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WASH4P Gene

protein-coding   GIFtS: 21
GCID: GC16M000064

WAS Protein Family Homolog 4 Pseudogene

(Previous names: family with sequence similarity 39, member C pseudogene)
(Previous symbol: FAM39CP)
  Search for WASH4P
at  

(According to 1HGNC, 2Entrez Gene,
3UniProtKB/Swiss-Prot, 4UniProtKB/TrEMBL, 5OMIM, 6GeneLoc, 7Ensembl, 8DME, 9miRBase, 10fRNAdb, 12H-InvDB, 13NCBI, 14NONCODE, and/or 15RNAdb)
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This gene clusters with an RNA gene
Subcategory (RNA class): lncRNA

Quality score for the ORGUL clustered with this gene is 3

Aliases
WAS Protein Family Homolog 4 Pseudogene1 2
FAM39CP1 2 3
Family With Sequence Similarity 39, Member C Pseudogene1
CXYorf1P2
Protein FAM39CP3

External Ids:    HGNC: 141261   Entrez Gene: 3746772   Ensembl: ENSG000002347697   UniProtKB: A8MWX33   
ORGUL members:         

Export aliases for WASH4P gene to outside databases

Previous GC identifer: GC16M000008


(According to Entrez Gene, GeneCards, Tocris Bioscience, Wikipedia's Gene Wiki, PharmGKB,
UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL)
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GeneCards Summary for WASH4P Gene:
WASH4P (WAS protein family homolog 4 pseudogene) is a protein-coding gene.

UniProtKB/Swiss-Prot: WASH4_HUMAN, A8MWX3
Function: Acts as a nucleation-promoting factor at the surface of endosomes, where it recruits and activates the
Arp2/3 complex to induce actin polymerization, playing a key role in the fission of tubules that serve as
transport intermediates during endosome sorting (By similarity)




(According to GeneLoc and/or HGNC, and/or
Entrez Gene (NCBI build 37),
and/or miRBase,
Genomic Views according to UCSC (hg19) and Ensembl (release 75), Regulatory elements and Epigenetics data according to QIAGEN, and/or SwitchGear Genomics)
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Regulatory elements:
   Search for regulatory transcription factor binding sites for WASH4P
         Other transcription factors

Browse SwitchGear Promoter luciferase reporter plasmids
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Epigenetics:
DNA Methylation CpG Assay Predesigned for Pyrosequencing in human, mouse, rat WASH4P


Genomic Location:
Genomic View: UCSC Golden Path with GeneCards custom track

Entrez Gene cytogenetic band: 16p13.3   Ensembl cytogenetic band:  16p13.3   HGNC cytogenetic band: 16p13.3

WASH4P Gene in genomic location: bands according to Ensembl, locations according to (and/or Entrez Gene and/or Ensembl if different)
WASH4P gene location

GeneLoc information about chromosome 16         GeneLoc Exon Structure

GeneLoc location for GC16M000064:  view genomic region     (about GC identifiers)

Start:
64,043 bp from pter      End:
69,452 bp from pter
Size:
5,410 bases      Orientation:
minus strand

