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Category   Symbol Source: HGNC EntrezGene Ensembl GeneCards RNA genes CroW21
GIFtS

WASH4P Gene

protein-coding   GIFtS: 20
GCID: GC16M000064

WAS Protein Family Homolog 4 Pseudogene

(Previous names: family with sequence similarity 39, member C pseudogene)
(Previous symbol: FAM39CP)
  Search for WASH4P
at  

(According to 1HGNC, 2Entrez Gene,
3UniProtKB/Swiss-Prot, 4UniProtKB/TrEMBL, 5OMIM, 6GeneLoc, 7Ensembl, 8DME, 9miRBase, 10fRNAdb, 12H-InvDB, 13NCBI, 14NONCODE, and/or 15RNAdb)
About This Section

Aliases
WAS Protein Family Homolog 4 Pseudogene1 2
FAM39CP1 2 3
Family With Sequence Similarity 39, Member C Pseudogene1
CXYorf1P2
Protein FAM39CP3

External Ids:    HGNC: 141261   Entrez Gene: 3746772   Ensembl: ENSG000002347697   UniProtKB: A8MWX33   

Export aliases for WASH4P gene to outside databases

Previous GC identifer: GC16M000008


(According to Entrez Gene, Tocris Bioscience, Wikipedia's Gene Wiki, PharmGKB,
UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL)
About This Section

GeneCards Summary for WASH4P Gene: 
WASH4P (WAS protein family homolog 4 pseudogene) is a protein-coding gene.

UniProtKB/Swiss-Prot: WASH4_HUMAN, A8MWX3
Function: Acts as a nucleation-promoting factor at the surface of endosomes, where it recruits and activates the
Arp2/3 complex to induce actin polymerization, playing a key role in the fission of tubules that serve as
transport intermediates during endosome sorting (By similarity)




(According to GeneLoc and/or HGNC, and/or
Entrez Gene (NCBI build 37),
and/or miRBase,
Genomic Views according to UCSC (hg19) and Ensembl (release 73), Regulatory elements and Epigenetics data according to QIAGEN, SABiosciences, and/or SwitchGear Genomics)
About This Section
Regulatory elements:
   Search SABiosciences Regulatory transcription factor binding sites for WASH4P
         Other transcription factors

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Epigenetics:
QIAGEN PyroMark CpG Assay predesigned Pyrosequencing DNA Methylation assays in human, mouse, rat WASH4P


Genomic Location:
Genomic View: UCSC Golden Path with GeneCards custom track

Entrez Gene cytogenetic band: 16p13.3   Ensembl cytogenetic band:  16p13.3   HGNC cytogenetic band: 16p13.3

WASH4P Gene in genomic location: bands according to Ensembl, locations according to (and/or Entrez Gene and/or Ensembl if different)
WASH4P gene location

GeneLoc information about chromosome 16         GeneLoc Exon Structure

GeneLoc location for GC16M000064:  view genomic region     (about GC identifiers)

Start:
64,043 bp from pter      End:
69,452 bp from pter
Size:
5,410 bases      Orientation:
minus strand

(According to UniProtKB, HORDE, neXtProt, Ensembl, and/or Reactome, Modification sites according to PhosphoSitePlus, Specific Peptides from DME, Protein expression images according to data from SPIRE 1MOPED, 2PaxDb, and 3MAXQB RefSeq according to NCBI, PDB rendering according to OCA and/or Proteopedia, Recombinant Proteins from EMD Millipore, R&D Systems, GenScript, Enzo Life Sciences, OriGene, Novus Biologicals, Sino Biological, ProSpec, and/or Cloud-Clone Corp.,
Biochemical Assays by EMD Millipore, R&D Systems, OriGene, GenScript, Cell Signaling Technology, Enzo Life Sciences, and/or Cloud-Clone Corp., Ontologies according to Gene Ontology Consortium 01 Oct 2013 and Entrez Gene, Antibodies by EMD Millipore, R&D Systems, GenScript, Cell Signaling Technology, OriGene, Novus Biologicals, Thermo Fisher Scientific, LSBio, Abcam, and/or Cloud-Clone Corp.)
About This Section

