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WDR19 Gene

protein-coding GIFtS: 51

GC04P038860

WD repeat domain 19
Symbol approved by the HUGO Gene Nomenclature Committee (HGNC) database

(According to ¹HGNC, ²Entrez Gene,
³UniProtKB/Swiss-Prot, ⁴UniProtKB/TrEMBL, ⁵OMIM, ⁶GeneLoc , and/or ⁷Ensembl, ⁸miRBase)
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Aliases
FLJ23127 ^¹, ²
KIAA1638 ^¹, ², ³
ORF26 ^¹, ²
PWDMP ^²
Pwdmp ^¹

Descriptions
WD repeat domain 19 ^²
WD repeat membrane protein PWDMP ^²

External Ids
HGNC: 18340¹
Entrez Gene: 57728²
UniProtKB: Q8NEZ3³
Ensembl: ENSG00000157796⁷

Search outside databases for aliases for WDR19 gene

Previous GC identifers: GC00U991193 GC04P039006

(According to Entrez Gene, Wikipedia's Gene Wiki,
UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL)
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EntrezGene summary for WDR19:

This gene encodes a member of the WD repeat protein family. WD repeats are minimally conserved

regions of approximately 40 amino acids typically bracketed by gly-his and trp-asp (GH-WD), which

may facilitate formation of heterotrimeric or multiprotein complexes. Members of this family are

involved in a variety of cellular processes, including cell cycle progression, signal

transduction, apoptosis, and gene regulation. This protein contains six WD repeats, a clathrin

heavy-chain repeat, and three transmembrane domains. This gene is conserved from C. elegans to

human. It may participate in androgen-regulated signaling mechanisms or in the vesicular

trafficking of androgen-regulated secretory processes. Alternatively spliced transcript variants

encoding distinct isoforms have been reported but the full-length nature of one of these variants

has not been defined. [provided by RefSeq]

(According to GeneLoc and/or HGNC, and/or
Entrez Gene (NCBI build 36),
and/or miRBase,
Genomic Views according to UCSC and Ensembl, Transcription factor binding sites according to SABiosciences)
About This Section

Genomic View:

UCSC Golden Path with GeneCards custom track

Transcription factor binding sites upstream to the WDR19 gene

Entrez Gene cytogenetic band: 4p14 Ensembl cytogenetic band: 4p14 HGNC cytogenetic band: 4p14

WDR19 Gene in genomic location: bands according to Ensembl, locations according to (and/or Entrez Gene and/or Ensembl if different)

GeneLoc gene densities for chromosome 4 GeneLoc Exon Structure

GeneLoc location for GC04P038860: (about GC identifiers)

Start:	38,858,173 bp from pter
End:	38,963,825 bp from pter
Size:	105,653 bases
Orientation:	plus strand

RefSeq DNA sequence:

NC_000004.10 NT_016297.15

(According to ¹UniProtKB, and/or Ensembl, Phosphorylation sites according to ²Phosphosite, RefSeq according to NCBI, PDB rendering according to OCA and/or Proteopedia, Recombinant Proteins from Invitrogen, Millipore, Sigma-Aldrich, R&D Systems, Enzo Life Sciences, Abnova, OriGene and/or, Abcam,
Biochemical Assays by Invitrogen, Millipore, R&D Systems, Cell Signaling Technology, and/or Enzo Life Sciences, Ontologies according to Gene Ontology Consortium 01 Apr 2009 and Entrez Gene, Antibodies by Invitrogen, Millipore, Sigma-Aldrich, R&D Systems, Cell Signaling Technology, Abcam, Abnova, and/or Novus Biologicals)
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UniProtKB/Swiss-Prot: WDR19_HUMAN, Q8NEZ3 (See protein sequence)

Recommended Name: WD repeat-containing protein 19

Size: 1342 amino acids; 151581 Da

Developmental stage: Expressed in fetal lung

Secondary accessions: Q8N5B4 Q9H5S0 Q9HCD4

Alternative splicing: 2 isoforms: Q8NEZ3-1 Q8NEZ3-2

Post-translational modifications:

View phosphorylation sites using PhosphoSite²

REFSEQ proteins: NP_079408.3

ENSEMBL proteins:

