SYPL1 Gene
protein-coding GIFtS: 50
GCID: GC07M105730
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synaptophysin-like 1(Previous name: synaptophysin-like protein ) (Previous symbol: SYPL)
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Aliases for SYPL1 gene
(According to
1HGNC,
2Entrez Gene,
3UniProtKB/Swiss-Prot,
4UniProtKB/TrEMBL, 5OMIM, 6GeneLoc,
7Ensembl,
8DME,
9miRBase,
and/or 10fRNAdb) About This Section
|
| Aliases |
|---|
| Synaptophysin-Like 11 2 | | Pantophysin3 | | SYPL1 2 3 | | Synaptophysin-Like Protein 12 | | Synaptophysin-Like Protein1 | | Pantophysin3 | | H-SP12 | | |
Export aliases for SYPL1 gene to outside databasesPrevious GC identifers: GC07M105325 GC07M105518 GC07M100090 |
Summaries for SYPL1 gene(According to Entrez Gene,
Tocris Bioscience,
Wikipedia's
Gene Wiki,
PharmGKB,
UniProtKB/Swiss-Prot,
and/or
UniProtKB/TrEMBL)
About This Section
| Gene Wiki entry for SYPL1
|
Genomic Views for SYPL1 gene
(According to
GeneLoc and/or
HGNC, and/or
Entrez Gene (NCBI build 37),
and/or miRBase,
Genomic Views according to
UCSC (hg19) and
Ensembl (release 69),
Regulatory elements and Epigenetics data according to
QIAGEN,
SABiosciences, and/or
SwitchGear Genomics) About This Section
| RefSeq DNA sequence:- NC_000007.13 NC_018918.1 NT_007933.15 NT_079596.2
Regulatory elements: SABiosciences Regulatory transcription factor binding sites in the SYPL1 gene promoter: Nkx3-1 Nkx3-1 v2 Nkx3-1 v4 Nkx3-1 v3 Nkx3-1 v1 STAT5A Other transcription factors
Search SABiosciences Chromatin IP Primers for SYPL1
Epigenetics:
|  | QIAGEN PyroMark CpG Assay predesigned Pyrosequencing DNA Methylation assays in human, mouse, rat SYPL1 |
Genomic Location: Genomic View: UCSC Golden Path with GeneCards custom track
Entrez Gene cytogenetic band: 7q22.3 Ensembl cytogenetic band: 7q22.3 HGNC cytogenetic band: 7q22.2SYPL1 Gene in genomic location: bands according to Ensembl, locations according to
(and/or Entrez Gene and/or Ensembl if different)

GeneLoc information about chromosome 7 GeneLoc Exon Structure GeneLoc location for GC07M105730: view genomic region
(about GC identifiers)
Start:
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105,730,949 bp from pter |
End:
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105,753,057 bp from pter |
Size:
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22,109 bases |
Orientation:
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minus strand |
1 alternative location:
| Chr7-,CRA_TCAG 105,091,975-105,114,065 |
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Proteins for SYPL1 gene
(According to
1UniProtKB,
HORDE,
neXtProt,
Ensembl,
and/or Reactome,
Modification sites according to 2PhosphoSitePlus,
Specific Peptides from DME,
Protein expression images according to data from SPIRE MOPED and PaxDb,
RefSeq according to NCBI,
PDB rendering according to OCA and/or Proteopedia,
Recombinant Proteins
from
EMD Millipore,
R&D Systems,
GenScript,
Enzo Life Sciences,
OriGene,
Novus Biologicals,
Sino Biological,
ProSpec, and/or
Uscn,
Biochemical Assays by
EMD Millipore,
R&D Systems,
OriGene,
GenScript,
Cell Signaling Technology,
Enzo Life Sciences, and/or
Uscn,
Ontologies according to Gene
Ontology Consortium 01 Mar 2013 and
Entrez Gene,
Antibodies by
EMD Millipore,
R&D Systems,
GenScript,
Cell Signaling Technology,
OriGene,
Novus Biologicals,
Thermo Fisher Scientific,
Abcam, and/or
Uscn)
About This Section
| UniProtKB/Swiss-Prot: SYPL1_HUMAN, Q16563 (See
protein sequence)Recommended Name: Synaptophysin-like protein 1 Size: 259 amino acids; 28565 Da
