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Category   Symbol Source: HGNC EntrezGene Ensembl GeneCards RNA genes CroW21
GIFtS

TRAM1L1 Gene

protein-coding   GIFtS: 44
GCID: GC04M118004

translocation associated membrane protein 1-like 1
 Explore 3 diseases affiliated with
TRAM1L1 via
MalaCards
our new
 Human Malady Compendium 
Biological research products
for TRAM1L1    

Aliases
for TRAM1L1 gene

(According to 1HGNC, 2Entrez Gene,
3UniProtKB/Swiss-Prot, 4UniProtKB/TrEMBL, 5OMIM, 6GeneLoc, 7Ensembl, 8DME, 9miRBase, and/or 10fRNAdb)
About This Section
Aliases
Translocation Associated Membrane Protein 1-Like 11 2
MGC265681
Translocating Chain-Associated Membrane Protein 1-Like 12

External Ids:    HGNC: 283711   Entrez Gene: 1330222   Ensembl: ENSG000001745997   UniProtKB: Q8N6093   

Export aliases for TRAM1L1 gene to outside databases

Previous GC identifers: GC04M118463 GC04M118362 GC04M118224 GC04M113738


Summaries
for TRAM1L1 gene

(According to Entrez Gene, Tocris Bioscience, Wikipedia's Gene Wiki, PharmGKB,
UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL)
About This Section
UniProtKB/Swiss-Prot: TR1L1_HUMAN, Q8N609
Function: Stimulatory or required for the translocation of secretory proteins across the ER membrane (By similarity)




Genomic Views
for TRAM1L1 gene

(According to GeneLoc and/or HGNC, and/or
Entrez Gene (NCBI build 37),
and/or miRBase,
Genomic Views according to UCSC (hg19) and Ensembl (release 69), Regulatory elements and Epigenetics data according to QIAGEN, SABiosciences, and/or SwitchGear Genomics)
About This Section
RefSeq DNA sequence:
NC_000004.11  NC_018915.1  NT_016354.19  
Regulatory elements:
   SABiosciences Regulatory transcription factor binding sites in the TRAM1L1 gene promoter:
         TBP   STAT1   p53   STAT1beta   POU3F2   STAT1alpha   S8   FAC1   HFH-1   IRF-7A   
         Other transcription factors

SwitchGear Promoter luciferase reporter plasmid: TRAM1L1 promoter sequence
   Search SABiosciences Chromatin IP Primers for TRAM1L1

Epigenetics:
QIAGEN PyroMark CpG Assay predesigned Pyrosequencing DNA Methylation assays in human, mouse, rat TRAM1L1


Genomic Location:
Genomic View: UCSC Golden Path with GeneCards custom track

Entrez Gene cytogenetic band: 4q26   Ensembl cytogenetic band:  4q26   HGNC cytogenetic band: 4q26

TRAM1L1 Gene in genomic location: bands according to Ensembl, locations according to (and/or Entrez Gene and/or Ensembl if different)
TRAM1L1 gene location

GeneLoc information about chromosome 4         GeneLoc Exon Structure

GeneLoc location for GC04M118004:  view genomic region     (about GC identifiers)

Start:
 118,004,710 bp from pter      End:
 118,006,736 bp from pter
Size:
 2,027 bases      Orientation:
 minus strand

Proteins
for TRAM1L1 gene

(According to 1UniProtKB, HORDE, neXtProt, Ensembl, and/or Reactome, Modification sites according to 2PhosphoSitePlus, Specific Peptides from DME, Protein expression images according to data from SPIRE MOPED and PaxDb, RefSeq according to NCBI, PDB rendering according to OCA and/or Proteopedia, Recombinant Proteins from EMD Millipore, R&D Systems, GenScript, Enzo Life Sciences, OriGene, Novus Biologicals, Sino Biological, ProSpec, and/or Uscn,
Biochemical Assays by EMD Millipore, R&D Systems, OriGene, GenScript, Cell Signaling Technology, Enzo Life Sciences, and/or Uscn, Ontologies according to Gene Ontology Consortium 01 Mar 2013 and Entrez Gene, Antibodies by EMD Millipore, R&D Systems, GenScript, Cell Signaling Technology, OriGene, Novus Biologicals, Thermo Fisher Scientific, Abcam, and/or Uscn)
About This Section
UniProtKB/Swiss-Prot: TR1L1_HUMAN, Q8N609 (See protein sequence)
Recommended Name: Translocating chain-associated membrane protein 1-like 1  
Size: 369 amino acids; 42162 Da
Subcellular location: Endoplasmic reticulum membrane; Multi-pass membrane protein (By similarity)
Secondary accessions: Q8N2L7

