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Category   Symbol Source: HGNC EntrezGene Ensembl GeneCards RNA genes CroW21
GIFtS

ZSCAN4 Gene

protein-coding   GIFtS: 46
GCID: GC19P058180

Zinc Finger And SCAN Domain Containing 4

(Previous name: zinc finger protein 494)
(Previous symbol: ZNF494)
Microbiology & Infectious Diseases Congress
  Search for ZSCAN4
at  

(According to 1HGNC, 2Entrez Gene,
3UniProtKB/Swiss-Prot, 4UniProtKB/TrEMBL, 5OMIM, 6GeneLoc, 7Ensembl, 8DME, 9miRBase, 10fRNAdb, 12H-InvDB, 13NCBI, 14NONCODE, and/or 15RNAdb)
About This Section

Aliases
Zinc Finger And SCAN Domain Containing 41 2
ZNF4941 2 3 5
Zinc Finger Protein 4941 2 3
Zinc Finger And SCAN Domain-Containing Protein 42

External Ids:    HGNC: 237091   Entrez Gene: 2015162   Ensembl: ENSG000001805327   OMIM: 6134195   UniProtKB: Q8NAM63   

Export aliases for ZSCAN4 gene to outside databases

Previous GC identifers: GC19P062874 GC19P062875 GC19P054492


(According to Entrez Gene, Tocris Bioscience, Wikipedia's Gene Wiki, PharmGKB,
UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL)
About This Section

Entrez Gene summary for ZSCAN4 Gene:
The ZSCAN4 gene encodes a protein involved in telomere maintenance and with a key role in the critical feature of
mouse embryonic stem (ES) cells, namely, defying cellular senescence and maintaining normal karyotype for many
cell divisions in culture (Zalzman et al., 2010 (PubMed 20336070)).(supplied by OMIM, May 2010)

GeneCards Summary for ZSCAN4 Gene: 
ZSCAN4 (zinc finger and SCAN domain containing 4) is a protein-coding gene. GO annotations related to this gene include DNA binding and sequence-specific DNA binding transcription factor activity.

UniProtKB/Swiss-Prot: ZSCA4_HUMAN, Q8NAM6
Function: Embryonic stem (ES) cell-specific transcription factor required to regulate ES cell pluripotency. Binds
telomeres and plays a key role in genomic stability in ES cells by regulating telomere elongation. Acts as an
activator of spontaneous telomere sister chromatid exchange (T-SCE) and telomere elongation in undifferentiated
ES cells (By similarity)




(According to GeneLoc and/or HGNC, and/or
Entrez Gene (NCBI build 37),
and/or miRBase,
Genomic Views according to UCSC (hg19) and Ensembl (release 73), Regulatory elements and Epigenetics data according to QIAGEN, SABiosciences, and/or SwitchGear Genomics)
About This Section
RefSeq DNA sequence:
NC_000019.9  NC_018930.2  NT_011109.16  
Regulatory elements:
   SABiosciences Regulatory transcription factor binding sites in the ZSCAN4 gene promoter:
         SRF   Elk-1   FOXF2   SRF (504 AA)   POU3F2 (N-Oct-5a)   POU3F2 (N-Oct-5b)   POU3F2   FOXC1   FOXO1a   FOXO1   
         Other transcription factors

SwitchGear Promoter luciferase reporter plasmidZSCAN4 promoter sequence
   Search SABiosciences Chromatin IP Primers for ZSCAN4

Epigenetics:
QIAGEN PyroMark CpG Assay predesigned Pyrosequencing DNA Methylation assays in human, mouse, rat ZSCAN4


Genomic Location:
Genomic View: UCSC Golden Path with GeneCards custom track

Entrez Gene cytogenetic band: 19q13.43   Ensembl cytogenetic band:  19q13.43   HGNC cytogenetic band: 19q13.43

ZSCAN4 Gene in genomic location: bands according to Ensembl, locations according to (and/or Entrez Gene and/or Ensembl if different)
ZSCAN4 gene location