Subcellular location: Cytoplasmic vesicle membrane; Multi-pass membrane protein (By similarity). Melanosome.Note=Cytoplasmic transport vesicles (By similarity). Identified by mass spectrometry in melanosome fractions fromstage I to stage IV
Secondary accessions: Q96AR8Alternative splicing: 2 isoforms: Q16563-1 Q16563-2 Explore the universe of human proteins at neXtProt for SYPL1: NX_Q16563
Post-translational modifications:
View modification sites using PhosphoSitePlus2View neXtProt modification sites for NX_Q16563 SYPL1 Protein expression data from MOPED and PaxDb: About this image 
REFSEQ proteins (2 alternative transcripts):
NP_006745.1 NP_874384.1 ENSEMBL proteins: ENSP00000388336 ENSP00000011473 ENSP00000417473 ENSP00000419070 Human Recombinant Protein Products:
Gene Ontology (GO): 5/7 cellular component terms (GO ID links to tree view) (see all 7): About this table
SYPL1 for ontologies About GeneDecksing
SYPL1 Antibody Products: Assay Products for SYPL1: |
Protein
Domains / Families for SYPL1 gene(According to InterPro, ProtoNet,
UniProtKB, and/or BLOCKS,
Sets of similar genes according to GeneDecks)
About This Section
|
SYPL1 for domains About GeneDecksing
3 InterPro domains/families:Graphical View of Domain Structure for InterPro Entry Q16563ProtoNet protein and cluster: Q16563 1 Blocks protein family: IPB008253 Marvel
UniProtKB/Swiss-Prot: SYPL1_HUMAN, Q16563Similarity: Belongs to the synaptophysin/synaptobrevin familySimilarity: Contains 1 MARVEL domain |
Function for SYPL1 gene
(According to 1UniProtKB,
Genatlas,
LifeMap Discovery™,
IUBMB, and/or
2DME,
Human phenotypes from GenomeRNAi,
Animal models from MGI Mar 06 2013,
bound targets from SABiosciences,
miRNA Gene Targets from miRTarBase
shRNA from
OriGene,
RNAi from
EMD Millipore,
siRNAs from
OriGene,
QIAGEN,
microRNA from QIAGEN,
Gene Editing from DNA2.0,
Clones from EMD Millipore,
OriGene,
SwitchGear Genomics,
GenScript,
Sino Biological,
DNA2.0,
and Vector BioLabs,
Cell Lines from GenScript,
LifeMap BioReagents,
In Situ Hybridization Assays from Advanced Cell Diagnostics,
Ontologies according to Gene Ontology Consortium 01 Mar 2013 via
Entrez Gene.)
About This Section
|
Clone Products: |  | Browse Clones for the Expression of Recombinant Proteins Available from EMD Millipore | |  | OriGene Myc/DDK tagged cDNA clones in CMV expression vector in human, mouse, rat for SYPL1 (see all 4) OriGene untagged cDNA clones in CMV expression vector in human, mouse, rat for SYPL1 (see all 3) OriGene custom cloning services – gene synthesis, subcloning, mutagenesis,
variant library, vector shuttling 
| |  | GenScript: all cDNA clones in your preferred vector (see all 2): SYPL1 (NM_182715) | |  | Browse Sino Biological Human cDNA Clones | |  | DNA2.0 Custom Codon Optimized Gene
Synthesis Service for SYPL1 | |  | Vector BioLabs ready-to-use adenovirus/AAV for human, mouse, rat SYPL1  |
In Situ Assay Products: |
| Advanced Cell Diagnostics RNAscope RNA in situ hybridization assays for SYPL1 |
Gene Ontology (GO): 1 molecular function term (GO ID links to tree view): About this table | GO ID | Qualified GO term | Evidence | PubMed IDs |
|---|
| GO:0005215 | transporter activity |
IEA | -- |
SYPL1 for ontologies About GeneDecksing
|
Pathways & Interactions for SYPL1 gene
(Pathways according to
EMD Millipore,
R&D Systems,
Cell Signaling Technology,
KEGG,
PharmGKB,
BioSystems,
Reactome,
Tocris Bioscience,
GeneGo (Thomson Reuters),
QIAGEN,
and/or UniProtKB,
Sets of similar genes according to GeneDecks,
Interaction Networks according to
SABiosciences,
and/or STRING,
Interactions according to 1UniProtKB,
2MINT,
3I2D, and/or
4STRING,
with links to IntAct and
Ensembl,
Ontologies according to Gene Ontology Consortium 01 Mar 2013 via
Entrez Gene).