Explore the universe of human proteins at neXtProt for TRAM1L1: NX_Q8N609

Post-translational modifications:

  • View modification sites using PhosphoSitePlus2
  • View neXtProt modification sites for NX_Q8N609

    • TRAM1L1 Protein expression data from MOPED and PaxDb:    About this image 
    Estimated protein expression log10 (pmol).

    REFSEQ proteins: NP_689615.2  
    ENSEMBL proteins: 
     ENSP00000309402  

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    Uscn Proteins for TRAM1L1

    Gene Ontology (GO): 2 cellular component terms (GO ID links to tree view):    About this table

    GO IDQualified GO termEvidencePubMed IDs
    GO:0005789endoplasmic reticulum membrane IEA--
    GO:0016021integral to membrane IEA--


    TRAM1L1 for ontologies           About GeneDecksing



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    Protein Domains /
    Families
    for TRAM1L1 gene

    (According to InterPro, ProtoNet, UniProtKB, and/or BLOCKS, Sets of similar genes according to GeneDecks)
    About This Section
    TRAM1L1 for domains           About GeneDecksing

    3 InterPro domains/families:
     IPR016447 Translocation_assoc_membrane
     IPR013599 TRAM1
     IPR006634 TLC-dom

    Graphical View of Domain Structure for InterPro Entry Q8N609

    ProtoNet protein and cluster: Q8N609

    4 Blocks protein families:
    IPB005547 Longevity-assurance protein (LAG1)
    IPB006634 TRAM
    IPB011075 Tetracyclin repressor-like
    IPB013599 TRAM1-like protein


    UniProtKB/Swiss-Prot: TR1L1_HUMAN, Q8N609
    Similarity: Belongs to the TRAM family
    Similarity: Contains 1 TLC (TRAM/LAG1/CLN8) domain


    Function
    for TRAM1L1 gene

    (According to 1UniProtKB, Genatlas, LifeMap Discovery™, IUBMB, and/or 2DME, Human phenotypes from GenomeRNAi, Animal models from MGI Mar 06 2013,
    bound targets from SABiosciences, miRNA Gene Targets from miRTarBase shRNA from OriGene, RNAi from EMD Millipore, siRNAs from OriGene, QIAGEN, microRNA from QIAGEN, Gene Editing from DNA2.0, Clones from EMD Millipore, OriGene, SwitchGear Genomics, GenScript, Sino Biological, DNA2.0, and Vector BioLabs, Cell Lines from GenScript, LifeMap BioReagents, In Situ Hybridization Assays from Advanced Cell Diagnostics, Ontologies according to Gene Ontology Consortium 01 Mar 2013 via Entrez Gene.)
    About This Section
    Function Summary:

         UniProtKB/Swiss-Prot: TR1L1_HUMAN, Q8N609
    Function: Stimulatory or required for the translocation of secretory proteins across the ER membrane (By similarity)

    miRNA
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    Inhib. RNA
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    Gene Editing
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    1 GenomeRNAi human phenotype for TRAM1L1:
     Decreased Hepatitis C virus re 


    Pathways & Interactions
    for TRAM1L1 gene

    (Pathways according to EMD Millipore, R&D Systems, Cell Signaling Technology, KEGG, PharmGKB, BioSystems, Reactome, Tocris Bioscience, GeneGo (Thomson Reuters), QIAGEN, and/or UniProtKB, Sets of similar genes according to GeneDecks, Interaction Networks according to SABiosciences, and/or STRING, Interactions according to 1UniProtKB, 2MINT, 3I2D, and/or 4STRING, with links to IntAct and Ensembl, Ontologies according to Gene Ontology Consortium 01 Mar 2013 via Entrez Gene).
    About This Section



    Interactions:

        Search SABiosciences Gene Network CentralTM Interacting Genes and Proteins Networks for TRAM1L1

