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TIMP4 Gene

protein-coding GIFtS: 60

GC03M012169

TIMP metallopeptidase inhibitor 4
(Previous name: tissue inhibitor of metalloproteinase 4 )
Symbol approved by the HUGO Gene Nomenclature Committee (HGNC) database

(According to ¹HGNC, ²Entrez Gene,
³UniProtKB/Swiss-Prot, ⁴UniProtKB/TrEMBL, ⁵OMIM, ⁶GeneLoc , and/or ⁷Ensembl, ⁸miRBase)
About This Section

Aliases
TIMP-4 ^³

Descriptions
TIMP metallopeptidase inhibitor 4 ^²
Tissue inhibitor of metalloproteinases 4 ^³
tissue inhibitor of metalloproteinase 4 ^¹, ²

External Ids
HGNC: 11823¹
Entrez Gene: 7079²
UniProtKB: Q99727³
Ensembl: ENSG00000157150⁷

Search outside databases for aliases for TIMP4 gene

Previous GC identifers: GC03M012123 GC03M012038

(According to Entrez Gene, Wikipedia's Gene Wiki,
UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL)
About This Section

EntrezGene summary for TIMP4:

This gene belongs to the TIMP gene family. The proteins encoded by this gene family are inhibitors

of the matrix metalloproteinases, a group of peptidases involved in degradation of the

extracellular matrix. The secreted, netrin domain-containing protein encoded by this gene is

involved in regulation of platelet aggregation and recruitment and may play role in hormonal

regulation and endometrial tissue remodeling. [provided by RefSeq]

UniProtKB/Swiss-Prot: TIMP4_HUMAN, Q99727

Function: Complexes with metalloproteinases (such as collagenases) and irreversibly inactivates

them. Known to act on MMP-1, MMP-2, MMP-3, MMP-7 and MMP-9

Gene Wiki entry for TIMP4

(According to GeneLoc and/or HGNC, and/or
Entrez Gene (NCBI build 36),
and/or miRBase,
Genomic Views according to UCSC and Ensembl, Transcription factor binding sites according to SABiosciences)
About This Section

Genomic View:

UCSC Golden Path with GeneCards custom track

Transcription factor binding sites upstream to the TIMP4 gene

Entrez Gene cytogenetic band: 3p25 Ensembl cytogenetic band: 3p25.2 HGNC cytogenetic band: 3p25

TIMP4 Gene in genomic location: bands according to Ensembl, locations according to (and/or Entrez Gene and/or Ensembl if different)

GeneLoc gene densities for chromosome 3 GeneLoc Exon Structure

GeneLoc location for GC03M012169: (about GC identifiers)

Start:	12,169,568 bp from pter
End:	12,175,851 bp from pter
Size:	6,284 bases
Orientation:	minus strand

RefSeq DNA sequence:

NC_000003.10 NT_022517.17

(According to ¹UniProtKB, and/or Ensembl, Phosphorylation sites according to ²Phosphosite, RefSeq according to NCBI, PDB rendering according to OCA and/or Proteopedia, Recombinant Proteins from Invitrogen, Millipore, Sigma-Aldrich, R&D Systems, Enzo Life Sciences, Abnova, OriGene and/or, Abcam,
Biochemical Assays by Invitrogen, Millipore, R&D Systems, Cell Signaling Technology, and/or Enzo Life Sciences, Ontologies according to Gene Ontology Consortium 01 Apr 2009 and Entrez Gene, Antibodies by Invitrogen, Millipore, Sigma-Aldrich, R&D Systems, Cell Signaling Technology, Abcam, Abnova, and/or Novus Biologicals)
About This Section

UniProtKB/Swiss-Prot: TIMP4_HUMAN, Q99727 (See protein sequence)

Recommended Name: Metalloproteinase inhibitor 4 precursor

Size: 224 amino acids; 25503 Da

Subcellular location: Secreted

REFSEQ proteins: NP_003247.1

ENSEMBL proteins:

ENSP00000287814

Human Recombinant Proteins

	Browse Drug Discovery Central at Invitrogen for human recombinant proteins
	Millipore Purified and/or Recombinant TIMP4 Protein
	Browse Human Recombinant Proteins at Sigma-Aldrich
	R&D Systems Recombinant & Natural Proteins for TIMP4 (TIMP-4)
	Enzo Life Sciences proteins for TIMP4
	Recombinant Proteins from Abcam (TIMP4)
	Human Recombinant Proteins from Abnova (TIMP4)

OriGene Purified Recombinant Human Protein: TIMP4

2 Gene Ontology (GO) cellular component terms (links to tree view):

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0005576	extracellular region	NAS	--
GO:0005578	proteinaceous extracellular matrix	IEA	--

About this table

Antibodies for TIMP4:

	Browse Antibodies Central at Invitrogen
	Millipore Mono- and Polyclonal Antibodies for the study of TIMP4
	Sigma-Aldrich Antibodies for TIMP4
	R&D Systems Antibodies for TIMP4 (TIMP-4)
	Antibodies from Abcam (TIMP4), each with their Abpromise^SM
	Monoclonal and Polyclonal Antibodies from Abnova (TIMP4)
	Novus Biologicals Antibodies for TIMP4

Assays for TIMP4:

	Browse Invitrogen for biochemical assays
	Browse Kits and Assays available from Millipore
	R&D Systems ELISAs for TIMP4 (TIMP-4) (see all)
	Browse biochemical assays available from Enzo Life Sciences

(According to InterPro, ProtoNet, UniProtKB, and/or BLOCKS, Sets of similar genes according to GeneDecks)
About This Section

Graphical View of Domain Structure for InterPro Entry Q99727

ProtoNet protein and cluster: Q99727

1 Blocks protein family: IPB001820 Tissue inhibitor of metalloproteinase

UniProtKB/Swiss-Prot: TIMP4_HUMAN, Q99727

Similarity: Belongs to the protease inhibitor I35 (TIMP) family

Similarity: Contains 1 NTR domain

(According to MGI Jun 06 2009, UniProtKB, IUBMB,and/or Genatlas,
shRNA from OriGene, Sigma-Aldrich, RNAi from Sigma-Aldrich,
RNAi Products, Clones, and Q-PCR Products from Invitrogen, Millipore, OriGene, and/or Abnova, siRNAs from Applied Biosystems, SYBR primers from OriGene, Cell-based Assays from Millipore, Ontologies according to Gene Ontology Consortium 01 Apr 2009 via Entrez Gene.)
About This Section

Inhib. RNA:		Invitrogen RNAi Products for gene knock-down (TIMP4)
		Browse for Gene Knock-down Tools from Millipore
		Abnova Chimera RNAi Products for Gene knock-down (TIMP4)

OriGene 29mer shRNA kit in GFP-retroviral vector: NM_003256

Applied Biosystems Silencer® siRNAs for TIMP4

Sigma-Aldrich siRNA for TIMP4

Sigma-Aldrich shRNA Panels and shRNA for TIMP4

Explore Sigma-Aldrich super-pooled esiRNAs

Clones:		Invitrogen Clones for TIMP4
		Browse Clones for the Expression of Recombinant Proteins Available from Millipore

OriGene GFP tagged cDNA clone in CMV expression vector: NM_003256
Myc/DDK tagged cDNA clone in CMV expression vector: NM_003256
untagged cDNA clone in CMV expression vector: NM_003256

Primers:

Browse Quantitative PCR Central at Invitrogen for Q-PCR LUX™ Primers

OriGene genome-wide validated SYBR primer pairs: NM_003256

UniProtKB/Swiss-Prot: TIMP4_HUMAN, Q99727

Function: Complexes with metalloproteinases (such as collagenases) and irreversibly inactivates

them. Known to act on MMP-1, MMP-2, MMP-3, MMP-7 and MMP-9

Genatlas biochemistry entry for TIMP4:

matrix metalloproteinase,tissue inhibitor 4,abundantly expressed only in adult heart maybe embedded

in intron V of SYN2

1 MGI mutant phenotype (inferred from 1 allele) (MGI details for Timp4):

no phenotypic analysis

3 Gene Ontology (GO) molecular function terms (links to tree view):

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0004857	enzyme inhibitor activity	IEA	--
GO:0005515	protein binding	IEA	--
GO:0008191	metalloendopeptidase inhibitor activity	NAS	8939999

About this table

(Pathways according to Invitrogen (maps by GeneGo), Millipore, Cell Signaling Technology, Sigma-Aldrich, KEGG and/or UniProtKB,
Sets of similar genes according to GeneDecks, Proteins Network according to SABiosciences, Interactions according to ¹UniProtKB, ²MINT, and/or ³STRING, with links to IntAct and Ensembl, Ontologies according to Gene Ontology Consortium 01 Apr 2009 via Entrez Gene.)
About This Section

1 Sigma-Aldrich "Your Favorite Gene" Pathway for TIMP4 (Your Favorite Gene powered by Ingenuity)

Leukocyte Extravasation Signaling

GeneDecks TIMP4 for the pathways selected above

About GeneDecksing

Gene Network Central^TM Interacting Genes and Proteins Network for TIMP4

5/19 Interacting proteins for TIMP4 (ENSP00000287814³) via UniProtKB, MINT, and/or STRING (see all 19 )

Interactant		Interaction Details
GeneCard	External ID(s)	Interaction Details
MMP2	ENSP00000219070³	STRING (score=.99)
MMP14	ENSP00000308208³	STRING (score=.965)
MMP15	ENSP00000219271³	STRING (score=.959)
HPX	ENSP00000265983³	STRING (score=.879)
MMP9	ENSP00000361405³	STRING (score=.866)

About this table

1 Gene Ontology (GO) biological process term (links to tree view):

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0008150	biological_process	ND	--

About this table

(Chemical Compounds according to UniProtKB, Enzo Life Sciences, Sigma-Aldrich, Tocris Bioscience, and/or Novoseek and Drugs according to Enzo Life Sciences and/or PharmGKB)
About This Section

Browse drugs & compounds from Enzo Life Sciences

Browse Small Molecules at Sigma-Aldrich

Browse Tocris compounds for TIMP4
2 Novoseek chemical compound relationships for TIMP4 gene

Compound	Score	Articles	PubMed IDs for Articles with Shared Sentences (# sentences)
hydroxamate	54.19	1	11415441 (1)
1,10-phenanthroline	27.28	2	11415441 (1), 15013428 (1)

About this table

(GenBank/EMBL/DDBJ Accessions according to
Unigene (Build 219 Homo sapiens; Jun 2 2009) or GenBank,
RefSeq according to Entrez Gene,
DOTS (version 10), and/or AceView,
non coding RNAs according to RNAdb,
ESTs according to GeneTide,
exon structure from GeneLoc, alternative splicing isoforms according to ASD and/or ECgene,
RNAi Products from Invitrogen, Millipore, and/or Abnova,
siRNAs from Applied Biosystems, Sigma-Aldrich,
shRNA from Sigma-Aldrich, OriGene,
Tagged/untagged cDNA clones from OriGene,
Expression Assays from Applied Biosystems)
About This Section

Inhib. RNA:		Invitrogen RNAi Products for gene knock-down (TIMP4)
		Browse for Gene Knock-down Tools from Millipore
		Abnova Chimera RNAi Products for Gene knock-down (TIMP4)

OriGene 29mer shRNA kit in GFP-retroviral vector: NM_003256

Sigma-Aldrich siRNA for TIMP4

Sigma-Aldrich shRNA Panels and shRNA for TIMP4

Explore Sigma-Aldrich super-pooled esiRNAs

Applied Biosystems Silencer® siRNAs:

NM_003256

REFSEQ mRNAs for TIMP4 gene:

NM_003256.2

Applied Biosystems TaqMan ® Gene Expression Assays:

NM_003256

              OriGene GFP tagged cDNA clone in CMV expression vector: NM_003256
                                 Myc/DDK tagged cDNA clone in CMV expression vector: NM_003256
                                 untagged cDNA clone in CMV expression vector: NM_003256

Additional cDNA sequence:

AK313018.1 BC010553.1 BT019627.1 U76456.1

1 DOTS entry:

DT.308421

24/50 AceView cDNA sequences (see all 50 ):

BU732949 AW963378 BM759178 BC010553 BU615814 BM707859 AW292470 NM_003256

CD245370 BP381456 AI637967 CD518333 BX103242 CA424098 AW293304 U76456

AW196583 CB265444 BM129238 CB995893 AI359841 BM740043 BG910254 BE788008

highest scoring ESTs for TIMP4:

U76456 AA193414 BC010553 BF346229 BG819996 BG910254 BI752578 CB142663 CB265444 CB992438

Unigene Cluster for TIMP4:

TIMP metallopeptidase inhibitor 4

Hs.591665 [show with all ESTs]

Unigene Representative Sequence: BC010553

GeneLoc Exon Structure

1 Ensembl transcript including schematic representation:

ENST00000287814

(Experimental results according to ¹GeneNote and GNF BioGPS,
probe sets-to-genes annotations according to ²GeneAnnot , ³GeneTide , Sets of similar genes according to GeneDecks, Electronic Northern calculations according to data from UniGene (Build 219 Homo sapiens), SAGE tags according to CGAP, plus additional links to SOURCE, and/or GNF BioGPS, and/or EXPOLDB, and/or UniProtKB,
Expression Assays from Applied Biosystems )
About This Section

TIMP4 expression in normal and diseased human tissues

Applied Biosystems TaqMan ® Gene Expression Assays for TIMP4

¹ / ² / ³

4 probe-sets matching TIMP4 gene

Affymetrix probe-set	Array	# genes	Sensitivity	Specificity	Correlation	Length	Gb_Accession	Consensus	Uniqueness	Score	Rank
Affymetrix probe-set	Array	GeneAnnot data			GeneNote data		GeneTide data
819_at²^{, 3}	U95-A	1	1.00	1.00	0.99	0.82	U76456	1.00	1.00	1.00	1
51691_at²^{, 3}	U95-B	1	1.00	1.00	0.97	1.20	AA193414	0.80	1.00	0.91	1
206243_at²^{, 3}	U133-A	1	1.00	1.00	--	--	NM_003256	0.60	1.00	0.82	1
206243_at²	U133Plus2	1	1.00	1.00	--	--	--	--	--	--	--

GeneDecks TIMP4 for binary patterns associated with the probe-sets selected above

About GeneDecksing

About this table

Data from

(Publications) and GNF BioGPS
About these images
About these images

CGAP SAGE TAG: AGATGGTATA

SOURCE GeneReport for Unigene cluster: Hs.591665
Expression variation in blood from EXPOLDB for TIMP4
UniProtKB/Swiss-Prot: TIMP4_HUMAN, Q99727
Tissue specificity: Abundant in heart and present at low levels in many other tissues

(Orthologs according to ^1,2HomoloGene (²older version, for species not in ¹newer version), ³euGenes, ⁴SGD and/or ⁵MGI Jun 06 2009, with possible further links to Flybase and/or WormBase, Gene Trees according to Ensembl)
About This Section