GeneLoc information about chromosome 19         GeneLoc Exon Structure

GeneLoc location for GC19P058180:  view genomic region     (about GC identifiers)

Start:
58,180,303 bp from pter      End:
58,190,520 bp from pter
Size:
10,218 bases      Orientation:
plus strand

(According to UniProtKB, HORDE, neXtProt, Ensembl, and/or Reactome, Modification sites according to PhosphoSitePlus, Specific Peptides from DME, Protein expression images according to data from SPIRE 1MOPED, 2PaxDb, and 3MAXQB RefSeq according to NCBI, PDB rendering according to OCA and/or Proteopedia, Recombinant Proteins from EMD Millipore, R&D Systems, GenScript, Enzo Life Sciences, OriGene, Novus Biologicals, Sino Biological, ProSpec, and/or Cloud-Clone Corp.,
Biochemical Assays by EMD Millipore, R&D Systems, OriGene, GenScript, Cell Signaling Technology, Enzo Life Sciences, and/or Cloud-Clone Corp., Ontologies according to Gene Ontology Consortium 01 Oct 2013 and Entrez Gene, Antibodies by EMD Millipore, R&D Systems, GenScript, Cell Signaling Technology, OriGene, Novus Biologicals, Thermo Fisher Scientific, LSBio, Abcam, and/or Cloud-Clone Corp.)
About This Section

UniProtKB/Swiss-Prot: ZSCA4_HUMAN, Q8NAM6 (See protein sequence)
Recommended Name: Zinc finger and SCAN domain-containing protein 4  
Size: 433 amino acids; 48957 Da
Subcellular location: Nucleus (By similarity). Chromosome, telomere (By similarity)
Secondary accessions: Q3MIQ2

Explore the universe of human proteins at neXtProt for ZSCAN4: NX_Q8NAM6

Explore proteomics data for ZSCAN4 at MOPED 

Post-translational modifications:

  • View modification sites using PhosphoSitePlus
  • View neXtProt modification sites for NX_Q8NAM6

  • ZSCAN4 Protein expression data from MOPED1, PaxDb2 and MAXQB3 :    About this image 

    ZSCAN4 Protein Expression
    REFSEQ proteins: NP_689890.1  
    ENSEMBL proteins: 
     ENSP00000321963  

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    Gene Ontology (GO): 2 cellular component terms (GO ID links to tree view):    About this table

    GO IDQualified GO termEvidencePubMed IDs
    GO:0000784nuclear chromosome, telomeric region ISS--
    GO:0005634nucleus ----

    ZSCAN4 for ontologies           About GeneDecksing



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    (According to HGNC, IUPHAR, InterPro, ProtoNet, UniProtKB, and/or BLOCKS, Sets of similar genes according to GeneDecks)
    About This Section
    HGNC Gene Families: 
    ZNF: Zinc fingers, C2H2-type

    5 InterPro protein domains:
     IPR015880 Znf_C2H2-like
     IPR008916 Retrov_capsid_C
     IPR013087 Znf_C2H2/integrase_DNA-bd
     IPR007087 Znf_C2H2
     IPR003309 Tscrpt_reg_SCAN

    Graphical View of Domain Structure for InterPro Entry Q8NAM6

    ProtoNet protein and cluster: Q8NAM6

    1 Blocks protein domain: IPB003309 Transcriptional regulator SCAN

    UniProtKB/Swiss-Prot: ZSCA4_HUMAN, Q8NAM6
    Similarity: Contains 4 C2H2-type zinc fingers
    Similarity: Contains 1 SCAN box domain


