TTLL11 Gene
protein-coding GIFtS: 46
GCID: GC09M124584
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|
tubulin tyrosine ligase-like family, member 11(Previous name: chromosome 9 open reading frame 20 )
(Previous symbol: C9orf20)
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Aliases
for TTLL11 gene
(According to
1HGNC,
2Entrez Gene,
3UniProtKB/Swiss-Prot,
4UniProtKB/TrEMBL, 5OMIM, 6GeneLoc,
7Ensembl,
8DME,
9miRBase,
and/or 10fRNAdb)
About This Section
|
| Aliases |
|---|
| Tubulin Tyrosine Ligase-Like Family, Member 111 2 | | Chromosome 9 Open Reading Frame 201 | | C9orf201 2 3 | | Tubulin Polyglutamylase TTLL112 | | BA244O19.11 | | EC 6.-.-.-3 | | Tubulin--Tyrosine Ligase-Like Protein 112 3 | | |
Export aliases for TTLL11 gene to outside databasesPrevious GC identifers: GC09M121664 GC09M123624 GC09M123791 GC09M094322 |
Summaries
for TTLL11 gene(According to Entrez Gene,
Tocris Bioscience,
Wikipedia's
Gene Wiki,
PharmGKB,
UniProtKB/Swiss-Prot,
and/or
UniProtKB/TrEMBL)
About This Section
|
UniProtKB/Swiss-Prot: TTL11_HUMAN, Q8NHH1Function: Polyglutamase which preferentially modifies alpha-tubulin. Involved in the side-chain elongation step of thepolyglutamylation reaction rather than in the initiation step (By similarity). Required for CCSAP localization to bothspindle and cilia microtubules
|
Genomic Views
for TTLL11 gene
(According to
GeneLoc and/or
HGNC, and/or
Entrez Gene (NCBI build 37),
and/or miRBase,
Genomic Views according to
UCSC (hg19) and
Ensembl (release 69),
Regulatory elements and Epigenetics data according to
QIAGEN,
SABiosciences, and/or
SwitchGear Genomics)
About This Section
| RefSeq DNA sequence:- NC_000009.11 NC_018920.1 NT_008470.19
Regulatory elements:
 SABiosciences Regulatory transcription factor binding sites in the TTLL11 gene promoter:
GR Pbx1a p53 Meis-1b CP2 HOXA5 Meis-1a GR-alpha ATF6 Meis-1
Other transcription factors
Search SABiosciences Chromatin IP Primers for TTLL11
Epigenetics:
| | QIAGEN PyroMark CpG Assay predesigned Pyrosequencing DNA Methylation assays in human, mouse, rat TTLL11 |
Genomic Location:
Genomic View: UCSC Golden Path with GeneCards custom track
Entrez Gene cytogenetic band: 9q33.2 Ensembl cytogenetic band: 9q33.2 HGNC cytogenetic band: 9q34.11TTLL11 Gene in genomic location: bands according to Ensembl, locations according to
 (and/or Entrez Gene and/or Ensembl if different)
GeneLoc information about chromosome 9 GeneLoc Exon Structure GeneLoc location for GC09M124584: view genomic region
(about GC identifiers)
Start:
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124,584,204 bp from pter |
End:
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124,855,885 bp from pter |
Size:
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271,682 bases |
Orientation:
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minus strand |
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Proteins
for TTLL11 gene
(According to
1UniProtKB,
HORDE,
neXtProt,
Ensembl,
and/or Reactome,
Modification sites according to 2PhosphoSitePlus,
Specific Peptides from DME,
Protein expression images according to data from SPIRE MOPED and PaxDb,
RefSeq according to NCBI,
PDB rendering according to OCA and/or Proteopedia,
Recombinant Proteins
from
EMD Millipore,
R&D Systems,
GenScript,
Enzo Life Sciences,
OriGene,
Novus Biologicals,
Sino Biological,
ProSpec, and/or
Uscn,
Biochemical Assays by
EMD Millipore,
R&D Systems,
OriGene,
GenScript,
Cell Signaling Technology,
Enzo Life Sciences, and/or
Uscn,
Ontologies according to Gene
Ontology Consortium 01 Mar 2013 and
Entrez Gene,
Antibodies by
EMD Millipore,
R&D Systems,
GenScript,
Cell Signaling Technology,
OriGene,
Novus Biologicals,
Thermo Fisher Scientific,
Abcam, and/or
Uscn)
About This Section
|
UniProtKB/Swiss-Prot: TTL11_HUMAN, Q8NHH1 (See
protein sequence)Recommended Name: Tubulin polyglutamylase TTLL11 Size: 538 amino acids; 58035 Da