Orthologs for TIMP4 gene from 5 species

Organism	Gene	Locus	Description	Human Similarity	NCBI accessions
chimpanzee (Pan troglodytes)	TIMP4¹	--	TIMP metallopeptidase inhibitor 4	98.96(n) 99.11(a)	460176 XM_516284.2 XP_516284.1
cow (Bos taurus)	TIMP4¹	--	TIMP metallopeptidase inhibitor 4	89.43(n) 90.18(a)	317694 NM_001045871.1 NP_001039336.1
rat (Rattus norvegicus)	Timp4¹	--	tissue inhibitor of metalloproteinase 4	87.8(n) 89.73(a)	680130 XM_001056766.1 XP_001056766.1
mouse (Mus musculus)	Timp4^{1, 5}	6 (46.00 cM)⁵	tissue inhibitor of metalloproteinase 4^{1, 5}	87.65(n)¹ 88.84(a)¹	110595¹ NM_080639.3¹ NP_542370.3¹ AF282730⁵ AF345863⁵ (see all 21)
chicken (Gallus gallus)	TIMP4¹	--	TIMP metallopeptidase inhibitor 4	74.69(n) 76.89(a)	768711 XM_001232058.1 XP_001232059.1

About this table Species with no ortholog for TIMP4

ENSEMBL Gene Tree for TIMP4

(Paralogs according to ¹HomoloGene
and ²Ensembl, Pseudogenes according to ³Pseudogene.org)
About This Section

Paralogs for TIMP4 gene

TIMP3² TIMP1² TIMP2²

(According to the ¹NCBI SNP Database, ²Ensembl, ³PupaSUITE, and UniProtKB, Linkage Disequilibrium by HapMap, Genotyping Reagents from Applied Biosystems)
About This Section

(in which this Gene is Involved, According to OMIM, UniProtKB, Novoseek, PharmGKB, Genatlas, GeneTests, Blood group antigen gene mutations by BGMUT, HGMD, GAD, HuGE Navigator, BCGD, and/or TGDB.)
About This Section

OMIM: 601915

10/14 Novoseek disease relationships for TIMP4 gene (see all 14 )

Disease	Score	Articles	PubMed IDs for Articles with Shared Sentences (# sentences)
neovascularization	23.56	1	9791730 (1)
metastasis	22.17	4	12828172 (2), 15523699 (1)
cancer	13.72	9	17873896 (3), 15928670 (1), 9190892 (1), 18336787 (1) (see all 7)
arthritis	13.38	4	12483743 (3)
tumors	12.99	19	12798711 (3), 17873896 (2), 15713681 (2), 17009974 (2) (see all 11)
breast cancer	12.82	14	12828172 (3), 11289097 (3), 11178975 (2), 9190892 (1) (see all 5)
adenocarcinoma	11.13	3	17873896 (1), 12798711 (1), 11501766 (1)
necrosis	8.60	2	15713681 (2)
hyperplasia	0.00	1	14744773 (1)
inflammation	0.00	3	19036126 (1), 16521002 (1)

About this table

Human Genome Epidemiology Navigator: TIMP4 (4 documents)

(Possibly Related Articles in Doctor's Guide)
About This Section

(in PubMed. Associations of this gene to articles via ¹Novoseek, ²HGNC, ³Entrez Gene, ⁴UniProtKB/Swiss-Prot, ⁵UniProtKB/TrEMBL, ⁶GAD, and/or ⁷PharmGKB)
About This Section

10/106 PubMed articles for TIMP4 gene (see all 106 ):