    ZSCAN4 for domains           About GeneDecksing


    (According to 1UniProtKB, Genatlas, LifeMap Discovery™, IUBMB, and/or 2DME, Human phenotypes from GenomeRNAi, Animal models from MGI Mar 06 2013, inGenious Targeting Laboratory, genOway,
    bound targets from SABiosciences, miRNA Gene Targets from miRTarBase, shRNA from OriGene, RNAi from EMD Millipore, siRNAs from OriGene, QIAGEN, microRNA from QIAGEN, Gene Editing from DNA2.0, Sirion Biotech, Clones from EMD Millipore, OriGene, SwitchGear Genomics, GenScript, Sino Biological, DNA2.0, Vector BioLabs, and Sirion Biotech, Cell Lines from GenScript, LifeMap BioReagents, In Situ Hybridization Assays from Advanced Cell Diagnostics, Ontologies according to Gene Ontology Consortium 01 Oct 2013 via Entrez Gene.)
    About This Section

    Molecular Function:

         UniProtKB/Swiss-Prot Summary: ZSCA4_HUMAN, Q8NAM6
    Function: Embryonic stem (ES) cell-specific transcription factor required to regulate ES cell pluripotency. Binds
    telomeres and plays a key role in genomic stability in ES cells by regulating telomere elongation. Acts as an
    activator of spontaneous telomere sister chromatid exchange (T-SCE) and telomere elongation in undifferentiated
    ES cells (By similarity)

         Gene Ontology (GO): 4 molecular function terms (GO ID links to tree view):    About this table

    GO IDQualified GO termEvidencePubMed IDs
    GO:0003677DNA binding IEA--
    GO:0003700sequence-specific DNA binding transcription factor activity IEA--
    GO:0008270zinc ion binding ----
    GO:0046872metal ion binding IEA--
         
    ZSCAN4 for ontologies           About GeneDecksing


    Animal Models:
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    (Pathways according to EMD Millipore, R&D Systems, Cell Signaling Technology, KEGG, PharmGKB, BioSystems, Sino Biological, Reactome, Tocris Bioscience, GeneGo (Thomson Reuters), QIAGEN, and/or UniProtKB, Sets of similar genes according to GeneDecks, Interaction Networks according to SABiosciences, and/or STRING, Interactions according to 1UniProtKB, 2MINT, 3I2D, and/or 4STRING, with links to IntAct and Ensembl, Ontologies according to Gene Ontology Consortium 01 Oct 2013 via Entrez Gene).
    About This Section



    Interactions:

        Search SABiosciences Gene Network CentralTM Interacting Genes and Proteins Networks for ZSCAN4

    2 Interacting proteins for ZSCAN4 (Q8NAM62, 3) via UniProtKB, MINT, STRING, and/or I2D
    InteractantInteraction Details
    GeneCardExternal ID(s)
    SMAD1Q157972, 3MINT-62000 I2D: score=3 
    SMAD9O151982, 3MINT-62085 I2D: score=3 
    About this table

    Gene Ontology (GO): 4 biological process terms (GO ID links to tree view):    About this table

    GO IDQualified GO termEvidencePubMed IDs
    GO:0006351transcription, DNA-dependent IEA--
    GO:0006355regulation of transcription, DNA-dependent ----
    GO:0010833telomere maintenance via telomere lengthening ISS--
    GO:0016032viral process ----

    ZSCAN4 for ontologies           About GeneDecksing



    (Chemical Compounds according to UniProtKB, Enzo Life Sciences, EMD Millipore, Tocris Bioscience HMDB, BitterDB, and/or Novoseek, Ligands according to IUPHAR, and Drugs according to DrugBank, Enzo Life Sciences, and/or PharmGKB, with drugs/clinical trials/news search links to CenterWatch)
    About This Section
    Browse Small Molecules at EMD Millipore
    Browse drugs & compounds from Enzo Life Sciences

    Browse Tocris compounds for ZSCAN4 (ZSCA4)