About This Section
|
Interactions:
Search SABiosciences Gene Network CentralTM Interacting Genes and Proteins Networks for SYPL1
STRING Interaction
Network Preview (showing 1 interactants - click image to see more details)
 1 Interacting protein for SYPL1 (Q165633 ENSP000000114734) via UniProtKB, MINT, STRING, and/or I2DAbout this table
Gene Ontology (GO): 1 biological process term (GO ID links to tree view): About this table
SYPL1 for ontologies About GeneDecksing
|
Drugs & Compounds for SYPL1 gene(Chemical Compounds according to UniProtKB, Enzo Life Sciences,
EMD Millipore, Tocris Bioscience
HMDB,
BitterDB, and/or
Novoseek, and Drugs according to
DrugBank,
Enzo Life Sciences, and/or
PharmGKB, with drugs/clinical trials/news
search links to CenterWatch)
About This Section
|
Browse Tocris compounds for SYPL1 Search CenterWatch for drugs/clinical trials and news about SYPL1 
|
Transcripts for SYPL1 gene(Secondary structures according to
fRNAdb,
GenBank/EMBL/DDBJ Accessions according to
Unigene
(Build 235 Homo sapiens; Mar 10 2013) or GenBank, RefSeq according to Entrez Gene,
DOTS (version 10), and/or
AceView,
transcript ids from Ensembl
with links to UCSC,
exon structure from GeneLoc,
alternative splicing isoforms according to ASD and/or
ECgene,
RNAi Products from
EMD Millipore,
siRNAs from
OriGene,
QIAGEN,
shRNA from
OriGene,
microRNA from QIAGEN,
Tagged/untagged cDNA clones from
OriGene,
SwitchGear Genomics,
GenScript,
DNA2.0,
Vector BioLabs,
Primers from
OriGene,
SABiosciences, and/or
QIAGEN
) About This Section
| REFSEQ mRNAs for SYPL1 gene (2 alternative transcripts): NM_006754.3 NM_182715.2 Unigene Cluster for SYPL1: Synaptophysin-like 1 Hs.80919 [show with all ESTs]Unigene Representative Sequence: AK1282996 Ensembl transcripts including schematic representations, and UCSC links where relevant: ENST00000455385(uc003vdo.3) ENST00000011473(uc003vdp.3) ENST00000464029 ENST00000470347 ENST00000460770 ENST00000483448
Clone Products: |  | OriGene Myc/DDK tagged cDNA clones in CMV expression vector in human, mouse, rat for SYPL1 (see all 4) OriGene untagged cDNA clones in CMV expression vector in human, mouse, rat for SYPL1 (see all 3) OriGene custom cloning services – gene synthesis, subcloning, mutagenesis,
variant library, vector shuttling 
| |  | GenScript: all cDNA clones in your preferred vector (see all 2): SYPL1 (NM_182715) | |  | DNA2.0 Custom Codon Optimized Gene
Synthesis Service for SYPL1 | |  | Vector BioLabs ready-to-use adenovirus/AAV for human, mouse, rat SYPL1  |
Additional cDNA sequence: AK128279.1 AK128299.1 AK292230.1 BC016835.1 BC020938.1 BC061887.1 S72481.1 X61382.1 X68194.1 22 DOTS entries: DT.101984298 DT.100880282 DT.91778483 DT.75193648 DT.121053658 DT.100880280 DT.95357184 DT.95365565 DT.97846300 DT.100767573 DT.121053704 DT.91778476 DT.100880284 DT.100880278 DT.95365554 DT.95365566 DT.100880273 DT.100880274 DT.100880281 DT.121053655 DT.121053664 DT.121053694 24/588 AceView cDNA sequences (see all 588): AA312682 BF002255 BG290426 AA374693 AA419082 AI168091 AA383493 BC016835 CR616230 AI689919 BP354092 AI632714 AI631583 CB112057 AI392986 AI367336 BP356314 BE465961 H06708 BP340625 BP351726 CD723586 AW244137 BQ439312 GeneLoc Exon Structure
