNAs that target GCN1
-
- hsa-miR-1293 (MIRT035903)
- hsa-miR-1226-3p (MIRT036502)
- hsa-miR-18a-3p (MIRT040858)
- hsa-miR-484 (MIRT042010)
- hsa-miR-425-3p (MIRT042433)
- hsa-miR-423-3p (MIRT042534)
- hsa-miR-328-3p (MIRT043819)
- hsa-miR-185-5p (MIRT045354)
- hsa-miR-149-5p (MIRT045622)
- hsa-miR-222-3p (MIRT046637)
- hsa-miR-221-3p (MIRT046958)
- hsa-let-7b-5p (MIRT052324)
- hsa-let-7a-5p (MIRT052425)
miRNA Products
Inhibitory RNA Products
- Origene shrna, sirna, and RNAi products in human, mouse, rat for GCN1
- Browse OriGene Inhibitory RNA Products For GCN1
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Clone Products
- Applied Biological Materials (abm): Clones for GCN1 - Select products 50% OFF >
- * GCN1 as ready-to-use vector or virus: ORF | Lenti- | Retro- | Adeno- | AAV- | Protein Vector - Browse All
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- Search GeneCopoeia for ORF cDNA, lentivirus, AAV viral particles and/or promoter clone products for GCN1
No data available for Enzyme Numbers (IUBMB) , Human Phenotype Ontology , Animal Models , Transcription Factor Targets and HOMER Transcription for GCN1 Gene
Localization for GCN1 Gene
Subcellular locations from UniProtKB/Swiss-Prot for GCN1 Gene
- Cytoplasm. Note=Associates with ribosomes in undifferentiated neuroblastoma cells and increases after neuronal differentiation. {ECO:0000250 UniProtKB:E9PVA8}.
| Compartment | Confidence |
|---|---|
| cytosol | 5 |
| plasma membrane | 3 |
| extracellular | 1 |
| cytoskeleton | 1 |
| mitochondrion | 1 |
| peroxisome | 1 |
| nucleus | 1 |
| endoplasmic reticulum | 1 |
| endosome | 1 |
| golgi apparatus | 1 |
- Cytosol (4)
| GO ID | Qualified GO term | Evidence | PubMed IDs |
|---|---|---|---|
| GO:0005737 | cytoplasm | NAS | 9234705 |
| GO:0005829 | cytosol | IDA | -- |
| GO:0005840 | ribosome | NAS | 9234705 |
| GO:0005844 | polysome | IEA,ISS | -- |
| GO:0016020 | membrane | HDA | 19946888 |
Pathways & Interactions for GCN1 Gene
Interacting Proteins for GCN1 Gene
GPS-Prot Interaction Network for GCN1
| GO ID | Qualified GO term | Evidence | PubMed IDs |
|---|---|---|---|
| GO:0006412 | translation | IEA | -- |
| GO:0006417 | regulation of translation | IEA | -- |
| GO:0033674 | positive regulation of kinase activity | IEA | -- |
| GO:0034198 | cellular response to amino acid starvation | IEA,ISS | -- |
| GO:0036003 | positive regulation of transcription from RNA polymerase II promoter in response to stress | IEA,ISS | -- |
No data available for Pathways by source and SIGNOR curated interactions for GCN1 Gene
Transcripts for GCN1 Gene
mRNA/cDNA for GCN1 Gene
- 1 REFSEQ mRNAs :
- 16 NCBI additional mRNA sequence :
- 9 Ensembl transcripts including schematic representations, and UCSC links to gene/alias where relevant :
-
- ENST00000300648 8681 nts (view in UCSC)
- ENST00000547263 366 nts (view in UCSC)
- ENST00000547369 595 nts (view in UCSC)
- ENST00000548132 498 nts (view in UCSC)
- ENST00000548821 504 nts (view in UCSC)
CRISPR Products
- Applied Biological Materials (abm): CRISPR Clones for GCN1 - Select products 50% OFF >
- * GCN1 CRISPR as ready-to-use vector or virus: ORF | Lenti- | Adeno- | AAV- | Protein Vector - Browse All
- * Gene synthesis, site-directed mutagenesis, subcloning, and more services - Browse All
-
OriGene CRISPR knockouts for GCN1
- Search GeneCopoeia for CRISPR/Cas9 clones, lentivirus and AAV products for GCN1
miRNA Products
Inhibitory RNA Products
- Origene shrna, sirna, and RNAi products in human, mouse, rat for GCN1
- Browse OriGene Inhibitory RNA Products For GCN1
- Search GeneCopoeia for shRNA, lentivirus and/or AAV clone products for GCN1
Clone Products
- Applied Biological Materials (abm): Clones for GCN1 - Select products 50% OFF >
- * GCN1 as ready-to-use vector or virus: ORF | Lenti- | Retro- | Adeno- | AAV- | Protein Vector - Browse All
- * Gene synthesis, site-directed mutagenesis, subcloning, and more services - Browse All
- Search GeneCopoeia for ORF cDNA, lentivirus, AAV viral particles and/or promoter clone products for GCN1
Relevant External Links for GCN1 Gene
- GeneLoc Exon Structure for
- GCN1
Expression for GCN1 Gene
Transcriptomic regulation report from SPP (The Signaling Pathways Project) for GCN1
mRNA Expression by UniProt/SwissProt for GCN1 Gene:
Q92616-GCN1_HUMAN
Tissue specificity:
Ubiquitously expressed (PubMed:9039502). Expressed in skeletal muscules, ovary and testis (PubMed:9234705).