Subcellular location: Cytoplasm, cytoskeleton, cilium basal body (By similarity)Explore the universe of human proteins at neXtProt for TTLL11: NX_Q8NHH1
Post-translational modifications:
View modification sites using PhosphoSitePlus2View neXtProt modification sites for NX_Q8NHH1
TTLL11 Protein expression data from MOPED and PaxDb: About this image 
REFSEQ proteins (2 alternative transcripts):
NP_001132914.1 NP_919228.2
ENSEMBL proteins: ENSP00000362883 ENSP00000362881 ENSP00000321346
Human Recombinant Protein Products:
Gene Ontology (GO): 3 cellular component terms (GO ID links to tree view): About this table
TTLL11 for ontologies About GeneDecksing
TTLL11 Antibody Products:
Assay Products for TTLL11: |
Protein
Domains /
Families
for TTLL11 gene(According to InterPro, ProtoNet,
UniProtKB, and/or BLOCKS,
Sets of similar genes according to GeneDecks)
About This Section
|
TTLL11 for domains About GeneDecksing
1 InterPro domain/family:Graphical View of Domain Structure for InterPro Entry Q8NHH1ProtoNet protein and cluster: Q8NHH1 1 Blocks protein family: IPB004344 Tubulin-tyrosine ligase
UniProtKB/Swiss-Prot: TTL11_HUMAN, Q8NHH1Similarity: Belongs to the tubulin--tyrosine ligase familySimilarity: Contains 1 TTL domain |
Function
for TTLL11 gene
(According to 1UniProtKB,
Genatlas,
LifeMap Discovery™,
IUBMB, and/or
2DME,
Human phenotypes from GenomeRNAi,
Animal models from MGI Mar 06 2013,
bound targets from SABiosciences,
miRNA Gene Targets from miRTarBase
shRNA from
OriGene,
RNAi from
EMD Millipore,
siRNAs from
OriGene,
QIAGEN,
microRNA from QIAGEN,
Gene Editing from DNA2.0,
Clones from EMD Millipore,
OriGene,
SwitchGear Genomics,
GenScript,
Sino Biological,
DNA2.0,
and Vector BioLabs,
Cell Lines from GenScript,
LifeMap BioReagents,
In Situ Hybridization Assays from Advanced Cell Diagnostics,
Ontologies according to Gene Ontology Consortium 01 Mar 2013 via
Entrez Gene.)
About This Section
|
Function Summary:
UniProtKB/Swiss-Prot: TTL11_HUMAN, Q8NHH1Function: Polyglutamase which preferentially modifies alpha-tubulin. Involved in the side-chain elongation step of thepolyglutamylation reaction rather than in the initiation step (By similarity). Required for CCSAP localization to bothspindle and cilia microtubulesEnzyme Number (IUBMB): EC 6.-.-.-1
Clone
Products: |  | Browse Clones for the Expression of Recombinant Proteins Available from EMD Millipore | |  | OriGene Myc/DDK tagged cDNA clones in CMV expression vector in human, mouse, rat for TTLL11 (see all 4)
OriGene untagged cDNA clones in CMV expression vector in human, mouse, rat for TTLL11 (see all 2)
OriGene custom cloning services – gene synthesis, subcloning, mutagenesis,
variant library, vector shuttling 
| |  | GenScript: all cDNA clones in your preferred vector (see all 2): TTLL11 (NM_194252) | |  | Browse Sino Biological Human cDNA Clones | |  | DNA2.0 Custom Codon Optimized Gene
Synthesis Service for TTLL11 | |  | Search Vector BioLabs for ready-to-use adenovirus/AAV for human, mouse, rat TTLL11 |
In Situ Assay
Products: |
 | Advanced Cell Diagnostics RNAscope RNA in situ hybridization assays for TTLL11 |
Gene Ontology (GO): 1 molecular function term (GO ID links to tree view): About this table | GO ID | Qualified GO term | Evidence | PubMed IDs |
|---|
| GO:0004835 | tubulin-tyrosine ligase activity |
IEA | -- |
TTLL11 for ontologies About GeneDecksing
1 GenomeRNAi human phenotype for TTLL11:
|
Pathways & Interactions
for TTLL11 gene
(Pathways according to
EMD Millipore,
R&D Systems,
Cell Signaling Technology,
KEGG,
PharmGKB,
BioSystems,
Reactome,
Tocris Bioscience,
GeneGo (Thomson Reuters),
QIAGEN,
and/or UniProtKB,
Sets of similar genes according to GeneDecks,
Interaction Networks according to
SABiosciences,
and/or STRING,
Interactions according to 1UniProtKB,
2MINT,
3I2D, and/or
4STRING,
with links to IntAct and
Ensembl,
Ontologies according to Gene Ontology Consortium 01 Mar 2013 via
Entrez Gene).