5/8 Alternative Splicing Database (ASD) splice patterns (SP) for SYPL1 (see all 8) About this scheme
| ExUns: | 1a | · | 1b | · | 1c | ^ | 2a | · | 2b | · | 2c | · | 2d | ^ | 3 | ^ | 4 | ^ | 5a | · | 5b | ^ | 6a | · | 6b | · | 6c | ^ | 7a | · | 7b | · | 7c | ^ | 8a | · | 8b | · | 8c | · | 8d | |
| SP1: | |   | |   | |   | - |   | |   | |   | - |   | |   | |   | |   | |   | |   | |   | |   | - |   | - |   | - |   | |   | |   | |   | |   | |
| SP2: | |   | |   | |   | - |   | |   | |   | - |   | |   | |   | |   | |   | |   | |   | |   | |   | |   | |   | |   | |   | |   | |   | |
| SP3: | |   | |   | |   | - |   | |   | |   | - |   | |   | |   | - |   | |   | |   | |   | |   | |   | |   | |   | |   | |   | |   | |   | |
| SP4: | |   | |   | |   | |   | |   | |   | - |   | |   | |   | |   | |   | |   | |   | |   | - |   | - |   | - |   | |   | |   | |   | |   | |
| SP5: | |   | |   | |   | |   | |   | |   | - |   | |   | - |   | |   | |   | |   | |   | |   | |   | |   | |   | |   | |   | |   | |   |
ECgene alternative splicing isoforms for SYPL1
|
Expression for SYPL1 gene
(RNA expression data according to
H-InvDB,
NONCODE,
miRBase, and
RNAdb,
Expression images according to data from
BioGPS,
Illumina Human BodyMap, and
CGAP
SAGE,
Sets of similar genes according to GeneDecks,
in vivo and in vitro expression data from LifeMap Discovery™,
plus additional links to
Genevestigator, and/or
SOURCE, and/or
BioGPS, and/or
UniProtKB,
PCR Arrays from
SABiosciences,
Primers from
OriGene,
SABiosciences, and/or
QIAGEN,
In Situ Hybridization Assays from Advanced Cell Diagnostics)
About This Section
| SYPL1 expression in normal human tissues (normalized intensities) See probesets specificity/sensitivity at GeneAnnot About this imageBioGPS CGAP TAG: ACAAGAATTG
About this image See SYPL1 Protein Expression from SPIRE MOPED and PaxDB Genevestigator expression for SYPL1
SOURCE GeneReport for Unigene cluster: Hs.80919 SABiosciences Custom PCR Arrays for SYPL1
Primer Products: |  | OriGene genome-wide validated SYBR primer pairs in human, mouse, rat for SYPL1 Browse OriGene validated miRNA SYBR primer pairs
| |  | SABiosciences RT2 qPCR Primer Assay in human, mouse, rat SYPL1 | |  | QIAGEN QuantiTect SYBR Green Assays in human, mouse, rat SYPL1 | |  | QIAGEN QuantiFast Probe-based Assays in human, mouse, rat SYPL1 | In Situ Assay Products: |
| Advanced Cell Diagnostics RNAscope RNA in situ hybridization assays for SYPL1 |
Orthologs for SYPL1 gene
(Orthologs according to
1,2HomoloGene (2older version, for species not in 1newer version),
3euGenes,
4SGD
,
5MGI Mar 06 2013,
with possible further links to
Flybase
and/or
WormBase,
and/or
6Ensembl pan taxonomic compara ,
Gene Trees according to Ensembl and
TreeFam)
About This Section
|
This gene was present in the common ancestor of animals.