ENSP00000382730 ENSP00000288634 ENSP00000382717

Human Recombinant Proteins

	Browse Drug Discovery Central at Invitrogen for human recombinant proteins
	Browse Purified and Recombinant Proteins at Millipore
	Browse Human Recombinant Proteins at Sigma-Aldrich
	Browse R&D Systems for human recombinant proteins
	Browse recombinant and purified proteins available from Enzo Life Sciences
	Recombinant Proteins from Abcam (WDR19)
	Human Recombinant Proteins from Abnova (WDR19)

Browse Origene for full length recombinant human proteins expressed in human HEK293 cells

Antibodies for WDR19:

	Browse Antibodies Central at Invitrogen
	Browse Millipore's Extensive Line of Mono- and Polyclonal Antibodies
	Browse Antibodies at Sigma-Aldrich
	Browse R&D Systems for Antibodies
	Antibodies from Abcam (WDR19), each with their Abpromise^SM
	Monoclonal and Polyclonal Antibodies from Abnova (WDR19)
	Novus Biologicals Antibodies for WDR19

Assays for WDR19:

	Browse Invitrogen for biochemical assays
	Browse Kits and Assays available from Millipore
	Browse R&D Systems for biochemical assays
	Browse biochemical assays available from Enzo Life Sciences

(According to InterPro, ProtoNet, UniProtKB, and/or BLOCKS, Sets of similar genes according to GeneDecks)
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Graphical View of Domain Structure for InterPro Entry Q8NEZ3

ProtoNet protein and cluster: Q8NEZ3

UniProtKB/Swiss-Prot: WDR19_HUMAN, Q8NEZ3

Similarity: Contains 6 TPR repeats

Similarity: Contains 6 WD repeats

(According to MGI Jun 06 2009, UniProtKB, IUBMB,and/or Genatlas,
shRNA from OriGene, Sigma-Aldrich, RNAi from Sigma-Aldrich,
RNAi Products, Clones, and Q-PCR Products from Invitrogen, Millipore, OriGene, and/or Abnova, siRNAs from Applied Biosystems, SYBR primers from OriGene, Cell-based Assays from Millipore, Ontologies according to Gene Ontology Consortium 01 Apr 2009 via Entrez Gene.)
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Inhib. RNA:		Invitrogen RNAi Products for gene knock-down (WDR19)
		Browse for Gene Knock-down Tools from Millipore
		Abnova Chimera RNAi Products for Gene knock-down (WDR19)

OriGene 29mer shRNA kit in GFP-retroviral vector: NM_025132

Applied Biosystems Silencer® siRNAs for WDR19

Sigma-Aldrich siRNA for WDR19

Sigma-Aldrich shRNA for WDR19

Explore Sigma-Aldrich super-pooled esiRNAs

Clones:		Invitrogen Clones for WDR19
		Browse Clones for the Expression of Recombinant Proteins Available from Millipore

OriGene GFP tagged cDNA clone in CMV expression vector: NM_025132
Myc/DDK tagged cDNA clone in CMV expression vector: NM_025132
untagged cDNA clone in CMV expression vector: NM_025132

Primers:

Browse Quantitative PCR Central at Invitrogen for Q-PCR LUX™ Primers

OriGene genome-wide validated SYBR primer pairs: NM_025132

UniProtKB/Swiss-Prot: WDR19_HUMAN, Q8NEZ3

Induction: By androgenic hormones. Expression increased 3-fold in an androgen-stimulated

androgen-sensitive prostate adenocarcinoma cell line compared with androgen-deprived cells

1 Gene Ontology (GO) molecular function term (links to tree view):

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0005515	protein binding	IEA	--

About this table

(Pathways according to Invitrogen (maps by GeneGo), Millipore, Cell Signaling Technology, Sigma-Aldrich, KEGG and/or UniProtKB,
Sets of similar genes according to GeneDecks, Proteins Network according to SABiosciences, Interactions according to ¹UniProtKB, ²MINT, and/or ³STRING, with links to IntAct and Ensembl, Ontologies according to Gene Ontology Consortium 01 Apr 2009 via Entrez Gene.)
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1 Interacting protein for WDR19 (ENSP00000288634³) via UniProtKB, MINT, and/or STRING

Interactant		Interaction Details
GeneCard	External ID(s)	Interaction Details
VIL1	ENSP00000248444³	STRING (score=.7)