(According to 1UniProtKB, HORDE, 2neXtProt, Ensembl, and/or Reactome, Modification sites according to PhosphoSitePlus, Specific Peptides from DME, RefSeq according to NCBI, PDB rendering according to OCA and/or Proteopedia, Recombinant Proteins from EMD Millipore, R&D Systems, GenScript, Enzo Life Sciences, OriGene, Novus Biologicals, Sino Biological, ProSpec, and/or Cloud-Clone Corp.,
Biochemical Assays by EMD Millipore, R&D Systems, OriGene, GenScript, Cell Signaling Technology, Enzo Life Sciences, and/or Cloud-Clone Corp., Antibodies by EMD Millipore, R&D Systems, Cell Signaling Technology, OriGene, Novus Biologicals, Thermo Fisher Scientific, LSBio, Abcam, Cloud-Clone Corp, and/or others.)
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UniProtKB/Swiss-Prot: WASH4_HUMAN, A8MWX3 (See protein sequence)
Recommended Name: Putative WAS protein family homolog 4  
Size: 477 amino acids; 51595 Da
Subunit: Component of the WASH complex, composed of F-actin-capping protein subunit alpha (CAPZA1, CAPZA2 or
CAPZA3), F-actin-capping protein subunit beta (CAPZB), WASH (WASH1, WASH2P, WASH3P, WASH4P or WASH6P), FAM21
(FAM21A, FAM21B or FAM21C), KIAA1033, KIAA0196 and CCDC53. Interacts (via WHD1 region) with FAM21C; the
interaction is direct (By similarity)
Miscellaneous: WASH genes duplicated to multiple chromosomal ends during primate evolution, with highest copy
number reached in humans, whose WASH repertoires probably vary extensively among individuals (PubMed:18159949).
It is therefore difficult to determine which gene is functional or not. The telomeric region of chromosome 9p is
paralogous to the pericentromeric regions of chromosome 9 as well as to 2q. Paralogous regions contain 7
transcriptional units. Duplicated WASH genes are also present in the Xq/Yq pseudoautosomal region, as well as on
chromosome 1 and 15. The chromosome 16 copy seems to be a pseudogene
Caution: Could be the product of a pseudogene

Explore the universe of human proteins at neXtProt for WASH4P: NX_A8MWX3

Explore proteomics data for WASH4P at MOPED


See WASH4P Protein Expression from SPIRE MOPED, PaxDB, and MaxQB


ENSEMBL proteins: 
 ENSP00000317542   ENSP00000454316   ENSP00000457756   ENSP00000457950   ENSP00000441402  

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(According to HGNC, IUPHAR, InterPro, ProtoNet, UniProtKB, and/or BLOCKS, Sets of similar genes according to GeneDecks)
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HGNC Gene Families:
WASH: WAS protein homologs

2 InterPro protein domains:
 IPR028290 WASH1
 IPR021854 WASH1_WAHD

Graphical View of Domain Structure for InterPro Entry A8MWX3

ProtoNet protein and cluster: A8MWX3

UniProtKB/Swiss-Prot: WASH4_HUMAN, A8MWX3
Similarity: Belongs to the WASH1 family
Similarity: Contains 1 WH2 domain


WASH4P for domains           About GeneDecksing


(According to 1UniProtKB, Genatlas, LifeMap Discovery™, IUBMB, and/or 2DME, Human phenotypes from GenomeRNAi, Animal models from MGI Mar 06 2013, inGenious Targeting Laboratory, genOway,
transcription factor targeting from QIAGEN and/or HOMER, miRNA Gene Targets from miRTarBase, shRNA from OriGene, siRNAs from OriGene, QIAGEN, microRNA from QIAGEN, SwitchGear Genomics, Gene Editing from DNA2.0, Clones from OriGene, GenScript, Sino Biological, DNA2.0, and Vector BioLabs, Cell Lines from GenScript, ESI BIO, In Situ Hybridization Assays from Advanced Cell Diagnostics, Ontologies according to Gene Ontology Consortium 01 Apr 2014 via Entrez Gene.)
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Molecular Function:

     UniProtKB/Swiss-Prot Summary: WASH4_HUMAN, A8MWX3
Function: Acts as a nucleation-promoting factor at the surface of endosomes, where it recruits and activates the
Arp2/3 complex to induce actin polymerization, playing a key role in the fission of tubules that serve as
transport intermediates during endosome sorting (By similarity)

     Gene Ontology (GO): 2 molecular function terms:    About this table

GO IDQualified GO termEvidencePubMed IDs
GO:0003779actin binding ----
GO:0043014alpha-tubulin binding ----
     
WASH4P for ontologies           About GeneDecksing


Animal Models:
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(According to UniProtKB, COMPARTMENTS Subcellular localization database, Ontologies according to Gene Ontology Consortium 01 Apr 2014 via Entrez Gene.)
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Subcellular locations from UniProtKB/Swiss-Prot
WASH4_HUMAN, A8MWX3: Early endosome membrane (By similarity). Recycling endosome membrane (By similarity).
Note=Localization to the endodome membrane is mediated via its interaction with FAM21 (By similarity)
Subcellular locations from COMPARTMENTS: 