Orthologs for SYPL1 gene from 5/17 species (see all 17) About this table
ENSEMBL Gene Tree for SYPL1 (if available) TreeFam Gene Tree for SYPL1 (if available)  |
Paralogs for SYPL1 gene(Paralogs according to
1HomoloGene, 2Ensembl, and 3SIMAP, Pseudogenes according to 4Pseudogene.org Build 68) About This Section
| Paralogs for SYPL1 gene
- SYNPR2 SYP2 SYPL22
9 SIMAP similar genes for SYPL1 using alignment to 3 protein entries: SYPL1_HUMAN (see all proteins):SYPL ZNF74 AURA2 FAM3A ZNF564 CRCP SYPL2 SYNPR SYP
SYPL1 for paralogs About GeneDecksing
3 Pseudogenes.org Pseudogenes for SYPL1 PGOHUM00000236996 PGOHUM00000237826 PGOHUM00000243403
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Genomic Variants for SYPL1 gene(SNPs/Variants according to the
1NCBI SNP Database,
2Ensembl,
3PupaSUITE,
UniProtKB, and
DNA2.0,
Linkage Disequilibrium by HapMap,
Structural Variations(CNVs/InDels/Inversions) from the Database of Genomic Variants, Mutations from the Human Gene
Mutation Database (HGMD) and the Locus Specific Mutation
Databases (LSDB), Blood group antigen gene mutations by BGMUT,
Resequencing Primers from QIAGEN,
Cancer Mutation PCR Arrays and Assays and Copy Number PCR Arrays from SABiosciences)
About This Section
|
| Genomic Data | Transcription Related Data | Allele Frequencies | | SNP ID | Valid | Clinical significance | Chr 7 pos | Sequence | # | AA Chg | Type | More | # | Allele freq | Pop | Total sample | More |
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HapMap Linkage Disequilibrium report for SYPL1 (105730949 - 105753057 bp)
Structural Variations (Copy Number Variations, Insertions/Deletions, Inversions) Database of Genomic Variants (DGV): 3 variations for SYPL1 3 Indels: 12955 42410 62712  | SABiosciences Cancer Mutation PCR Assays |
|  | QIAGEN SeqTarget long-range PCR primers in human, mouse, rat for resequencing SYPL1 |
|
Disorders
/ Diseases for SYPL1 gene
(in which this Gene is Involved, According to MalaCards,
OMIM, UniProtKB,
the University of Copenhagen DISEASES
database,
Genatlas, GeneTests,
GAD,
HuGE Navigator,
and/or TGDB.)
About This Section
|
SYPL1 for disorders About GeneDecksing
2 diseases for SYPL1: About MalaCardsneuronitis alcoholism Export disorders for SYPL1 gene to outside databases
|
Publications for SYPL1 gene (in
PubMed.
Associations of this gene to articles via
1Entrez Gene,
2UniProtKB/Swiss-Prot,
3HGNC,
4GAD,
5PharmGKB,
6HMDB,
7DrugBank,
8UniProtKB/TrEMBL,
9 Novoseek, and/or
10fRNAdb)
About This Section
|
PubMed articles for SYPL1 gene, integrated from 9 sources (see all 23): (articles sorted by number of sources associating them with SYPL1) | |  | Utopia: connect your pdf to the dynamic world of online information |
- Expression of the synaptophysin gene family is not restricted to neuronal and neuroendocrine differentiation in rat and human. (PubMed id 8034131)1, 2, 9 Leube R.E. (1994)
- Proteomic and bioinformatic characterization of the biogenesis and function of melanosomes. (PubMed id 17081065)1, 2 Chi A....Hunt D.F. (2006)
- The status, quality, and expansion of the NIH full-length cDNA project: the Mammalian Gene Collection (MGC). (PubMed id 15489334)1, 2 Gerhard D.S....Malek J. (2004)
- The DNA sequence of human chromosome 7. (PubMed id 12853948)1, 2 Hillier L.W.... Wilson R.K. (2003)
- Molecular cloning of a cDNA encoding a novel protein related to the neuronal vesicle protein synaptophysin. (PubMed id 1730063)1, 2 Zhong C.Z.... Runge M.S. (1992)
- Pantophysin is a phosphoprotein component of adipocyte transport vesicles and associates with GLUT4-containing vesicles. (PubMed id 10636906)1, 9 Brooks C.C....Cheatham B. (2000)
- Pantophysin is a ubiquitously expressed synaptophysin homologue and defines constitutive transport vesicles. (PubMed id 8707851)1, 9 Haass N.K....Leube R.E. (1996)
- Initial characterization of the human central proteome. (PubMed id 21269460)2 Burkard T.R.... Colinge J. (2011)
- Mass spectrometric analysis of lysine ubiquitylation r eveals promiscuity at site level. (PubMed id 21139048)1 Danielsen J.M....Nielsen M.L. (2011)