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2 Gene Ontology (GO) biological process terms (links to tree view):

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0006886	intracellular protein transport	IEA	--
GO:0016192	vesicle-mediated transport	IEA	--

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(Chemical Compounds according to UniProtKB, Enzo Life Sciences, Sigma-Aldrich, Tocris Bioscience, and/or Novoseek and Drugs according to Enzo Life Sciences and/or PharmGKB)
About This Section

Browse drugs & compounds from Enzo Life Sciences

Browse Small Molecules at Sigma-Aldrich

Browse Tocris compounds for WDR19

(GenBank/EMBL/DDBJ Accessions according to
Unigene (Build 219 Homo sapiens; Jun 2 2009) or GenBank,
RefSeq according to Entrez Gene,
DOTS (version 10), and/or AceView,
non coding RNAs according to RNAdb,
ESTs according to GeneTide,
exon structure from GeneLoc, alternative splicing isoforms according to ASD and/or ECgene,
RNAi Products from Invitrogen, Millipore, and/or Abnova,
siRNAs from Applied Biosystems, Sigma-Aldrich,
shRNA from Sigma-Aldrich, OriGene,
Tagged/untagged cDNA clones from OriGene,
Expression Assays from Applied Biosystems)
About This Section

Inhib. RNA:		Invitrogen RNAi Products for gene knock-down (WDR19)
		Browse for Gene Knock-down Tools from Millipore
		Abnova Chimera RNAi Products for Gene knock-down (WDR19)

OriGene 29mer shRNA kit in GFP-retroviral vector: NM_025132

Sigma-Aldrich siRNA for WDR19

Sigma-Aldrich shRNA for WDR19

Explore Sigma-Aldrich super-pooled esiRNAs

Applied Biosystems Silencer® siRNAs:

NM_025132

REFSEQ mRNAs for WDR19 gene:

NM_025132.3

Applied Biosystems TaqMan ® Gene Expression Assays:

NM_025132

              OriGene GFP tagged cDNA clone in CMV expression vector: NM_025132
                                 Myc/DDK tagged cDNA clone in CMV expression vector: NM_025132
                                 untagged cDNA clone in CMV expression vector: NM_025132

Additional cDNA sequence:

AB046858.1 AK026780.1 AK225588.1 AK294730.1 AK296112.1 AK304161.1 AK308803.1 AY029257.1

BC032578.1 BC131501.1

12 DOTS entries:

DT.114006 DT.92040399 DT.100026571 DT.100755175 DT.92416906 DT.92057945 DT.99971287 DT.92418702

DT.100736238 DT.121259470 DT.95343830 DT.97839733

24/97 AceView cDNA sequences (see all 97 ):

BU948584 AW450839 BU621014 BP376333 AA281901 AA281724 AA445953 BQ923434

AB046858 BM719838 BU686662 BU679219 AL702961 AI969388 AA934791 AA363808

BQ181511 AL711455 AY029257 BQ950404 BU948827 BE046347 NM_025132 AA363807

highest scoring ESTs for WDR19:

AA581393 AA007605 AA281724 AA464597 AA653325 AA858065 AA934791 AI355348 AI769134 AI969388

Unigene Cluster for WDR19:

WD repeat domain 19

Hs.438482 [show with all ESTs]

Unigene Representative Sequence: NM_025132

GeneLoc Exon Structure

3 Ensembl transcripts including schematic representations:

ENST00000399836 ENST00000288634 ENST00000399820

(Experimental results according to ¹GeneNote and GNF BioGPS,
probe sets-to-genes annotations according to ²GeneAnnot , ³GeneTide , Sets of similar genes according to GeneDecks, Electronic Northern calculations according to data from UniGene (Build 219 Homo sapiens), SAGE tags according to CGAP, plus additional links to SOURCE, and/or GNF BioGPS, and/or EXPOLDB, and/or UniProtKB,
Expression Assays from Applied Biosystems )
About This Section