Evidence on tissue expression from TISSUES for GCN1 Gene
- Liver(4.9)
- Skin(4.6)
- Nervous system(4.5)
- Muscle(4.4)
- Adrenal gland(3.4)
- Heart(2.9)
- Eye(2.8)
- Pancreas(2.6)
- Kidney(2.1)
Primer Products
-
OriGene qPCR primer pairs for GCN1
Gene Expression Assay Products
No data available for mRNA expression in normal human tissues , mRNA expression in embryonic tissues and stem cells from LifeMap Discovery , mRNA differential expression in normal tissues , Protein differential expression in normal tissues , Protein expression , Protein tissue co-expression partners and Phenotype-based relationships between genes and organs from Gene ORGANizer for GCN1 Gene
Orthologs for GCN1 Gene
This gene was present in the common ancestor of eukaryotes.
| Organism | Taxonomy | Gene | Similarity | Type | Details |
|---|---|---|---|---|---|
| Chimpanzee (Pan troglodytes) |
Mammalia | GCN1L1 30 31 |
|
OneToOne | |
| Dog (Canis familiaris) |
Mammalia | GCN1L1 30 31 |
|
OneToOne | |
| Cow (Bos Taurus) |
Mammalia | GCN1L1 30 31 |
|
OneToOne | |
| Oppossum (Monodelphis domestica) |
Mammalia | GCN1L1 31 |
|
OneToOne | |
| Rat (Rattus norvegicus) |
Mammalia | Gcn1l1 30 |
|
||
| Mouse (Mus musculus) |
Mammalia | Gcn1l1 30 31 |
|
OneToOne | |
| Gcn1 17 |
|
||||
| Platypus (Ornithorhynchus anatinus) |
Mammalia | GCN1L1 31 |
|
OneToOne | |
| Chicken (Gallus gallus) |
Aves | GCN1L1 30 31 |
|
OneToOne | |
| Lizard (Anolis carolinensis) |
Reptilia | GCN1L1 31 |
|
OneToOne | |
| Tropical Clawed Frog (Silurana tropicalis) |
Amphibia | gcn1l1 30 |
|
||
| African clawed frog (Xenopus laevis) |
Amphibia | Xl.6112 30 |
|
||
| Zebrafish (Danio rerio) |
Actinopterygii | gcn1l1 30 31 |
|
OneToOne | |
| African malaria mosquito (Anopheles gambiae) |
Insecta | AgaP_AGAP001700 30 |
|
||
| Fruit Fly (Drosophila melanogaster) |
Insecta | CG17514 30 31 |
|
OneToOne | |
| Worm (Caenorhabditis elegans) |
Secernentea | gcn-1 30 31 |
|
OneToOne | |
| Baker's yeast (Saccharomyces cerevisiae) |
Saccharomycetes | GCN1 30 31 33 |
|
OneToOne | |
| K. Lactis Yeast (Kluyveromyces lactis) |
Saccharomycetes | KLLA0F12430g 30 |
|
||
| A. gosspyii yeast (Eremothecium gossypii) |
Saccharomycetes | AGOS_AFR169W 30 |
|
||
| Thale Cress (Arabidopsis thaliana) |
eudicotyledons | ILA 30 |
|
||
| Rice (Oryza sativa) |
Liliopsida | Os03g0721200 30 |
|
||
| Os.15153 30 |
|
||||
| Bread mold (Neurospora crassa) |
Ascomycetes | NCU05803 30 |
|
||
| Sea Squirt (Ciona savignyi) |
Ascidiacea | -- 31 |
|
OneToOne | |
| Fission Yeast (Schizosaccharomyces pombe) |
Schizosaccharomycetes | SPAC18G6.05c 30 |
|
- Species where no ortholog for GCN1 was found in the sources mined by GeneCards:
-
- Actinobacteria (Mycobacterium tuberculosis)
- Alicante grape (Vitis vinifera)
- Alpha proteobacteria (Wolbachia pipientis)
- Amoeba (Dictyostelium discoideum)
- Archea (Pyrococcus horikoshii)
- Barley (Hordeum vulgare)
- Beta proteobacteria (Neisseria meningitidis)
- Chromalveolata (Phytophthora infestans)
- Common water flea (Daphnia pulex)
- Corn (Zea mays)
- E. coli (Escherichia coli)
- Filamentous fungi (Aspergillus nidulans)
- Firmicute Bacteria (Streptococcus pneumoniae)