    Gene Ontology (GO): 1 biological process term (GO ID links to tree view):    About this table

    GO IDQualified GO termEvidencePubMed IDs
    GO:0015031protein transport IEA--


    TRAM1L1 for ontologies           About GeneDecksing



    Drugs & Compounds
    for TRAM1L1 gene

    (Chemical Compounds according to UniProtKB, Enzo Life Sciences, EMD Millipore, Tocris Bioscience HMDB, BitterDB, and/or Novoseek, and Drugs according to DrugBank, Enzo Life Sciences, and/or PharmGKB, with drugs/clinical trials/news search links to CenterWatch)
    About This Section
    Browse Small Molecules at EMD Millipore
    Browse drugs & compounds from Enzo Life Sciences

    Browse Tocris compounds for TRAM1L1
    Search CenterWatch for drugs/clinical trials and news about TRAM1L1 / TR1L1 

    Transcripts
    for TRAM1L1 gene

    (Secondary structures according to fRNAdb,
    GenBank/EMBL/DDBJ Accessions according to
    Unigene (Build 235 Homo sapiens; Mar 10 2013) or GenBank,
    RefSeq according to Entrez Gene,
    DOTS (version 10), and/or AceView, transcript ids from Ensembl with links to UCSC,
    exon structure from GeneLoc, alternative splicing isoforms according to ASD and/or ECgene,
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    About This Section
    REFSEQ mRNAs for TRAM1L1 gene: 
    NM_152402.2  

    Unigene Cluster for TRAM1L1:

    Translocation associated membrane protein 1-like 1
    Hs.570737  [show with all ESTs]
    Unigene Representative Sequence: BC030831
    1 Ensembl transcript including schematic representation, and UCSC links where relevant:
    ENST00000310754(uc003ibv.4)

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    hsa-miR-3675-3p hsa-miR-216b hsa-miR-660
    SwitchGear 3'UTR luciferase reporter plasmid: TRAM1L1 3' UTR sequence
    Inhib. RNA
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    OriGene 29mer shRNA kits in GFP-retroviral vector in human, mouse, rat for TRAM1L1 (see all 7)
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    Clone
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    Primer
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    Additional cDNA sequence: 

    AK074617.1 BC030831.1 

    1 DOTS entry:

    DT.40255174 

    20 AceView cDNA sequences:

    AK074617 BX097238 AV725026 BX423039 AI138349 CR596564 BM663681 AW078666 
    CA415689 NM_152402 BC030831 BG714759 AA169554 AW611506 CD679598 AW023347 
    AI335840 BE938743 BE938727 BE938736 

    GeneLoc Exon Structure


    Expression
    for TRAM1L1 gene

    (RNA expression data according to H-InvDB, NONCODE, miRBase, and RNAdb, Expression images according to data from BioGPS, Illumina Human BodyMap, and CGAP SAGE, Sets of similar genes according to GeneDecks, in vivo and in vitro expression data from LifeMap Discovery™, plus additional links to Genevestigator, and/or SOURCE, and/or BioGPS, and/or UniProtKB,
    PCR Arrays from SABiosciences, Primers from OriGene, SABiosciences, and/or QIAGEN, In Situ Hybridization Assays from Advanced Cell Diagnostics)
    About This Section

    TRAM1L1 expression in normal human tissues (normalized intensities)
    See probesets specificity/sensitivity at GeneAnnot
    About this imageBioGPS
    CGAP TAG: --

    Microarray
    RNAseq (Illumina Body Map)
    (100×FPKM)½
    SAGE (Serial Analysis of Gene Expression)
    About this image

    TRAM1L1 expression in embryonic tissues and stem cells
    Expression by the Database of Embryonic development, Stem cell research, and Regenerative medicine    About this table
    2 LifeMap In Vivo Development Anatomical Compartments/Cells 
    Tissue Anatomical Compartment CellCategory (developmental path)
    EyeInner Nuclear LayerMature Horizontal CellsHorizontal, Retina
    EyeInner Nuclear LayerMature Rod Bipolar CellsBipolar, Retina
    Expression: Positive    Negative     Selective marker
    Experimental details: Curated     Microarrays     In-situ hybridization