CompartmentConfidence
endosome4
mitochondrion2
cytosol1
nucleus1

Gene Ontology (GO): Selected cellular component terms (see all 6):    About this table

GO IDQualified GO termEvidencePubMed IDs
GO:0005769early endosome ----
GO:0031901early endosome membrane ----
GO:0043231intracellular membrane-bounded organelle ----
GO:0055037recycling endosome ----
GO:0055038recycling endosome membrane ----

WASH4P for ontologies           About GeneDecksing


(SuperPaths according to PathCards, Pathways according to R&D Systems, Cell Signaling Technology, KEGG, PharmGKB, BioSystems, Sino Biological, Reactome, Tocris Bioscience, GeneGo (Thomson Reuters), QIAGEN, and/or UniProtKB, Sets of similar genes according to GeneDecks, Interaction Networks according to QIAGEN, and/or STRING, Interactions according to 1UniProtKB, 2MINT, 3I2D, and/or 4STRING, with links to IntAct and Ensembl, Ontologies according to Gene Ontology Consortium 01 Apr 2014 via Entrez Gene).
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    Custom Pathway & Disease-focused RT2 Profiler PCR Arrays for WASH4P
Interactions:

    Search GeneGlobe Interaction Network for WASH4P

STRING Interaction Network Preview (showing 5 interactants - click image to see 12)

Selected Interacting proteins for WASH4P (ENSP000003175424) via UniProtKB, MINT, STRING, and/or I2D (see all 12)
InteractantInteraction Details
GeneCardExternal ID(s)
CAPZA1ENSP000002631684STRING: ENSP00000263168
FAM21AENSP000002826334STRING: ENSP00000282633
FAM21BENSP000003512594STRING: ENSP00000351259
FAM21CENSP000003375414STRING: ENSP00000337541
FAM21DENSP000003097864STRING: ENSP00000309786
About this table

Gene Ontology (GO): 3 biological process terms:    About this table

GO IDQualified GO termEvidencePubMed IDs
GO:0016197endosomal transport ----
GO:0034314Arp2/3 complex-mediated actin nucleation ----
GO:0042147retrograde transport, endosome to Golgi ----

WASH4P for ontologies           About GeneDecksing



(Chemical Compounds according to UniProtKB, Enzo Life Sciences, EMD Millipore, Tocris Bioscience, ApexBio, HMDB, BitterDB, and/or Novoseek, Ligands according to IUPHAR, and Drugs according to DrugBank, Enzo Life Sciences, and/or PharmGKB)
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Browse Tocris compounds for WASH4P (WASH4)



(Secondary structures according to fRNAdb,
GenBank/EMBL/DDBJ Accessions according to
Unigene (Build 236 Homo sapiens; Apr 25 2013) or GenBank,
RefSeq according to Entrez Gene,
DOTS (version 10), and/or AceView, transcript ids from Ensembl with links to UCSC,
exon structure from GeneLoc, alternative splicing isoforms according to ASD and/or ECgene,
siRNAs from OriGene, QIAGEN, shRNA from OriGene, microRNA from QIAGEN, SwitchGear Genomics,
Tagged/untagged cDNA clones from OriGene, GenScript, DNA2.0, Vector BioLabs, Primers from OriGene, and/or QIAGEN )
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13 Ensembl transcripts including schematic representations, and UCSC links where relevant:
ENST00000326592(uc002cfg.1) ENST00000474799 ENST00000470283 ENST00000468574
ENST00000564273 ENST00000477365 ENST00000495251 ENST00000462860 ENST00000470725
ENST00000487568 ENST00000483390 ENST00000538848 ENST00000494585
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Selected AceView cDNA sequences (see all 139):

AI830275 AI683609 AA514350 BM711336 BQ888995 BV202473 BI833485 CA406456 
BQ879313 BV201963 AI784591 BV185364 AA707896 BV185365 BG677534 BV201999 
AA278412 BV201958 CA392349 CD722548 BQ637147 BV201965 AA382059 AA579996 