    Search CenterWatch for drugs/clinical trials and news about ZSCAN4 / ZSCA4

    (Secondary structures according to fRNAdb,
    GenBank/EMBL/DDBJ Accessions according to
    Unigene (Build 236 Homo sapiens; Apr 25 2013) or GenBank,
    RefSeq according to Entrez Gene,
    DOTS (version 10), and/or AceView, transcript ids from Ensembl with links to UCSC,
    Conferences by KenesGroup, exon structure from GeneLoc, alternative splicing isoforms according to ASD and/or ECgene,
    RNAi Products from EMD Millipore,
    siRNAs from OriGene, QIAGEN, shRNA from OriGene, microRNA from QIAGEN,
    Tagged/untagged cDNA clones from OriGene, SwitchGear Genomics, GenScript, DNA2.0, Vector BioLabs, Sirion Biotech, Primers from OriGene, SABiosciences, and/or QIAGEN )
    About This Section

    REFSEQ mRNAs for ZSCAN4 gene: 
    NM_152677.2  

    Unigene Cluster for ZSCAN4:

    Zinc finger and SCAN domain containing 4
    Hs.469663  [show with all ESTs]
    Unigene Representative Sequence: NM_152677
    1 Ensembl transcript including schematic representation, and UCSC links where relevant:
    ENST00000318203(uc002qpu.3)
    Congresses - knowledge worth sharing:  
    European Congress of Clinical Microbiology and Infectious Diseases (ECCMID) 10 - 13 May 2014

    miRNA
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    Search QIAGEN for miScript miRNA Assays for microRNAs that regulate ZSCAN4
    SwitchGear 3'UTR luciferase reporter plasmidZSCAN4 3' UTR sequence
    Inhib. RNA
    Products:
         
    Browse for Gene Knock-down Tools from EMD Millipore
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      QIAGEN QuantiTect SYBR Green Assays in human, mouse, rat ZSCAN4
      QIAGEN QuantiFast Probe-based Assays in human, mouse, rat ZSCAN4

    Additional mRNA sequence: 

    AK092424.1 BC101738.1 BC101740.1 

    1 DOTS entry:

    DT.75129322 

    6 AceView cDNA sequences:

    AK092424 AX747538 NM_152677 AI138628 BX118882 BG572132 

    GeneLoc Exon Structure


    (RNA expression data according to H-InvDB, NONCODE, miRBase, and RNAdb, Expression images according to data from BioGPS, Illumina Human BodyMap, and CGAP SAGE, Sets of similar genes according to GeneDecks, in vivo and in vitro expression data from LifeMap Discovery™, plus additional links to Genevestigator, and/or SOURCE, and/or BioGPS, and/or UniProtKB,
    PCR Arrays from SABiosciences, Primers from OriGene, SABiosciences, and/or QIAGEN, In Situ Hybridization Assays from Advanced Cell Diagnostics)
    About This Section

    ZSCAN4 expression in normal human tissues (normalized intensities)
    See probesets specificity/sensitivity at GeneAnnot
    About this imageBioGPS <intensity>2/3
    CGAP TAG: GAATTTGTCA
    ZSCAN4 Expression
    About this image


    See ZSCAN4 Protein Expression from SPIRE MOPED and PaxDB
    Genevestigator expression for ZSCAN4

    SOURCE GeneReport for Unigene cluster: Hs.469663
        SABiosciences Custom PCR Arrays for ZSCAN4
    Primer
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    In Situ
    Assay Products:
     

     
    Advanced Cell Diagnostics RNAscope RNA in situ hybridization assays for ZSCAN4

    (Orthologs according to 1,2HomoloGene (2older version, for species not in 1newer version), 3euGenes, 4SGD , 5MGI Mar 06 2013, with possible further links to Flybase and/or WormBase, and/or 6Ensembl pan taxonomic compara , Gene Trees according to Ensembl and TreeFam)
    About This Section
      --

    ENSEMBL Gene Tree for ZSCAN4 (if available)
    TreeFam Gene Tree for ZSCAN4 (if available)