    See TRAM1L1 Protein Expression from SPIRE MOPED and PaxDB
    Genevestigator expression for TRAM1L1

    SOURCE GeneReport for Unigene cluster: Hs.570737
        SABiosciences Custom PCR Arrays for TRAM1L1
    Primer
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    In Situ
    Assay Products:     
         		Advanced Cell Diagnostics RNAscope RNA in situ hybridization assays for TRAM1L1

    Orthologs
    for TRAM1L1 gene

    (Orthologs according to 1,2HomoloGene (2older version, for species not in 1newer version), 3euGenes, 4SGD , 5MGI Mar 06 2013, with possible further links to Flybase and/or WormBase, and/or 6Ensembl pan taxonomic compara , Gene Trees according to Ensembl and TreeFam)
    About This Section
    This gene was present in the common ancestor of animals.

    Orthologs for TRAM1L1 gene from 6/18 species (see all 18)    About this table
    Organism Taxonomic
    classification    		 Gene Description Human
    Similarity    		 Orthology
    Type    		 Details
    mouse
    (Mus musculus)    		 Mammalia Tram1l11 , 5 translocation associated membrane protein 1-like 11, 5 77.16(n)1
    76.24(a)1    		   3 (54.68 cM)5
    2298011  NM_146140.31  NP_666252.11 
     1243210375 
    chicken
    (Gallus gallus)    		 Aves TRAM16
    		 translocation associated membrane protein 1
    		 59(a)
    		 1 → many
    		 2(121363156-121381698)
    lizard
    (Anolis carolinensis)    		 Reptilia --
    		 --
    		 65(a)
    		 1 → many
    		 4(32376180-32408683)
    zebrafish
    (Danio rerio)    		 Actinopterygii tram16
    		 translocating chain-associating membrane protein 1...
    		 57(a)
    		 1 → many
    		 24(14118964-14138957)
    fruit fly
    (Drosophila melanogaster)    		 Insecta TRAM6
    		 TRAM
    		 35(a)
    		 1 → many
    		 X(829265-831855)
    worm
    (Caenorhabditis elegans)    		 Secernentea tram-16
    		 TRAM (TRanslocating chain-Associating Membrane) pr...
    		 29(a)
    		 1 → many
    		 IV(10219409-10221260)


    ENSEMBL Gene Tree for TRAM1L1 (if available)
    TreeFam Gene Tree for TRAM1L1 (if available) 

    Paralogs
    for TRAM1L1 gene

    (Paralogs according to 1HomoloGene,
    2Ensembl, and 3SIMAP, Pseudogenes according to 4Pseudogene.org Build 68)
    About This Section
    Paralogs for TRAM1L1 gene
    TRAM12  TRAM22  
    2 SIMAP similar genes for TRAM1L1 using alignment to 1 protein entry:     TR1L1_HUMAN:
    TRAM1    TRAM2

    TRAM1L1 for paralogs           About GeneDecksing



    Genomic Variants
    for TRAM1L1 gene

    (SNPs/Variants according to the 1NCBI SNP Database, 2Ensembl, 3PupaSUITE, UniProtKB, and DNA2.0, Linkage Disequilibrium by HapMap, Structural Variations(CNVs/InDels/Inversions) from the Database of Genomic Variants, Mutations from the Human Gene Mutation Database (HGMD) and the Locus Specific Mutation Databases (LSDB), Blood group antigen gene mutations by BGMUT, Resequencing Primers from QIAGEN, Cancer Mutation PCR Arrays and Assays and Copy Number PCR Arrays from SABiosciences)
    About This Section
    SABiosciences Cancer Mutation PCR Assays
    QIAGEN SeqTarget long-range PCR primers in human, mouse, rat for resequencing TRAM1L1
    DNA2.0 Custom Variant and Variant Library Synthesis for TRAM1L1

    Disorders / Diseases
    for TRAM1L1 gene

    (in which this Gene is Involved, According to MalaCards, OMIM, UniProtKB, the University of Copenhagen DISEASES database, Genatlas, GeneTests, GAD, HuGE Navigator, and/or TGDB.)
    About This Section
    TRAM1L1 for disorders           About GeneDecksing

    3 diseases for TRAM1L1:    About MalaCards
    alcohol dependence    alcoholism    obesity