UniProtKB/Swiss-Prot: WASH4_HUMAN, A8MWX3 (See protein sequence)
Recommended Name: Putative WAS protein family homolog 4  
Size: 477 amino acids; 51595 Da
Subunit: Component of the WASH complex, composed of F-actin-capping protein subunit alpha (CAPZA1, CAPZA2 or
CAPZA3), F-actin-capping protein subunit beta (CAPZB), WASH (WASH1, WASH2P, WASH3P, WASH4P or WASH6P), FAM21
(FAM21A, FAM21B or FAM21C), KIAA1033, KIAA0196 and CCDC53. Interacts (via WHD1 region) with FAM21C; the
interaction is direct (By similarity)
Subcellular location: Early endosome membrane (By similarity). Recycling endosome membrane (By similarity).
Note=Localization to the endodome membrane is mediated via its interaction with FAM21 (By similarity)
Miscellaneous: WASH genes duplicated to multiple chromosomal ends during primate evolution, with highest copy
number reached in humans, whose WASH repertoires probably vary extensively among individuals (PubMed:18159949).
It is therefore difficult to determine which gene is functional or not. The telomeric region of chromosome 9p is
paralogous to the pericentromeric regions of chromosome 9 as well as to 2q. Paralogous regions contain 7
transcriptional units. Duplicated WASH genes are also present in the Xq/Yq pseudoautosomal region, as well as on
chromosome 1 and 15. The chromosome 16 copy seems to be a pseudogene
Caution: Could be the product of a pseudogene

Explore the universe of human proteins at neXtProt for WASH4P: NX_A8MWX3

Explore proteomics data for WASH4P at MOPED 

WASH4P Protein expression data from MOPED1, PaxDb2 and MAXQB3 :    About this image 

WASH4P Protein Expression

ENSEMBL proteins: 
 ENSP00000317542   ENSP00000454316   ENSP00000457756   ENSP00000457950   ENSP00000441402  

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Gene Ontology (GO): 5/6 cellular component terms (GO ID links to tree view) (see all 6):    About this table

GO IDQualified GO termEvidencePubMed IDs
GO:0005769early endosome ----
GO:0031901early endosome membrane ----
GO:0043231intracellular membrane-bounded organelle ----
GO:0055037recycling endosome ----
GO:0055038recycling endosome membrane ----

WASH4P for ontologies           About GeneDecksing



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(According to HGNC, IUPHAR, InterPro, ProtoNet, UniProtKB, and/or BLOCKS, Sets of similar genes according to GeneDecks)
About This Section
HGNC Gene Families: 
WASH: WAS protein homologs

1 InterPro protein domain:
 IPR021854 WASH_WASD

Graphical View of Domain Structure for InterPro Entry A8MWX3

ProtoNet protein and cluster: A8MWX3

UniProtKB/Swiss-Prot: WASH4_HUMAN, A8MWX3
Similarity: Belongs to the WASH1 family
Similarity: Contains 1 WH2 domain


WASH4P for domains           About GeneDecksing


(According to 1UniProtKB, Genatlas, LifeMap Discovery™, IUBMB, and/or 2DME, Human phenotypes from GenomeRNAi, Animal models from MGI Mar 06 2013, inGenious Targeting Laboratory, genOway,
bound targets from SABiosciences, miRNA Gene Targets from miRTarBase, shRNA from OriGene, RNAi from EMD Millipore, siRNAs from OriGene, QIAGEN, microRNA from QIAGEN, Gene Editing from DNA2.0, Sirion Biotech, Clones from EMD Millipore, OriGene, SwitchGear Genomics, GenScript, Sino Biological, DNA2.0, Vector BioLabs, and Sirion Biotech, Cell Lines from GenScript, LifeMap BioReagents, In Situ Hybridization Assays from Advanced Cell Diagnostics, Ontologies according to Gene Ontology Consortium 01 Oct 2013 via Entrez Gene.)
About This Section

Molecular Function:

     UniProtKB/Swiss-Prot Summary: WASH4_HUMAN, A8MWX3
Function: Acts as a nucleation-promoting factor at the surface of endosomes, where it recruits and activates the
Arp2/3 complex to induce actin polymerization, playing a key role in the fission of tubules that serve as
transport intermediates during endosome sorting (By similarity)

     Gene Ontology (GO): 2 molecular function terms (GO ID links to tree view):    About this table

GO IDQualified GO termEvidencePubMed IDs
GO:0003779actin binding ----
GO:0043014alpha-tubulin binding ----
     