    (Paralogs according to 1HomoloGene,
    2Ensembl, and 3SIMAP, Pseudogenes according to 4Pseudogene.org Build 68)
    About This Section
    Paralogs for ZSCAN4 gene
    9 SIMAP similar genes for ZSCAN4 using alignment to 1 protein entry:     ZSCA4_HUMAN:
    ZNF    ZNF501    ZNF664    ZNF697    ZNF589    ZNF702P
    ZNF32    ZNF844    ZNF323

    ZSCAN4 for paralogs           About GeneDecksing



    (SNPs/Variants according to the 1NCBI SNP Database, 2Ensembl, 3PupaSUITE, 4UniProtKB, and DNA2.0, Linkage Disequilibrium by HapMap, Structural Variations(CNVs/InDels/Inversions) from the Database of Genomic Variants, Mutations from the Human Gene Mutation Database (HGMD) and the Locus Specific Mutation Databases (LSDB), Blood group antigen gene mutations by BGMUT, Resequencing Primers from QIAGEN, Cancer Mutation PCR Arrays and Assays and Copy Number PCR Arrays from SABiosciences)
    About This Section

    10/339 SNPs in ZSCAN4 are shown (see all 339)    About this table     
    Genomic DataTranscription Related DataAllele Frequencies
    SNP IDValidClinical
    significance
    Chr 19 posSequence#AA
    Chg
    TypeMore#Allele
    freq
    PopTotal
    sample
    More
    ----------
    rs730645801,2
    C--58178398(+) CAAGTT/CGAGAT 1 -- us2k14Minor allele frequency- C:0.17NA WA EA 360
    rs129835961,2
    C,F,A,H--58178505(+) ATCCTT/CATTTG 1 -- us2k121Minor allele frequency- C:0.28NS EA NA CSA WA 2348
    rs596506521,2
    C,F--58178576(+) CCTAAG/CTTGGG 1 -- us2k14Minor allele frequency- C:0.16WA CSA 124
    rs1874615531,2
    --58178594(+) CAGGTA/CATTTC 1 -- us2k10--------
    rs562044061,2
    C,F--58178854(+) ATTTAG/AGATCT 1 -- us2k15Minor allele frequency- A:0.17NA CSA WA EA 362
    rs1468886881,2
    --58179023(+) AGTTAC/TACACC 1 -- us2k10--------
    rs1919050401,2
    --58179122(+) TCCTTA/GCAGCT 1 -- us2k10--------
    rs1396708111,2
    C--58179143(+) CTCAAG/TGAACT 1 -- us2k10--------
    rs104038511,2
    C,F,H--58179234(+) TCTACA/GTTTTT 1 -- us2k128Minor allele frequency- G:0.50NS EA NA WA CSA 2776
    rs129764391,2
    C--58179251(+) TTTTTG/TTTTGT 1 -- us2k1 tfbs3 trp34Minor allele frequency- T:0.00NA 8

    HapMap Linkage Disequilibrium report for ZSCAN4 (58180303 - 58190520 bp)

    Structural Variations
         Database of Genomic Variants (DGV) 2 variations for ZSCAN4:    About this table     
    Variant IDTypeSubtypePubMed ID
    dgv4028n71CNV Loss21882294
    nsv912528CNV Gain21882294

    SABiosciences Cancer Mutation PCR Assays
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    (in which this Gene is Involved, According to MalaCards, OMIM, UniProtKB, the University of Copenhagen DISEASES database, Conferences by KenesGroup, Genatlas, GeneTests, GAD, HuGE Navigator, and/or TGDB.)
    About This Section

    ZSCAN4 for disorders           About MalaCards
    OMIM gene information: 613419    OMIM disorders: --


    ZSCAN4 for disorders           About GeneDecksing


    Export disorders for ZSCAN4 gene to outside databases

    (in PubMed. Associations of this gene to articles via 1Entrez Gene, 2UniProtKB/Swiss-Prot, 3HGNC, 4GAD, 5PharmGKB, 6HMDB, 7DrugBank, 8UniProtKB/TrEMBL, 9 Novoseek, and/or 10fRNAdb)
    About This Section