Cloning of the human tissue inhibitor of metalloproteinase-4 gene (TIMP4) and localization of the TIMP4 and Timp4 genes to human chromosome 3p25 and mouse chromosome 6, respectively. (PubMed id 9693046)^{1, 2, 3, 4} Olson T.M....Apte S.S. (1998)
Molecular cloning and characterization of human tissue inhibitor of metalloproteinase 4. (PubMed id 8939999)^{1, 2, 3, 4} Greene J.... Shi Y.E. (1996)
The status, quality, and expansion of the NIH full-length cDNA project: the Mammalian Gene Collection (MGC). (PubMed id 15489334)^{3, 4} Gerhard D.S....Malek J. (2004)
Specific, high affinity binding of tissue inhibitor of metalloproteinases-4 (TIMP-4) to the COOH-terminal hemopexin-like domain of human gelatinase A. TIMP-4 binds progelatinase A and the COOH-terminal domain in a similar manner to TIMP-2. (PubMed id 9182583)^{1, 3} Bigg H.F....Overall C.M. (1997)
Endometrial TIMP-4 mRNA is expressed in the stroma, while TIMP-4 protein accumulates in the epithelium and is released to the uterine fluid. (PubMed id 16809379)^{1, 3} Pilka R....Casslen B. (2006)
Tissue inhibitor of metalloproteinases 4 (TIMP4) is involved in inflammatory processes of human cardiovascular pathology. (PubMed id 16521002)^{1, 3} Koskivirta I....Jarvelainen H. (2006)
Identification, regulation and role of tissue inhibitor of metalloproteinases-4 (TIMP-4) in human platelets. (PubMed id 12466243)^{1, 3} Radomski A....Radomski M.W. (2002)
Imbalance of tissue inhibitors of metalloproteinases (TIMP) - 1 and - 4 serum levels, in patients with inflammatory bowel disease. (PubMed id 19036126)^{1, 3} Kapsoritakis A.N....Potamianos S.P. (2008)
E. coli expression of TIMP-4 and comparative kinetic studies with TIMP-1 and TIMP-2: insights into the interactions of TIMPs and matrix metalloproteinase 2 (gelatinase A). (PubMed id 12475252)^{1, 3} Troeberg L....Nagase H. (2002)
Promoter Polymorphism (rs3755724, -55C/T) of Tissue Inhibitor of Metalloproteinase 4 (TIMP4) as a Risk Factor for Kawasaki Disease with Coronary Artery Lesions in a Korean Population. (PubMed id 19048177)^{1, 3} Ban J.Y....Chung J.H. (2008)

(in PubMed, OMIM, and NCBI Bookshelf)
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		AND	OR
Aliases
Free Text

Query String		PubMed OMIM NCBI Bookshelf
	(Note: In FireFox, select the above section and copy using Ctrl-C)

(According to Entrez Gene, HGNC, AceView, euGenes, Ensembl, miRBase, ECgene, and/or H-InvDB)
About This Section

Entrez Gene: 7079	HGNC: 11823	AceView: TIMP4	Ensembl:ENSG00000157150	euGenes: HUgn7079
ECgene: TIMP4	H-InvDB: TIMP4

(According to HUGE)
About This Section

(According to ATLAS, HORDE, IMGT, MTDB, LEIDEN, UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL,
Wikipedia and/or GeneReviews via UniProtKB/Swiss-Prot)
About This Section

Name	Description
ATLAS Chromosomes in Cancer entry for TIMP4	Genetics and Cytogenetics in Oncology and Haematology

Licensable Technologies for TIMP4
(Available from WIS Yeda, Salk, Tufts)

About This Section

(Reagents available from Applied Biosystems, Antibodies and assays by Cell Signaling Technology, Abcam, Novus Biologicals,
Sigma-Aldrich, R&D Systems, Millipore, Abnova, and/or Invitrogen, Clones available from OriGene,and/or Invitrogen, Drugs and/or compounds by Sigma-Aldrich,
Enzo Life Sciences, and/or Tocris Bioscience)
About This Section

Products for TIMP4:

TaqMan ® Gene Expression Assays

TaqMan ® Genotyping Assays

Free SNP selection tool

Invitrogen iPath Pathways	Invitrogen BLOCK-iT™ RNAi
Invitrogen Antibodies	Invitrogen Assays
Invitrogen Clones	Invitrogen Q-PCR Products
Invitrogen Human Recombinant Kinases	Invitrogen Custom Antibody and Peptide Service
Invitrogen Proteins / Assays / Screening Services	Search Invitrogen catalog for TIMP4-related products

	Millipore Custom Antibody & Bulk Services
	Millipore Preclinical / Clinical Development Services
	Millipore Immunoassay Services
	Millipore Target Screening & Profiling Services