- Glycoproteomics analysis of human liver tissue by combination of multiple enzyme digestion and hydrazide chemistry. (PubMed id 19159218)2 Chen R.... Zou H. (2009)
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External Searches for SYPL1 gene
(in PubMed,
OMIM, and NCBI Bookshelf) About This Section
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Genome Databases showing SYPL1 gene
(According to
Entrez Gene,
HGNC,
AceView,
euGenes,
Ensembl,
miRBase,
ECgene,
Kegg,
and/or
H-InvDB)
About This Section
|
|
Other Databases showing SYPL1 gene
(According to HUGE)
About This Section
| -- |
Specialized Databases showing SYPL1 gene(According to PharmGKB,
ATLAS, HORDE, IMGT, LEIDEN, UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL, Wikipedia and/or GeneReviews via UniProtKB/Swiss-Prot) About This Section
|
| Name | Description |
| PharmGKB entry for SYPL1 | Pharmacogenomics, SNPs, Pathways |
|
| | |
About This Section
| Patent Information for SYPL1 gene: Search GeneIP for patents involving SYPL1
GeneCards and IP: Japan Patent Office Licenses GeneCards European Patent Office Licenses GeneCards Improving the IP Search
|
Products for SYPL1 gene(Antibodies, recombinant proteins, and assays from EMD Millipore, R&D Systems, OriGene, QIAGEN, GenScript, Cell Signaling Technology, SABiosciences, Novus Biologicals, Sino Biological, Enzo Life Sciences, Abcam, ProSpec, Uscn, Thermo Fisher Scientific, Gene Editing from DNA2.0, Clones from EMD Millipore, OriGene, GenScript, Sino Biological, DNA2.0, SwitchGear Genomics, Vector BioLabs, Cell lines from GenScript and LifeMap BioReagents, PCR Arrays from SABiosciences, Drugs and/or compounds from EMD Millipore, Tocris Bioscience, and/or
Enzo Life Sciences), In Situ Hybridization Assays from Advanced Cell Diagnostics About This Section
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| | | | OriGene Antibodies for SYPL1 | | OriGene shRNA RFP for SYPL1 | | OriGene 29mer shRNA kits in GFP-retroviral vector in human, mouse, rat for SYPL1 | | OriGene genome-wide validated SYBR primer pairs in human, mouse, rat for SYPL1 | | OriGene Protein Over-expression Lysate for SYPL1 | | Browse OriGene Fluorogenic Cell Assay Kits | | OriGene siRNA for SYPL1 | | OriGene 3'-UTR Clone for SYPL1 | | OriGene untagged cDNA clones in CMV expression vector in human, mouse, rat for SYPL1 | | OriGene Myc/DDK tagged cDNA clones in CMV expression vector in human, mouse, rat for SYPL1 | | Browse OriGene GFP tagged cDNA clones in CMV expression vector | | Browse OriGene MicroRNA Expression Plasmids | | Browse OriGene basic RS shRNAs | | Browse OriGene validated miRNA SYBR primer pairs | | Browse OriGene full length recombinant human proteins expressed in human HEK293 cells | | OriGene custom cloning services - gene synthesis, subcloning, mutagenesis,
variant library, vector shuttling | | OriGene Custom Antibody Services for SYPL1 | | OriGene Custom Protein Services for SYPL1 | | OriGene Custom Immunoassay Development | | |
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| | QIAGEN Custom miScript Target Protector blocks miRNA-binding site of in human, mouse, rat SYPL1 | | QIAGEN SeqTarget long-range PCR primers in human, mouse, rat for resequencing SYPL1 | | QIAGEN PyroMark CpG Assay predesigned Pyrosequencing DNA Methylation assays in human, mouse, rat SYPL1 | | QIAGEN FlexiTube/FlexiPlate siRNA for gene silencing in human, mouse, rat SYPL1 | | QIAGEN QuantiFast Probe-based Assays in human, mouse, rat SYPL1 | | QIAGEN QuantiTect SYBR Green Assays in human, mouse, rat SYPL1 |
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| | | Search Tocris compounds for SYPL1 |
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 | | SYPL1 Proteins, Antibodies, CLIAs, and ELISAs |
| | | | Advanced Cell Diagnostics RNAscope RNA in situ hybridization assays for SYPL1 |
|  |  |  | | | ThermoFisher Antibodies for SYPL1 |
| | Vector BioLabs ready-to-use adenovirus/AAV for human, mouse, rat SYPL1 |
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