About This Section
|
Interactions:
Search SABiosciences Gene Network CentralTM Interacting Genes and Proteins Networks for TTLL11
1 Interacting protein for TTLL11 (Q8NHH12) via UniProtKB, MINT, STRING, and/or I2DAbout this table
Gene Ontology (GO): 3 biological process terms (GO ID links to tree view): About this table | GO ID | Qualified GO term | Evidence | PubMed IDs |
|---|
| GO:0006464 | cellular protein modification process |
IEA | -- | | GO:0018095 | protein polyglutamylation |
-- | -- | | GO:0051013 | microtubule severing |
-- | -- |
TTLL11 for ontologies About GeneDecksing
|
Drugs & Compounds
for TTLL11 gene(Chemical Compounds according to UniProtKB, Enzo Life Sciences,
EMD Millipore, Tocris Bioscience
HMDB,
BitterDB, and/or
Novoseek, and Drugs according to
DrugBank,
Enzo Life Sciences, and/or
PharmGKB, with drugs/clinical trials/news
search links to CenterWatch)
About This Section
|
 Browse Tocris compounds for TTLL11
Search CenterWatch for drugs/clinical trials and news about TTLL11 / TTL11
|
Transcripts
for TTLL11 gene(Secondary structures according to
fRNAdb,
GenBank/EMBL/DDBJ Accessions according to
Unigene
(Build 235 Homo sapiens; Mar 10 2013) or GenBank,
RefSeq according to Entrez Gene,
DOTS (version 10), and/or
AceView,
transcript ids from Ensembl
with links to UCSC,
exon structure from GeneLoc,
alternative splicing isoforms according to ASD and/or
ECgene,
RNAi Products from
EMD Millipore,
siRNAs from
OriGene,
QIAGEN,
shRNA from
OriGene,
microRNA from QIAGEN,
Tagged/untagged cDNA clones from
OriGene,
SwitchGear Genomics,
GenScript,
DNA2.0,
Vector BioLabs,
Primers from
OriGene,
SABiosciences, and/or
QIAGEN
)
About This Section
|
REFSEQ mRNAs for TTLL11 gene (2 alternative transcripts): NM_001139442.1 NM_194252.2 Unigene Cluster for TTLL11: Tubulin tyrosine ligase-like family, member 11 Hs.640663 [show with all ESTs]Unigene Representative Sequence: NM_0011394426 Ensembl transcripts including schematic representations, and UCSC links where relevant: ENST00000486362 ENST00000373778 ENST00000474723 ENST00000373776(uc004blt.1 uc004blu.1)
ENST00000487468 ENST00000321582(uc011lym.1 uc011lyl.2 uc004blr.3)
Clone
Products: | | OriGene Myc/DDK tagged cDNA clones in CMV expression vector in human, mouse, rat for TTLL11 (see all 4)
OriGene untagged cDNA clones in CMV expression vector in human, mouse, rat for TTLL11 (see all 2)
OriGene custom cloning services – gene synthesis, subcloning, mutagenesis,
variant library, vector shuttling
| | | GenScript: all cDNA clones in your preferred vector (see all 2): TTLL11 (NM_194252) | | | DNA2.0 Custom Codon Optimized Gene
Synthesis Service for TTLL11 | | | Search Vector BioLabs for ready-to-use adenovirus/AAV for human, mouse, rat TTLL11 |