WDR19 expression in normal and diseased human tissues

Applied Biosystems TaqMan ® Gene Expression Assays for WDR19

¹ / ² / ³

6 probe-sets matching WDR19 gene

Affymetrix probe-set	Array	# genes	Sensitivity	Specificity	Correlation	Length	Gb_Accession	Consensus	Uniqueness	Score	Rank
Affymetrix probe-set	Array	GeneAnnot data			GeneNote data		GeneTide data
56146_at²^{, 3}	U95-B	1	0.94	1.00	1.00	1.00	AA581393	1.00	1.00	1.00	1
232163_at²^{, 3}	U133-B	1	1.00	1.00	--	--	AI982884	0.60	1.00	0.82	1
220917_s_at²^{, 3}	U133-A	1	1.00	1.00	--	--	NM_025132	0.60	1.00	0.82	1
232163_at²	U133Plus2	1	1.00	1.00	--	--	--	--	--	--	--
1555314_at²	U133Plus2	1	1.00	1.00	--	--	--	--	--	--	--
220917_s_at²	U133Plus2	1	1.00	1.00	--	--	--	--	--	--	--

GeneDecks WDR19 for binary patterns associated with the probe-sets selected above

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Data from

(Publications) and GNF BioGPS
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CGAP SAGE TAG: TAACCTTGCT

SOURCE GeneReport for Unigene cluster: Hs.438482
UniProtKB/Swiss-Prot: WDR19_HUMAN, Q8NEZ3
Tissue specificity: Some isoforms are tissue-specific. Highly expressed in the prostate. Lower
expression in the cerebellum, pituitary gland, fetal lung, and pancreas. In normal prostate,
expressed in both basal and luminal epithelial cells. No expression detected in fibromuscular
stromal cells, endothelial cells, or infiltrating lymphocytes. Uniformed expression in prostate
adenocarcinoma cells

(Orthologs according to ^1,2HomoloGene (²older version, for species not in ¹newer version), ³euGenes, ⁴SGD and/or ⁵MGI Jun 06 2009, with possible further links to Flybase and/or WormBase, Gene Trees according to Ensembl)
About This Section

Orthologs for WDR19 gene from 5/11 species (see all 11 )

Organism	Gene	Locus	Description	Human Similarity	NCBI accessions
dog (Canis familiaris)	WDR19¹	--	WD repeat domain 19	91.02(n) 92.95(a)	479113 XM_536260.2 XP_536260.2
chimpanzee (Pan troglodytes)	WDR19¹	--	WD repeat domain 19	99.65(n) 99.85(a)	461172 XM_517152.2 XP_517152.2
cow (Bos taurus)	WDR19¹	--	WD repeat domain 19	90.31(n) 91.57(a)	533633 XM_613099.3 XP_613099.3
rat (Rattus norvegicus)	Wdr19¹	--	WD repeat domain 19	85.58(n) 89.93(a)	305349 XM_223412.3 XP_223412.3
mouse (Mus musculus)	Wdr19^{1, 5}	5⁵	WD repeat domain 19^{1, 5}	85.83(n)¹ 89.39(a)¹	213081¹ NM_153391.1¹ NP_700440.1¹ AA388062⁵ AA879550⁵ (see all 22)

About this table Species with no ortholog for WDR19

ENSEMBL Gene Tree for WDR19

(Paralogs according to ¹HomoloGene
and ²Ensembl, Pseudogenes according to ³Pseudogene.org)
About This Section

(According to the ¹NCBI SNP Database, ²Ensembl, ³PupaSUITE, and UniProtKB, Linkage Disequilibrium by HapMap, Genotyping Reagents from Applied Biosystems)
About This Section

10/588 NCBI SNPs in WDR19 are shown (see all 588 )
(Click for Applied Biosystems TaqMan ® Genotyping Assay) (see all 157)