WASH4P for ontologies           About GeneDecksing


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(Pathways according to EMD Millipore, R&D Systems, Cell Signaling Technology, KEGG, PharmGKB, BioSystems, Sino Biological, Reactome, Tocris Bioscience, GeneGo (Thomson Reuters), QIAGEN, and/or UniProtKB, Sets of similar genes according to GeneDecks, Interaction Networks according to SABiosciences, and/or STRING, Interactions according to 1UniProtKB, 2MINT, 3I2D, and/or 4STRING, with links to IntAct and Ensembl, Ontologies according to Gene Ontology Consortium 01 Oct 2013 via Entrez Gene).
About This Section



Interactions:

    Search SABiosciences Gene Network CentralTM Interacting Genes and Proteins Networks for WASH4P

STRING Interaction Network Preview (showing 5 interactants - click image to see 12)

5/12 Interacting proteins for WASH4P (ENSP000003175424) via UniProtKB, MINT, STRING, and/or I2D (see all 12)
InteractantInteraction Details
GeneCardExternal ID(s)
CAPZA1ENSP000002631684STRING: ENSP00000263168
CCDC53ENSP000002400794STRING: ENSP00000240079
FAM21AENSP000002826334STRING: ENSP00000282633
FAM21BENSP000003512594STRING: ENSP00000351259
FAM21CENSP000003375414STRING: ENSP00000337541
About this table

Gene Ontology (GO): 3 biological process terms (GO ID links to tree view):    About this table

GO IDQualified GO termEvidencePubMed IDs
GO:0016197endosomal transport ----
GO:0034314Arp2/3 complex-mediated actin nucleation ----
GO:0042147retrograde transport, endosome to Golgi ----

WASH4P for ontologies           About GeneDecksing



(Chemical Compounds according to UniProtKB, Enzo Life Sciences, EMD Millipore, Tocris Bioscience HMDB, BitterDB, and/or Novoseek, Ligands according to IUPHAR, and Drugs according to DrugBank, Enzo Life Sciences, and/or PharmGKB, with drugs/clinical trials/news search links to CenterWatch)
About This Section
Browse Small Molecules at EMD Millipore
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Search CenterWatch for drugs/clinical trials and news about WASH4P / WASH4

(Secondary structures according to fRNAdb,
GenBank/EMBL/DDBJ Accessions according to
Unigene (Build 236 Homo sapiens; Apr 25 2013) or GenBank,
RefSeq according to Entrez Gene,
DOTS (version 10), and/or AceView, transcript ids from Ensembl with links to UCSC,
Conferences by KenesGroup, exon structure from GeneLoc, alternative splicing isoforms according to ASD and/or ECgene,
RNAi Products from EMD Millipore,
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About This Section

13 Ensembl transcripts including schematic representations, and UCSC links where relevant:
ENST00000326592(uc002cfg.1) ENST00000474799 ENST00000470283 ENST00000468574
ENST00000564273 ENST00000477365 ENST00000495251 ENST00000462860 ENST00000470725
ENST00000487568 ENST00000483390 ENST00000538848 ENST00000494585
miRNA
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24/139 AceView cDNA sequences (see all 139):

AI830275 AI683609 AA514350 CA392349 BV201958 BG677534 AA278412 BV201965 
AA707896 BV201999 BV202473 BV185365 BV185364 CA406456 BV201963 AA579996 
CD722548 AA382059 BQ214648 AI784591 BV201982 BQ637147 BV202491 BM711336 

GeneLoc Exon Structure


(RNA expression data according to H-InvDB, NONCODE, miRBase, and RNAdb, Expression images according to data from BioGPS, Illumina Human BodyMap, and CGAP SAGE, Sets of similar genes according to GeneDecks, in vivo and in vitro expression data from LifeMap Discovery™, plus additional links to Genevestigator, and/or SOURCE, and/or BioGPS, and/or UniProtKB,
PCR Arrays from SABiosciences, Primers from OriGene, SABiosciences, and/or QIAGEN, In Situ Hybridization Assays from Advanced Cell Diagnostics)
About This Section

WASH4P expression in normal human tissues (normalized intensities)
See probesets specificity/sensitivity at GeneAnnot
About this imageBioGPS <intensity>2/3
CGAP TAG: --
WASH4P Expression
About this image


See WASH4P Protein Expression from SPIRE MOPED and PaxDB    SABiosciences Custom PCR Arrays for WASH4P
Primer
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In Situ
Assay Products:
 

 
Advanced Cell Diagnostics RNAscope RNA in situ hybridization assays for WASH4P

(Orthologs according to 1,2HomoloGene (2older version, for species not in 1newer version), 3euGenes, 4SGD , 5MGI Mar 06 2013, with possible further links to Flybase and/or WormBase, and/or 6Ensembl pan taxonomic compara , Gene Trees according to Ensembl and TreeFam)
About This Section

This gene was present in the common ancestor of animals.