Additional cDNA sequence: AF521886.1 AF521887.1 BC089452.1 BC171751.1 4 DOTS entries: DT.99947626 DT.40265942 DT.208691 DT.99947228 24/36 AceView cDNA sequences (see all 36): AI741070 AF521886 AI571330 AW629275 AI150621 CF128404 AK125178 BM929722 AF521887 NM_194252 AA609430 BF439913 AA861826 BX092305 BX282469 BG771772 AI185724 AA676222 AW015518 BF108868 BF594279 AI769213 AI472951 BQ103840
GeneLoc Exon Structure
4 Alternative Splicing Database (ASD) splice patterns (SP) for TTLL11 About this scheme
| ExUns: | 1a | · | 1b | ^ | 2 | ^ | 3 | ^ | 4a | · | 4b | ^ | 5 | ^ | 6 | ^ | 7a | · | 7b | ^ | 8 | ^ | 9 | ^ | 10 | ^ | 11 |
|---|
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| SP1: | | | | | | | - | | | | | | | | | | - | | - | | - | | | | | | | | |
| SP2: | | | | | | | | | | | | | | | | | | | | | | | | | | | | | |
| SP3: | | | | | | | | | | | | | | | | | | | | | - | | | | | | | | |
| SP4: | | | | | | | | | | | | | | | | | | | | | | | | | | | | |
ECgene alternative splicing isoforms for TTLL11
|
Expression
for TTLL11 gene
(RNA expression data according to
H-InvDB,
NONCODE,
miRBase, and
RNAdb,
Expression images according to data from
BioGPS,
Illumina Human BodyMap, and
CGAP
SAGE,
Sets of similar genes according to GeneDecks,
in vivo and in vitro expression data from LifeMap Discovery™,
plus additional links to
Genevestigator, and/or
SOURCE, and/or
BioGPS, and/or
UniProtKB,
PCR Arrays from
SABiosciences,
Primers from
OriGene,
SABiosciences, and/or
QIAGEN,
In Situ Hybridization Assays from Advanced Cell Diagnostics)
About This Section
| TTLL11 expression in normal human tissues (normalized intensities)
See probesets specificity/sensitivity at GeneAnnot
About this imageBioGPS
CGAP TAG: --
About this image
See TTLL11 Protein Expression from SPIRE MOPED and PaxDB
Genevestigator expression for TTLL11
SOURCE GeneReport for Unigene cluster: Hs.640663
SABiosciences Custom PCR Arrays for TTLL11
Primer
Products: | | OriGene genome-wide validated SYBR primer pairs in human, mouse, rat for TTLL11
Browse OriGene validated miRNA SYBR primer pairs
| | | SABiosciences RT2 qPCR Primer Assay in human, mouse, rat TTLL11 | | | QIAGEN QuantiTect SYBR Green Assays in human, mouse, rat TTLL11 | | | QIAGEN QuantiFast Probe-based Assays in human, mouse, rat TTLL11 | In Situ
Assay Products: |
| Advanced Cell Diagnostics RNAscope RNA in situ hybridization assays for TTLL11 |
Orthologs
for TTLL11 gene
(Orthologs according to
1,2HomoloGene (2older version, for species not in 1newer version),
3euGenes,
4SGD
,
5MGI Mar 06 2013,
with possible further links to
Flybase
and/or
WormBase,
and/or
6Ensembl pan taxonomic compara ,
Gene Trees according to Ensembl and
TreeFam)
About This Section
|
This gene was present in the common ancestor of animals.