GeneLoc Exon Structure


(RNA expression data according to H-InvDB, NONCODE, miRBase, and RNAdb, Expression images according to data from BioGPS, Illumina Human BodyMap, and CGAP SAGE, Sets of similar genes according to GeneDecks, in vivo and in vitro expression data from LifeMap Discovery™, Protein expression images according to data from SPIRE 1MOPED, 2PaxDb, and 3MaxQB, plus additional links to SOURCE, and/or BioGPS, and/or UniProtKB,
PCR Arrays from QIAGEN, Primers from OriGene, and/or QIAGEN, In Situ Hybridization Assays from Advanced Cell Diagnostics)
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WASH4P expression in normal human tissues (normalized intensities)
See probesets specificity/sensitivity at GeneAnnot
About this imageBioGPS <intensity>2/3
CGAP TAG: --
WASH4P Expression
About this image

WASH4P Protein expression data from MOPED1, PaxDb2 and MaxQB3    About this image

WASH4P Protein Expression
    Custom PCR Arrays for WASH4P
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In Situ
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(Orthologs according to 1,2HomoloGene (2older version, for species not in 1newer version), 3euGenes, 4SGD , 5MGI Mar 06 2013, with possible further links to Flybase and/or WormBase, and/or 6Ensembl pan taxonomic compara , Gene Trees according to Ensembl and TreeFam)
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This gene was present in the common ancestor of animals.

Orthologs for WASH4P gene from Selected species (see all 11)    About this table
Organism Taxonomic
classification
Gene Description Human
Similarity
Orthology
Type
Details
chicken
(Gallus gallus)
Aves WASH16
Gallus gallus WAS protein family homolog 1 (WASH1)...
69(a)
1 ↔ 1
1(60080916-60120121)
lizard
(Anolis carolinensis)
Reptilia WASH4P6
WAS protein family homolog 4 pseudogene
66(a)
1 ↔ 1
5(11324639-11363802)
zebrafish
(Danio rerio)
Actinopterygii wash6
WAS protein family homolog
59(a)
1 ↔ 1
18(5565808-5588950) ENSDARG00000063457
fruit fly
(Drosophila melanogaster)
Insecta wash6
washout
27(a)
1 ↔ 1
2R(8068145-8069878)
        Species with no ortholog for WASH4P

ENSEMBL Gene Tree for WASH4P (if available)
TreeFam Gene Tree for WASH4P (if available)

(Paralogs according to 1HomoloGene,
2Ensembl, and 3SIMAP, Pseudogenes according to 4Pseudogene.org Build 68)
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Paralogs for WASH4P gene
8 SIMAP similar genes for WASH4P using alignment to 4 protein entries:     WASH4_HUMAN (see all proteins):
WASH2P    WASH    WASH1    WASH3P    WASH6P    DKFZp686C24272
DKFZp434K1323    FLJ00075

WASH4P for paralogs           About GeneDecksing


Selected Pseudogenes.org Pseudogenes for WASH4P (see all 9)
PGOHUM00000262196 PGOHUM00000262197 PGOHUM00000262200 PGOHUM00000262201 PGOHUM00000262264


(SNPs/Variants according to the 1NCBI SNP Database, 2Ensembl, 3PupaSUITE, 4UniProtKB, and DNA2.0, Linkage Disequilibrium by HapMap, Structural Variations(CNVs/InDels/Inversions) from the Database of Genomic Variants, Mutations from the Human Gene Mutation Database (HGMD), the Human Cytochrome P450 Allele Nomenclature Database, and the Locus Specific Mutation Databases (LSDB), Blood group antigen gene mutations by BGMUT, Resequencing Primers, Cancer Mutation PCR Arrays and Assays, and Copy Number PCR Arrays from QIAGEN)
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Site Specific Mutation Identification with PCR Assays
Search QIAGEN SeqTarget long-range PCR primers for resequencing WASH4P
DNA2.0 Custom Variant and Variant Library Synthesis for WASH4P

(in which this Gene is Involved, According to MalaCards, OMIM, UniProtKB, the University of Copenhagen DISEASES database, Genatlas, GeneTests, GAD, HuGE Navigator, and/or TGDB.)
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  --