    PubMed articles for ZSCAN4 gene integrated from 9 sources:
    (articles sorted by number of sources associating them with ZSCAN4)
        Utopia: connect your pdf to the dynamic
    world of online information

    1. The status, quality, and expansion of the NIH full-length cDNA project: the Mammalian Gene Collection (MGC). (PubMed id 15489334)1, 2 Gerhard D.S....Malek J. (2004)
    2. Complete sequencing and characterization of 21,243 full-length human cDNAs. (PubMed id 14702039)1, 2 Ota T.... Sugano S. (2004)
    3. Zscan4 regulates telomere elongation and genomic stab ility in ES cells. (PubMed id 20336070)1 Zalzman M....Ko M.S. (2010)
    4. An atlas of combinatorial transcriptional regulation in mouse and man. (PubMed id 20211142)1 Ravasi T....Hayashizaki Y. (2010)
    5. Functional proteomics mapping of a human signaling pathway. (PubMed id 15231748)1 Colland F....Gauthier J.M. (2004)
    6. Generation and initial analysis of more than 15,000 full-length human and mouse cDNA sequences. (PubMed id 12477932)1 Strausberg R.L....Marra M.A. (2002)

    (in PubMed, OMIM, and NCBI Bookshelf)
    About This Section
     ANDOR
    Aliases
    Free Text  

      Query String
    PubMed
    OMIM
    NCBI Bookshelf
      (Note: In FireFox, select the above section and copy using Ctrl-C)

    (According to Entrez Gene, HGNC, AceView, euGenes, Ensembl, miRBase, ECgene, Kegg, and/or H-InvDB)
    About This Section
    Entrez Gene: 201516 HGNC: 23709 AceView: ZNF494 Ensembl:ENSG00000180532 euGenes: HUgn201516
    ECgene: ZSCAN4 H-InvDB: ZSCAN4

    (According to HUGE)
    About This Section
      --

    (According to PharmGKB, ATLAS, HORDE, IMGT, LEIDEN, UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL,
    Wikipedia and/or GeneReviews via UniProtKB/Swiss-Prot)
    About This Section
    NameDescription
    PharmGKB entry for ZSCAN4 Pharmacogenomics, SNPs, Pathways

    (Patent information from GeneIP,
    Licensable technologies from WIS Yeda, Salk, Tufts,
    IP news from LifeMap Sciences, Inc.)
    About This Section
    Patent Information for ZSCAN4 gene:
    Search GeneIP for patents involving ZSCAN4

    GeneCards and IP:
    Japan Patent Office Licenses GeneCards     European Patent Office Licenses GeneCards     Improving the IP Search



    (Antibodies, recombinant proteins, and assays from EMD Millipore, R&D Systems, OriGene, QIAGEN, GenScript, Cell Signaling Technology, SABiosciences, Novus Biologicals, Sino Biological, Enzo Life Sciences, Abcam, ProSpec, Cloud-Clone Corp., Thermo Fisher Scientific, LSBio, Gene Editing from DNA2.0 and Sirion Biotech, Clones from EMD Millipore, OriGene, GenScript, Sino Biological, DNA2.0, SwitchGear Genomics, Vector BioLabs, Sirion Biotech, Cell lines from GenScript, and LifeMap BioReagents, PCR Arrays from SABiosciences, Drugs and/or compounds from EMD Millipore, Tocris Bioscience, and/or Enzo Life Sciences, In Situ Hybridization Assays from
    Advanced Cell Diagnostics, Animal models from inGenious Targeting Laboratory, genOway)
    About This Section

     
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    GeneCards Homepage - Last full update: 23 Oct 2013 - Incrementals: 4 Nov 2013 , 7 Nov 2013 , 23 Jan 2014

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    Category
    (GIFtS: 73)
    transforming growth factor, beta 1
    GIFtS Group
    The GeneCards human gene database gene index: 1 3 5 6 A B C D E F G H I J K L M N O P Q R S T U V W X Y Z 


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