    Human Genome Epidemiology (HuGE) Navigator: TRAM1L1 (1 document)

    Export disorders for TRAM1L1 gene to outside databases

    Publications
    for TRAM1L1 gene

    (in PubMed. Associations of this gene to articles via 1Entrez Gene, 2UniProtKB/Swiss-Prot, 3HGNC, 4GAD, 5PharmGKB, 6HMDB, 7DrugBank, 8UniProtKB/TrEMBL, 9 Novoseek, and/or 10fRNAdb)
    About This Section

    PubMed articles for TRAM1L1 gene integrated from 9 sources:
    (articles sorted by number of sources associating them with TRAM1L1)    		    Utopia: connect your pdf to the dynamic
    world of online information

    1. Signal sequence and keyword trap in silico for selection of full- length human cDNAs encoding secretion or membrane proteins from oligo- capped cDNA libraries. (PubMed id 16303743)1, 2 Otsuki T....Isogai T. (2005)
    2. Generation and initial analysis of more than 15,000 full-length human and mouse cDNA sequences. (PubMed id 12477932)1, 3 Strausberg R.L....Marra M.A. (2002)
    3. Genome-wide association for abdominal subcutaneous and visceral adipose reveals a novel locus for visceral fat in women. (PubMed id 22589738)1 Fox C.S....Borecki I.B. (2012)
    4. A systematic gene-based screen of chr4q22-q32 identif ies association of a novel susceptibility gene, DKK2, with the quantitative tra it of alcohol dependence symptom counts. (PubMed id 20332099)1 Kalsi G....Riley B.P. (2010)
    5. Common body mass index-associated variants confer ris k of extreme obesity. (PubMed id 19553259)1 Cotsapas C....Daly M.J. (2009)
    6. Diversification of transcriptional modulation: large-scale identification and characterization of putative alternative promoters of human genes. (PubMed id 16344560)1 Kimura K.... Sugano S. (2006)
    7. TRAM2 protein interacts with endoplasmic reticulum Ca2+ pump Serca2b and is necessary for collagen type I synthesis. (PubMed id 14749390)1 Stefanovic B....Brenner D.A. (2004)
    8. The status, quality, and expansion of the NIH full-length cDNA project: the Mammalian Gene Collection (MGC). (PubMed id 15489334)2 Gerhard D.S....Malek J. (2004)

    External Searches for TRAM1L1 gene

    (in PubMed, OMIM, and NCBI Bookshelf)
    About This Section
     ANDOR
    Aliases
    Free Text  
      Query String
    		PubMed
    OMIM
    NCBI Bookshelf
      (Note: In FireFox, select the above section and copy using Ctrl-C)

    Genome Databases showing TRAM1L1 gene

    (According to Entrez Gene, HGNC, AceView, euGenes, Ensembl, miRBase, ECgene, Kegg, and/or H-InvDB)
    About This Section
    Entrez Gene: 133022 HGNC: 28371 AceView: TRAM1L1 Ensembl:ENSG00000174599 euGenes: HUgn133022
    ECgene: TRAM1L1 H-InvDB: TRAM1L1

    Other Databases showing TRAM1L1 gene

    (According to HUGE)
    About This Section
    		   --

    Specialized Databases showing TRAM1L1 gene

    (According to PharmGKB, ATLAS, HORDE, IMGT, LEIDEN, UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL,
    Wikipedia and/or GeneReviews via UniProtKB/Swiss-Prot)
    About This Section
    NameDescription
    PharmGKB entry for TRAM1L1 Pharmacogenomics, SNPs, Pathways
    ATLAS Chromosomes in Cancer entry for TRAM1L1 Genetics and Cytogenetics in Oncology and Haematology

    Intellectual Property
    for TRAM1L1 gene

    (Patent information from GeneIP,
    Licensable technologies from WIS Yeda, Salk, Tufts,
    IP news from LifeMap Sciences, Inc.)
    About This Section
    		Patent Information for TRAM1L1 gene:
    Search GeneIP for patents involving TRAM1L1

    GeneCards and IP:
    Japan Patent Office Licenses GeneCards     European Patent Office Licenses GeneCards     Improving the IP Search



    Products
    for TRAM1L1 gene

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    About This Section

     
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