AB	SNP ID	Valid	Chr 4 pos	Sequence	Recs	AA Chg	Type	More	Recs	Allele freq	Pop	Total sample	More
AB	Genomic Data				Transcription Data				Allele Frequencies
				--	--	--		--	--				--
	rs1451819^1,2	A,C,F,H,O	38860127(-)	`TGGGCC/TAAATA`	1	--	ng5¹		8		MN NA EU EA WA	744
	rs10030029^1,2	A,C,F,H	38963950(+)	`GTTTAC/TACAAA`	1	--	ng3¹		5		NS EU EA WA	576
	rs10030213^1,2	A,C,F,H	38964052(+)	`TTATAC/TCTATC`	1	--	ng3¹		7		NA EU EA WA	560
	rs1451820^1,2	C,F,H,O	38859842(-)	`TTGTAT/CAGTAT`	1	--	ng5¹		13		MN EA NA EU WA	2464
	rs4367182^1,2	A,C,F,H	38860166(+)	`GTCTTT/CTTTCC`	1	--	ng5¹ trp³		9		NA EU EA WA	736
	rs16995209^1,2	C,F,H	38944144(+)	`ACGATG/AGCATG`	1	S/G	mis¹ ese³		11		NA EU EA WA	974
--	rs17288883^1,2	C,F	38964134(-)	`AACAA-/ACAA TGTAT`	1	--	ng3¹		1		NS	170
	rs7564^1,2	C,F	38963717(+)	`TTACAC/TACTCT`	1	--	ut3¹		7		MN EA NA NS	1586
--	rs17335661^1,2	F	38963964(-)	`CTTAAT/CAGCTG`	1	--	ng3¹		1		NS	180
--	rs10027977^1,2	C	38964250(+)	`GGAGTA/CCATAC`	1	--	ng3¹		0	--	--	--	--

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HapMap Linkage Disequilibrium images for WDR19 (up to first 250kb)

(in which this Gene is Involved, According to OMIM, UniProtKB, PharmGKB, Genatlas, GeneTests, Blood group antigen gene mutations by BGMUT, HGMD, GAD, HuGE Navigator, BCGD, and/or TGDB.)
About This Section

OMIM: 608151

(Possibly Related Articles in Doctor's Guide)
About This Section

(in PubMed. Associations of this gene to articles via ¹Novoseek, ²HGNC, ³Entrez Gene, ⁴UniProtKB/Swiss-Prot, ⁵UniProtKB/TrEMBL, ⁶GAD, and/or ⁷PharmGKB)
About This Section

9 PubMed articles for WDR19 gene:

Isolation and characterization of human and mouse WDR19,a novel WD- repeat protein exhibiting androgen-regulated expression in prostate epithelium. (PubMed id 12906858)^{1, 2, 3, 4} Lin B.... Nelson P.S. (2003)
The status, quality, and expansion of the NIH full-length cDNA project: the Mammalian Gene Collection (MGC). (PubMed id 15489334)^{3, 4} Gerhard D.S....Malek J. (2004)
Complete sequencing and characterization of 21,243 full-length human cDNAs. (PubMed id 14702039)^{3, 4} Ota T....Sugano S. (2004)
Prediction of the coding sequences of unidentified human genes. XVIII. The complete sequences of 100 new cDNA clones from brain which code for large proteins in vitro. (PubMed id 10997877)^{3, 4} Nagase T....Ohara O. (2000)
WDR19 expression is increased in prostate cancer compared with normal cells, but low-intensity expression in cancers is associated with shorter time to biochemical failures and local recurrence. (PubMed id 18316561)³ Lin B....Akslen L.A. (2008)
Generation and initial analysis of more than 15,000 full-length human and mouse cDNA sequences. (PubMed id 12477932)³ Strausberg R.L....Marra M.A. (2002)
Construction of expression-ready cDNA clones for KIAA genes: manual curation of 330 KIAA cDNA clones. (PubMed id 12168954)⁴ Nakajima D.... Nagase T. (2002)
Construction and characterization of a full length-enriched and a 5'- end-enriched cDNA library. (PubMed id 9373149)³ Suzuki Y.... Sugano S. (1997)
Oligo-capping: a simple method to replace the cap structure of eukaryotic mRNAs with oligoribonucleotides. (PubMed id 8125298)³ Maruyama K. and Sugano S. (1994)

(in PubMed, OMIM, and NCBI Bookshelf)
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		AND	OR
Aliases
Free Text

Query String		PubMed OMIM NCBI Bookshelf
	(Note: In FireFox, select the above section and copy using Ctrl-C)

(According to Entrez Gene, HGNC, AceView, euGenes, Ensembl, miRBase, ECgene, and/or H-InvDB)
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Entrez Gene: 57728	HGNC: 18340	AceView: WDR19	Ensembl:ENSG00000157796	euGenes: HUgn57728
ECgene: WDR19	H-InvDB: WDR19

(According to HUGE)
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HUGE: KIAA1638