	Predesigned and custom siRNAs for TIMP4		Antibodies for TIMP4
	Explore super-pooled esiRNAs		Browse proteins at Sigma-Aldrich
	Lentivirus-delivered shRNAs for TIMP4		Browse small molecules at Sigma-Aldrich
	"Your Favorite Gene" Pathways		feedback

	Antibodies		Primer Pairs
	Cell Culture Products		ELISAs
	Flow Cytometry Kits		Protease Activity Assays & Reagents
	Cell Selection/Detection Kits & Reagents		ELISA/Assay Development Kits & Reagents
	Multiplex/Array Assay Kits & Reagents		ELISpot Kits & Development Modules
	Recombinant & Natural Proteins

	Recombinant Proteins (TIMP4)
	Antibodies (TIMP4)
	Chimera RNAi (TIMP4)
	Custom Service for Mouse Mab
	Custom Service for Rabbit Pab from Full-length Protein

Search for Antibodies & Assays

	Recombinant Proteins (TIMP4)
	Antibodies (TIMP4)

	Tagged/untagged cDNA clones
	Validated SYBR primer pairs
	Purified proteins from HEK293T cells
	Western blot validated antibodies
	shRNA in GFP-retroviral vector

Search Tocris compounds for TIMP4

	Proteins for TIMP4
	Search www.enzolifesicences.com for proteins, assays, substrates, inhibitors & antibodies

Antibodies for TIMP4

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TIMP4 Gene

protein-coding GIFtS: 60

Aliases &
Descriptions
for TIMP4

Summaries
for TIMP4

Genomic Location
for TIMP4

Proteins
for TIMP4

Protein Domains/
Families
for TIMP4

Gene Function
for TIMP4

Pathways & Interactions
for TIMP4

Drugs & Compounds
for TIMP4

Transcripts
for TIMP4

Expression
for TIMP4

Orthologs
for TIMP4

Paralogs
for TIMP4

SNPs/Variants
for TIMP4

Disorders & Mutations
for TIMP4

Medical News
for TIMP4

Publications
for TIMP4

Search for TIMP4

Genome Databases showing TIMP4

Other Databases showing TIMP4

Specialized Databases showing TIMP4

Licensable Technologies
for TIMP4

Services
for TIMP4

Gene Index: 3 5 A B C D E F G H I J K L M N O P Q R S T U V W X Y Z

TIMP4 Gene

protein-coding GIFtS: 60

Aliases & Descriptionsfor TIMP4

Summariesfor TIMP4

Genomic Locationfor TIMP4

Proteinsfor TIMP4

Protein Domains/Familiesfor TIMP4

Gene Functionfor TIMP4

Pathways & Interactionsfor TIMP4

Drugs & Compoundsfor TIMP4

Transcriptsfor TIMP4

Expression for TIMP4

Orthologsfor TIMP4

Paralogsfor TIMP4

SNPs/Variantsfor TIMP4

Disorders & Mutationsfor TIMP4

Medical Newsfor TIMP4

Publicationsfor TIMP4

Search for TIMP4

Genome Databases showing TIMP4

Other Databases showing TIMP4

Specialized Databases showing TIMP4

Licensable Technologiesfor TIMP4

Servicesfor TIMP4

Gene Index: 3 5 A B C D E F G H I J K L M N O P Q R S T U V W X Y Z

Aliases &
Descriptions
for TIMP4

Summaries
for TIMP4

Genomic Location
for TIMP4

Proteins
for TIMP4

Protein Domains/
Families
for TIMP4

Gene Function
for TIMP4

Pathways & Interactions
for TIMP4

Drugs & Compounds
for TIMP4

Transcripts
for TIMP4

Expression
for TIMP4

Orthologs
for TIMP4

Paralogs
for TIMP4

SNPs/Variants
for TIMP4

Disorders & Mutations
for TIMP4

Medical News
for TIMP4

Publications
for TIMP4

Licensable Technologies
for TIMP4

Services
for TIMP4