(in PubMed. Associations of this gene to articles via 1Entrez Gene, 2UniProtKB/Swiss-Prot, 3HGNC, 4GAD, 5PharmGKB, 6HMDB, 7DrugBank, 8UniProtKB/TrEMBL, 9 Novoseek, and/or 10fRNAdb)
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PubMed articles for WASH4P gene integrated from 10 sources:
(articles sorted by number of sources associating them with WASH4P)
    Utopia: connect your pdf to the dynamic
world of online information

  1. Human subtelomeric WASH genes encode a new subclass of the WASP family. (PubMed id 18159949)1, 2, 3 Linardopoulou E.V.... Trask B.J. (PLoS Genet. 2007)
  2. The sequence and analysis of duplication-rich human chromosome 16. (PubMed id 15616553)1, 2 Martin J.... Pennacchio L.A. (Nature 2004)
  3. Characterization of the murine orthologue of a novel human subtelomeric multigene family. (PubMed id 11701968)1, 3 Gianfrancesco F....D'Urso M. (Cytogenet. Cell Genet. 2001)
  4. The relationship between chromosome structure and function at a human telomeric region. (PubMed id 9054936)1, 3 Flint J.... Higgs D.R. (Nat. Genet. 1997)
  5. Toward a confocal subcellular atlas of the human proteome. (PubMed id 18029348)1 Barbe L....Andersson-Svahn H. (amp 2008)
  6. Differentially regulated and evolved genes in the fully sequenced Xq/Yq pseudoautosomal region. (PubMed id 10655549)2 Ciccodicola A.... D'Urso M. (Hum. Mol. Genet. 2000)

(in PubMed, OMIM, and NCBI Bookshelf)
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 ANDOR
Aliases
Free Text  

  Query String
PubMed
OMIM
NCBI Bookshelf
  (Note: In FireFox, select the above section and copy using Ctrl-C)

(According to Entrez Gene, HGNC, AceView, euGenes, Ensembl, miRBase, ECgene, Kegg, and/or H-InvDB)
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Entrez Gene: 374677 HGNC: 14126 AceView: CXYorf1P Ensembl:ENSG00000234769 euGenes: HUgn374677
ECgene: WASH4P H-InvDB: WASH4P

(According to HUGE)
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  --

(According to PharmGKB, ATLAS, HORDE, IMGT, LEIDEN, UniProtKB/Swiss-Prot, UniProtKB/TrEMBL, and/or others, e.g. Wikipedia and GeneReviews, via UniProtKB/Swiss-Prot)
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  --

(Patent information from GeneIP,
Licensable technologies from WIS Yeda, Salk, Tufts,
IP news from LifeMap Sciences, Inc.)
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Patent Information for WASH4P gene:
Search GeneIP for patents involving WASH4P

GeneCards and IP:
Japan Patent Office Licenses GeneCards     European Patent Office Licenses GeneCards     Improving the IP Search



(Antibodies, recombinant proteins, and assays from EMD Millipore, R&D Systems, OriGene, QIAGEN, GenScript, Cell Signaling Technology, Novus Biologicals, Sino Biological, Enzo Life Sciences, Abcam, ProSpec, Cloud-Clone Corp., Thermo Fisher Scientific, LSBio, Gene Editing from DNA2.0. Clones from OriGene, GenScript, Sino Biological, DNA2.0, SwitchGear Genomics, Vector BioLabs, Cell lines from GenScript, and ESI BIO, PCR Arrays from QIAGEN, Drugs and/or compounds from EMD Millipore, Tocris Bioscience, Enzo Life Sciences, and/or ApexBio, In Situ Hybridization Assays from
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 Browse SwitchGear Promoter luciferase reporter plasmids for WASH4P
 Search ThermoFisher Antibodies for WASH4P
 Search Vector BioLabs for ready-to-use adenovirus/AAV for human, mouse, rat WASH4P
 inGenious Targeting Laboratory: Let us create your new Knockout/Knockin mouse model for WASH4P
 inGenious Targeting Laboratory: Contact us about creating complex and humanized mouse models for WASH4P
 Search LSBio for Antibodies for WASH4P
Customized transgenic rodents for:
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GeneCards Homepage - Last full update: 7 May 2014 - Incrementals: 9 May 2014 , 2 Jun 2014 , 26 Jun 2014 , 30 Jun 2014

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