Orthologs for WASH4P gene from 5/12 species (see all 12)    About this table
Organism Taxonomic
classification
Gene Description Human
Similarity
Orthology
Type
Details
chicken
(Gallus gallus)
Aves WASH16
WAS protein family homolog 1
68(a)
1 ↔ 1
1(60080916-60120121)
lizard
(Anolis carolinensis)
Reptilia WASH4P6
WAS protein family homolog 4 pseudogene
63(a)
1 ↔ 1
5(11324639-11363802)
zebrafish
(Danio rerio)
Actinopterygii wash6
WAS protein family homolog
57(a)
1 ↔ 1
18(5565808-5588950)
fruit fly
(Drosophila melanogaster)
Insecta wash6
washout
26(a)
1 ↔ 1
2R(8068145-8069878)
worm
(Caenorhabditis elegans)
Secernentea ddl-26
Protein DDL-2
20(a)
1 ↔ 1
II(13542835-13547629)
        Species with no ortholog for WASH4P

ENSEMBL Gene Tree for WASH4P (if available)
TreeFam Gene Tree for WASH4P (if available)

(Paralogs according to 1HomoloGene,
2Ensembl, and 3SIMAP, Pseudogenes according to 4Pseudogene.org Build 68)
About This Section
Paralogs for WASH4P gene
8 SIMAP similar genes for WASH4P using alignment to 4 protein entries:     WASH4_HUMAN (see all proteins):
WASH2P    WASH    WASH1    WASH3P    DKFZp686C24272    WASH6P
DKFZp434K1323    FLJ00075

WASH4P for paralogs           About GeneDecksing


5/9 Pseudogenes.org Pseudogenes for WASH4P (see all 9)
PGOHUM00000262196 PGOHUM00000262197 PGOHUM00000262200 PGOHUM00000262201 PGOHUM00000262264


(SNPs/Variants according to the 1NCBI SNP Database, 2Ensembl, 3PupaSUITE, 4UniProtKB, and DNA2.0, Linkage Disequilibrium by HapMap, Structural Variations(CNVs/InDels/Inversions) from the Database of Genomic Variants, Mutations from the Human Gene Mutation Database (HGMD) and the Locus Specific Mutation Databases (LSDB), Blood group antigen gene mutations by BGMUT, Resequencing Primers from QIAGEN, Cancer Mutation PCR Arrays and Assays and Copy Number PCR Arrays from SABiosciences)
About This Section
SABiosciences Cancer Mutation PCR Assays
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(in which this Gene is Involved, According to MalaCards, OMIM, UniProtKB, the University of Copenhagen DISEASES database, Conferences by KenesGroup, Genatlas, GeneTests, GAD, HuGE Navigator, and/or TGDB.)
About This Section
  --

(in PubMed. Associations of this gene to articles via 1Entrez Gene, 2UniProtKB/Swiss-Prot, 3HGNC, 4GAD, 5PharmGKB, 6HMDB, 7DrugBank, 8UniProtKB/TrEMBL, 9 Novoseek, and/or 10fRNAdb)
About This Section

PubMed articles for WASH4P gene integrated from 9 sources:
(articles sorted by number of sources associating them with WASH4P)
    Utopia: connect your pdf to the dynamic
world of online information

  1. Human subtelomeric WASH genes encode a new subclass of the WASP family. (PubMed id 18159949)1, 2, 3 Linardopoulou E.V.... Trask B.J. (2007)
  2. The sequence and analysis of duplication-rich human chromosome 16. (PubMed id 15616553)1, 2 Martin J.... Pennacchio L.A. (2004)
  3. Characterization of the murine orthologue of a novel human subtelomeric multigene family. (PubMed id 11701968)1, 3 Gianfrancesco F....D'Urso M. (2001)
  4. The relationship between chromosome structure and function at a human telomeric region. (PubMed id 9054936)1, 3 Flint J.... Higgs D.R. (1997)
  5. Toward a confocal subcellular atlas of the human proteome. (PubMed id 18029348)1 Barbe L....Andersson-Svahn H. (2008)
  6. Differentially regulated and evolved genes in the fully sequenced Xq/Yq pseudoautosomal region. (PubMed id 10655549)2 Ciccodicola A.... D'Urso M. (2000)