- Green Algae (Chlamydomonas reinhardtii)
- Honey Bee (Apis mellifera)
- Loblloly Pine (Pinus taeda)
- Malaria Parasite (Plasmodium falciparum)
- Medicago Trunc (Medicago Truncatula)
- Moss (Physcomitrella patens)
- Orangutan (Pongo pygmaeus)
- Pig (Sus scrofa)
- Rainbow Trout (Oncorhynchus mykiss)
- Rice Blast Fungus (Magnaporthe grisea)
- Schistosome Parasite (Schistosoma mansoni)
- Sea Anemone (Nematostella vectensis)
- Sea Vase (Ciona intestinalis)
- Sea Urchin (Strongylocentrotus purpuratus)
- Sorghum (Sorghum bicolor)
- Soybean (Glycine max)
- Stem Rust Fungus (Puccinia graminis)
- Sugarcane (Saccharum officinarum)
- Tomato (Lycopersicon esculentum)
- Toxoplasmosis (Toxoplasma gondii)
- Trichoplax (Trichoplax adhaerens)
- Wheat (Triticum aestivum)
Evolution for GCN1 Gene
- ENSEMBL:
- Gene Tree for GCN1 (if available)
- TreeFam:
- Gene Tree for GCN1 (if available)
- Aminode:
- Evolutionary constrained regions (ECRs) for GCN1: view image
Paralogs for GCN1 Gene
Variants for GCN1 Gene
Sequence variations, with clinical significance, from ClinVar and Humsavar, with links to dbSNP for GCN1 Gene
| SNP ID | Clinical significance and condition | Chr 12 pos | Variation | AA Info | Type |
|---|---|---|---|---|---|
| 692308 | Uncertain Significance: Fetal akinesia sequence; Arthrogryposis multiplex congenita | 120,155,290(-) | G/T | MISSENSE_VARIANT | |
| 710178 | Benign: not provided | 120,178,643(-) | G/A | SYNONYMOUS_VARIANT | |
| 712125 | Benign: not provided | 120,137,596(-) | G/A | SYNONYMOUS_VARIANT | |
| 713633 | Benign: not provided | 120,178,894(-) | G/A | SYNONYMOUS_VARIANT | |
| 713995 | Likely Benign: not provided | 120,173,684(-) | G/A | SYNONYMOUS_VARIANT |
Residual Variation Intolerance Score:
0.271% of all genes are more intolerant (likely to be disease-causing)
Gene Damage Index Score:
3.74;
57.73% of all genes are more intolerant (likely to be disease-causing)
Additional Variant Information for GCN1 Gene
- SNPedia medical, phenotypic, and genealogical associations of SNPs for
- GCN1
SNP Genotyping and Copy Number Assay Products
No data available for Polymorphic Variants from UniProtKB/Swiss-Prot for GCN1 Gene
Disorders for GCN1 Gene
| Disorder | Aliases | PubMed IDs |
|---|---|---|
| ogden syndrome |
|
|
| myopathy, centronuclear, 2 |
|
|
| myopathy, centronuclear, 1 |
|
|
| hemophagocytic lymphohistiocytosis, familial, 3 |
|
|
| neuroblastoma |
|
|
Search GCN1 in MalaCards
View complete list of genes associated with diseases
Additional Disease Information for GCN1
No data available for UniProtKB/Swiss-Prot and Genatlas for GCN1 Gene
Publications for GCN1 Gene
- Evidence that GCN1 and GCN20, translational regulators of GCN4, function on elongating ribosomes in activation of eIF2alpha kinase GCN2. (PMID: 9234705) Marton MJ … Hinnebusch AG (Molecular and cellular biology 1997) 2 3 4 23
- Perturbations in actin dynamics reconfigure protein complexes that modulate GCN2 activity and promote an eIF2 response. (PMID: 27852836) Silva RC … Castilho BA (Journal of cell science 2016) 2 3