Orthologs for TTLL11 gene from 4/18 species (see all 18) About this table
ENSEMBL Gene Tree for TTLL11 (if available)
TreeFam Gene Tree for TTLL11 (if available) |
Paralogs
for TTLL11 gene(Paralogs according to
1HomoloGene,
2Ensembl, and 3SIMAP, Pseudogenes according to 4Pseudogene.org Build 68)
About This Section
| Paralogs for TTLL11 gene
- TTLL102 TTLL42 TTLL32 TTLL12 TTLL52 TTLL92 TTLL72 TTLL62
- TTL2 TTLL82 TTLL22 TTLL132 ARPC4-TTLL32
5 SIMAP similar genes for TTLL11 using alignment to 4 protein entries: TTL11_HUMAN (see all proteins):TTLL6 TTLL13 TTLL7 TTLL5 TTLL4
TTLL11 for paralogs About GeneDecksing
|
Genomic Variants
for TTLL11 gene(SNPs/Variants according to the
1NCBI SNP Database,
2Ensembl,
3PupaSUITE,
UniProtKB, and
DNA2.0,
Linkage Disequilibrium by HapMap,
Structural Variations(CNVs/InDels/Inversions) from the Database of Genomic Variants, Mutations from the Human Gene
Mutation Database (HGMD) and the Locus Specific Mutation
Databases (LSDB), Blood group antigen gene mutations by BGMUT,
Resequencing Primers from QIAGEN,
Cancer Mutation PCR Arrays and Assays and Copy Number PCR Arrays from SABiosciences)
About This Section
|
| Genomic Data | Transcription Related Data | Allele Frequencies | | SNP ID | Valid | Clinical
significance | Chr 9 pos | Sequence | # | AA
Chg | Type | More | # | Allele
freq | Pop | Total
sample | More |
|---|
HapMap Linkage Disequilibrium report for TTLL11 (124584204 - 124834204 bp, first 250kb of TTLL11)
Structural Variations (Copy Number Variations, Insertions/Deletions, Inversions)
Database of Genomic Variants (DGV): 9 variations for TTLL11
2 CNVs: 6713 59507
7 Indels: 42554 52704 47642 13199 29050 41680 41206
Human Gene Mutation Database (HGMD): TTLL11
| SABiosciences Cancer Mutation PCR Assays |
| | QIAGEN SeqTarget long-range PCR primers in human, mouse, rat for resequencing TTLL11 |
|
Disorders
/ Diseases
for TTLL11 gene
(in which this Gene is Involved, According to MalaCards,
OMIM, UniProtKB,
the University of Copenhagen DISEASES
database,
Genatlas, GeneTests,
GAD,
HuGE Navigator,
and/or TGDB.)
About This Section
|
TTLL11 for disorders About GeneDecksing
2  diseases for TTLL11: About MalaCardscleft lip/palate cleft lip
Human Genome Epidemiology (HuGE) Navigator: TTLL11 (2 documents)
Export disorders for TTLL11 gene to outside databases
|
Publications
for TTLL11 gene (in
PubMed.
Associations of this gene to articles via
1Entrez Gene,
2UniProtKB/Swiss-Prot,
3HGNC,
4GAD,
5PharmGKB,
6HMDB,
7DrugBank,
8UniProtKB/TrEMBL,
9 Novoseek, and/or
10fRNAdb)
About This Section
|
PubMed articles for TTLL11 gene integrated from 9 sources:
(articles sorted by number of sources associating them with TTLL11) | |  | Utopia: connect your pdf to the dynamic
world of online information |
- Tubulin polyglutamylase enzymes are members of the TTL domain protein family. (PubMed id 15890843)1, 3 Janke C.... Edde B. (2005)
- DNA sequence and analysis of human chromosome 9. (PubMed id 15164053)1, 2 Humphray S.J.... Dunham I. (2004)
- CSAP localizes to polyglutamylated microtubules and promotes proper cilia function and zebrafish development. (PubMed id 22493317)2 Backer C.B.... Cheeseman I.M. (2012)
- Personalized smoking cessation: interactions between nicotine dose, dependence and quit-success genotype score. (PubMed id 20379614)1 Rose J.E....Uhl G.R. (2010)
- Follow-up association studies of chromosome region 9q and nonsyndromic cleft lip/palate. (PubMed id 20583170)1 Letra A....Vieira A.R. (2010)
- Complete sequencing and characterization of 21,243 full-length human cDNAs. (PubMed id 14702039)1 Ota T.... Sugano S. (2004)
- Generation and initial analysis of more than 15,000 full-length human and mouse cDNA sequences. (PubMed id 12477932)1 Strausberg R.L....Marra M.A. (2002)
- Hepatitis C virus core protein interacts with 14-3-3 protein and activates the kinase Raf-1. (PubMed id )2 Aoki H.... Hino O. (2000)