(in PubMed, OMIM, and NCBI Bookshelf)
About This Section
 ANDOR
Aliases
Free Text  

  Query String
PubMed
OMIM
NCBI Bookshelf
  (Note: In FireFox, select the above section and copy using Ctrl-C)

(According to Entrez Gene, HGNC, AceView, euGenes, Ensembl, miRBase, ECgene, Kegg, and/or H-InvDB)
About This Section
Entrez Gene: 374677 HGNC: 14126 AceView: CXYorf1P Ensembl:ENSG00000234769 euGenes: HUgn374677
ECgene: WASH4P H-InvDB: WASH4P

(According to HUGE)
About This Section
  --

(According to PharmGKB, ATLAS, HORDE, IMGT, LEIDEN, UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL,
Wikipedia and/or GeneReviews via UniProtKB/Swiss-Prot)
About This Section
  --

(Patent information from GeneIP,
Licensable technologies from WIS Yeda, Salk, Tufts,
IP news from LifeMap Sciences, Inc.)
About This Section
Patent Information for WASH4P gene:
Search GeneIP for patents involving WASH4P

GeneCards and IP:
Japan Patent Office Licenses GeneCards     European Patent Office Licenses GeneCards     Improving the IP Search



(Antibodies, recombinant proteins, and assays from EMD Millipore, R&D Systems, OriGene, QIAGEN, GenScript, Cell Signaling Technology, SABiosciences, Novus Biologicals, Sino Biological, Enzo Life Sciences, Abcam, ProSpec, Cloud-Clone Corp., Thermo Fisher Scientific, LSBio, Gene Editing from DNA2.0 and Sirion Biotech, Clones from EMD Millipore, OriGene, GenScript, Sino Biological, DNA2.0, SwitchGear Genomics, Vector BioLabs, Sirion Biotech, Cell lines from GenScript, and LifeMap BioReagents, PCR Arrays from SABiosciences, Drugs and/or compounds from EMD Millipore, Tocris Bioscience, and/or Enzo Life Sciences, In Situ Hybridization Assays from
Advanced Cell Diagnostics, Animal models from inGenious Targeting Laboratory, genOway)
About This Section

 
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 See all of Abcam's Antibodies, Kits and Proteins for WASH4P
 Custom Antibody / Protein Production Service
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 Abcam protocols and scientific support
 Browse ProSpec Recombinant Proteins




 Browse Proteins at Cloud-Clone Corp.
 Browse Antibodies at Cloud-Clone Corp.
 Browse ELISAs at Cloud-Clone Corp.
 Browse CLIAs at Cloud-Clone Corp.
 Search LifeMap BioReagents cell lines for WASH4P
 Gene Synthesis
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 Advanced Cell Diagnostics RNAscope RNA in situ hybridization assays for WASH4P
 Browse SwitchGear 3'UTR luciferase reporter plasmids for WASH4P
 Browse SwitchGear Promoter luciferase reporter plasmids for WASH4P
 Search ThermoFisher Antibodies for WASH4P
 Search Vector BioLabs for ready-to-use adenovirus/AAV for human, mouse, rat WASH4P
 inGenious Targeting Laboratory - Custom generated mouse model solutions for WASH4P
 inGenious Targeting Laboratory - Custom generated inducible mouse model solutions for WASH4P
Customized:
 lentivirus for stable overexpression of WASH4P
 lentivirus expression plasmids for stable overexpression of WASH4P
 adenovirus for overexpression of WASH4P
 Search LSBio for Antibodies for WASH4P
Customized transgenic rodents for:
 Humanization
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GeneCards Homepage - Last full update: 23 Oct 2013 - Incrementals: 4 Nov 2013 , 7 Nov 2013 , 23 Jan 2014

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(GIFtS: 73)
transforming growth factor, beta 1
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The GeneCards human gene database gene index: 1 3 5 6 A B C D E F G H I J K L M N O P Q R S T U V W X Y Z 


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