- The finished DNA sequence of human chromosome 12. (PMID: 16541075) Scherer SE … Baylor College of Medicine Human Genome Sequencing Center Sequence Production Team (Nature 2006) 3 4
- The status, quality, and expansion of the NIH full-length cDNA project: the Mammalian Gene Collection (MGC). (PMID: 15489334) Gerhard DS … MGC Project Team (Genome research 2004) 3 4
- Construction of expression-ready cDNA clones for KIAA genes: manual curation of 330 KIAA cDNA clones. (PMID: 12168954) Nakajima D … Nagase T (DNA research : an international journal for rapid publication of reports on genes and genomes 2002) 3 4
ExternalLinks
Products for GCN1 Gene
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- Novus Biologicals proteins for GCN1
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- Invitrogen Antibodies for GCN1 (AFLGC-GCN1)
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- VectorBuilder Stable cell line generation for GCN1
- antibodies-online: Show 28 available GCN1L1 Antibodies ranked by validation data
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- antibodies-online: Search results for available GCN1L1 related products ranked by validation data
- antibodies-online: Search results for available GCN1L1 related products ranked by validation data
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- Boster Bio Antibodies for GCN1
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- Biorbyt antibodies for GCN1
Sources for GCN1 Gene
- (1) GeneCards
- (2) HGNC
- (3) NCBI Entrez Gene
- (4) UniProtKB/Swiss-Prot
- (5) Ensembl
- (6) OMIM
- (7) GeneLoc
- (8) Gene Wiki
- (9) UCSC
- (10) PhosphoSitePlus
- (11) Cell Signaling Technology
- (12) GO
- (13) UniProtKB/TrEMBL
- (14) InterPro
- (15) ProtoNet
- (16) Blocks
- (17) MGI
- (18) IUBMB
- (19) KEGG
- (20) MINT
- (21) STRING
- (22) IntAct
- (23) Novoseek
- (24) PharmGKB
- (25) DrugBank
- (26) HMDB
- (27) ASD
- (28) SAGE
- (29) SOURCE
- (30) HomoloGene
- (31) PanEnsembl
- (32) euGenes
- (33) SGD
- (34) FlyBase
- (35) WormBase
- (36) Pseudogene
- (37) DGV
- (38) dbSNP
- (39) GenAtlas
- (40) HGMD
- (41) GAD
- (42) BGMUT
- (43) HuGE
- (44) Atlas
- (45) GenBank
- (46) HORDE
- (47) HUGE
- (48) IMGT
- (49) Leiden
- (50) miRBase
- (51) DME
- (52) OriGene
- (53) PubMed
- (54) R&D Systems
- (55) TGDB
- (56) Tocris
- (57) Abcam
- (58) Novus Biologicals
- (59) ProSpec
- (60) Sino Biological
- (61) GenScript
- (62) Qiagen
- (63) OCA
- (64) Proteopedia
- (65) MOPED
- (66) neXtProt
- (67) Reactome
- (68) GeneGo (Thomson Reuters)
- (69) DISEASES
- (70) SIMAP
- (71) GenomeRNAi
- (72) LifeMap
- (73) miRTarBase
- (74) MalaCards
- (75) Invitrogen
- (76) BitterDB
- (77) ESI-BIO
- (78) RefSeq
- (79) BioSystems
- (80) MaxQB
- (81) IUPHAR
- (82) BioGPS
- (83) Illumina
- (84) COMPARTMENTS
- (85) HOMER
- (86) PaxDb
- (87) ApexBio
- (88) Addgene
- (89) antibodies-online
- (90) CYP
- (91) NONCODE
- (92) TreeFam
- (93) PathCards
- (94) GeneReviews
- (95) GeneTex
- (96) Taconic Biosciences
- (97) GTEx
- (98) ProteomicsDB
- (99) SCBT
- (100) DGIdb
- (101) ClinicalTrials
- (102) RVIS
- (103) SIGNOR
- (104) diseasecard
- (105) NIH Rare Diseases
- (106) Orphanet
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