- Shotgun sequencing of the human transcriptome with ORF expressed sequence tags. (PubMed id 10737800)1 Dias Neto E....Simpson A.J.G. (2000)
|
External Searches for TTLL11 gene
(in PubMed,
OMIM, and NCBI Bookshelf)
About This Section
|
|
Genome Databases showing TTLL11 gene
(According to
Entrez Gene,
HGNC,
AceView,
euGenes,
Ensembl,
miRBase,
ECgene,
Kegg,
and/or
H-InvDB)
About This Section
|
|
Other Databases showing TTLL11 gene
(According to HUGE)
About This Section
| -- |
Specialized Databases showing TTLL11 gene(According to PharmGKB,
ATLAS, HORDE, IMGT, LEIDEN, UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL,
Wikipedia and/or GeneReviews via UniProtKB/Swiss-Prot)
About This Section
|
| Name | Description |
| PharmGKB entry for TTLL11 | Pharmacogenomics, SNPs, Pathways |
|
| | |
About This Section
| Patent Information for TTLL11 gene:
Search GeneIP for patents involving TTLL11
GeneCards and IP:
Japan Patent Office Licenses GeneCards European Patent Office Licenses GeneCards Improving the IP Search
|
Products
for TTLL11 gene(Antibodies, recombinant proteins, and assays from EMD Millipore, R&D Systems, OriGene, QIAGEN, GenScript, Cell Signaling Technology, SABiosciences, Novus Biologicals, Sino Biological, Enzo Life Sciences, Abcam, ProSpec, Uscn, Thermo Fisher Scientific, Gene Editing from DNA2.0, Clones from EMD Millipore, OriGene, GenScript, Sino Biological, DNA2.0, SwitchGear Genomics, Vector BioLabs, Cell lines from GenScript and LifeMap BioReagents, PCR Arrays from SABiosciences, Drugs and/or compounds from EMD Millipore, Tocris Bioscience, and/or
Enzo Life Sciences),
In Situ Hybridization Assays from
Advanced Cell Diagnostics
About This Section
|
 | |
 | |
 |
| | |  | Browse OriGene Antibodies | | OriGene shRNA RFP for TTLL11 | | | OriGene 29mer shRNA kits in GFP-retroviral vector in human, mouse, rat for TTLL11 | | OriGene genome-wide validated SYBR primer pairs in human, mouse, rat for TTLL11 | | | OriGene Protein Over-expression Lysate for TTLL11 | | Browse OriGene Fluorogenic Cell Assay Kits | | | OriGene siRNA for TTLL11 | | OriGene 3'-UTR Clone for TTLL11 | | | OriGene untagged cDNA clones in CMV expression vector in human, mouse, rat for TTLL11 | | OriGene Myc/DDK tagged cDNA clones in CMV expression vector in human, mouse, rat for TTLL11 | | | Browse OriGene GFP tagged cDNA clones in CMV expression vector | | Browse OriGene MicroRNA Expression Plasmids | | | Browse OriGene basic RS shRNAs | | Browse OriGene validated miRNA SYBR primer pairs | | | Browse OriGene full length recombinant human proteins expressed in human HEK293 cells | | OriGene custom cloning services - gene synthesis, subcloning, mutagenesis,
variant library, vector shuttling | | | OriGene Custom Antibody Services for TTLL11 | | OriGene Custom Protein Services for TTLL11 | | | OriGene Custom Immunoassay Development | | |
| |
 |
 |
| | | QIAGEN Custom miScript Target Protector blocks miRNA-binding site of in human, mouse, rat TTLL11 | | | QIAGEN SeqTarget long-range PCR primers in human, mouse, rat for resequencing TTLL11 | | | QIAGEN PyroMark CpG Assay predesigned Pyrosequencing DNA Methylation assays in human, mouse, rat TTLL11 | | | QIAGEN FlexiTube/FlexiPlate siRNA for gene silencing in human, mouse, rat TTLL11 | | | QIAGEN QuantiFast Probe-based Assays in human, mouse, rat TTLL11 | | | QIAGEN QuantiTect SYBR Green Assays in human, mouse, rat TTLL11 |
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 | | | TTLL11 Proteins, Antibodies, CLIAs, and ELISAs |
| | | | | Advanced Cell Diagnostics RNAscope RNA in situ hybridization assays for TTLL11 |
|  |  |  | | | | ThermoFisher Antibodies for TTLL11 |
| | | Search Vector BioLabs for ready-to-use adenovirus/AAV for human, mouse, rat TTLL11 |
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