(According to ATLAS, HORDE, IMGT, MTDB, LEIDEN, UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL,
Wikipedia and/or GeneReviews via UniProtKB/Swiss-Prot)
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Licensable Technologies for WDR19
(Available from WIS Yeda, Salk, Tufts)

About This Section

(Reagents available from Applied Biosystems, Antibodies and assays by Cell Signaling Technology, Abcam, Novus Biologicals,
Sigma-Aldrich, R&D Systems, Millipore, Abnova, and/or Invitrogen, Clones available from OriGene,and/or Invitrogen, Drugs and/or compounds by Sigma-Aldrich,
Enzo Life Sciences, and/or Tocris Bioscience)
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Products for WDR19:

TaqMan ® Gene Expression Assays

TaqMan ® Genotyping Assays

Free SNP selection tool

Invitrogen iPath Pathways	Invitrogen BLOCK-iT™ RNAi
Invitrogen Antibodies	Invitrogen Assays
Invitrogen Clones	Invitrogen Q-PCR Products
Invitrogen Human Recombinant Kinases	Invitrogen Custom Antibody and Peptide Service
Invitrogen Proteins / Assays / Screening Services	Search Invitrogen catalog for WDR19-related products

	Millipore Custom Antibody & Bulk Services
	Millipore Preclinical / Clinical Development Services
	Millipore Immunoassay Services
	Millipore Target Screening & Profiling Services

	Predesigned and custom siRNAs for WDR19		Browse antibodies at Sigma-Aldrich
	Explore super-pooled esiRNAs		Browse proteins at Sigma-Aldrich
	Lentivirus-delivered shRNAs for WDR19		Browse small molecules at Sigma-Aldrich
	"Your Favorite Gene" Pathways		feedback

	Antibodies		Primer Pairs
	Cell Culture Products		ELISAs
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	Multiplex/Array Assay Kits & Reagents		ELISpot Kits & Development Modules
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WDR19 Gene

protein-coding GIFtS: 51

Aliases &
Descriptions
for WDR19

Summaries
for WDR19

Genomic Location
for WDR19

Proteins
for WDR19

Protein Domains/
Families
for WDR19

Gene Function
for WDR19

Pathways & Interactions
for WDR19

Drugs & Compounds
for WDR19

Transcripts
for WDR19

Expression
for WDR19

Orthologs
for WDR19

Paralogs
for WDR19

SNPs/Variants
for WDR19

Disorders & Mutations
for WDR19

Medical News
for WDR19

Publications
for WDR19

Search for WDR19

Genome Databases showing WDR19

Other Databases showing WDR19

Specialized Databases showing WDR19

Licensable Technologies
for WDR19

Services
for WDR19

Gene Index: 3 5 A B C D E F G H I J K L M N O P Q R S T U V W X Y Z

WDR19 Gene

protein-coding GIFtS: 51

Aliases & Descriptionsfor WDR19

Summariesfor WDR19

Genomic Locationfor WDR19

Proteinsfor WDR19

Protein Domains/Familiesfor WDR19

Gene Functionfor WDR19

Pathways & Interactionsfor WDR19

Drugs & Compoundsfor WDR19

Transcriptsfor WDR19

Expression for WDR19

Orthologsfor WDR19

Paralogsfor WDR19

SNPs/Variantsfor WDR19

Disorders & Mutationsfor WDR19

Medical Newsfor WDR19

Publicationsfor WDR19

Search for WDR19

Genome Databases showing WDR19

Other Databases showing WDR19

Specialized Databases showing WDR19

Licensable Technologiesfor WDR19

Servicesfor WDR19

Gene Index: 3 5 A B C D E F G H I J K L M N O P Q R S T U V W X Y Z

Aliases &
Descriptions
for WDR19

Summaries
for WDR19

Genomic Location
for WDR19

Proteins
for WDR19

Protein Domains/
Families
for WDR19

Gene Function
for WDR19

Pathways & Interactions
for WDR19

Drugs & Compounds
for WDR19

Transcripts
for WDR19

Expression
for WDR19

Orthologs
for WDR19

Paralogs
for WDR19

SNPs/Variants
for WDR19

Disorders & Mutations
for WDR19

Medical News
for WDR19

Publications
for WDR19

Licensable Technologies
for WDR19

Services
for WDR19