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DENND5A Gene

protein-coding GIFtS: 46

GC11M009117

DENN/MADD domain containing 5A
(Previous name: RAB6 interacting protein 1 )
Symbol approved by the HUGO Gene Nomenclature Committee (HGNC) database
(Previous symbol: RAB6IP1)

(According to ¹HGNC, ²Entrez Gene,
³UniProtKB/Swiss-Prot, ⁴UniProtKB/TrEMBL, ⁵OMIM, ⁶GeneLoc , and/or ⁷Ensembl, ⁸miRBase)
About This Section

Aliases
FLJ22354 ^¹, ²
FLJ33829 ^¹, ²
FLJ43455 ^¹, ²
KIAA1091 ^¹, ², ³
RAB6IP1 ^², ³
Rab6IP1 ^³

Descriptions
DENN/MADD domain containing 5A ^²
RAB6 interacting protein 1 ^¹, ²
Rab6-interacting protein 1 ^³

External Ids
HGNC: 19344¹
Entrez Gene: 23258²
UniProtKB: Q6IQ26³
Ensembl: ENSG00000184014⁷

Search outside databases for aliases for DENND5A gene

(According to Entrez Gene, Wikipedia's Gene Wiki,
UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL)
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UniProtKB/Swiss-Prot: DEN5A_HUMAN, Q6IQ26

Function: May be involved in Rab6-mediated GTPase signaling

(According to GeneLoc and/or HGNC, and/or
Entrez Gene (NCBI build 36),
and/or miRBase,
Genomic Views according to UCSC and Ensembl, Transcription factor binding sites according to SABiosciences)
About This Section

Genomic View:

UCSC Golden Path with GeneCards custom track

Entrez Gene cytogenetic band: 11p15.4 Ensembl cytogenetic band: 11p15.4 HGNC cytogenetic band: 11p15.3

DENND5A Gene in genomic location: bands according to Ensembl, locations according to (and/or Entrez Gene and/or Ensembl if different)

GeneLoc gene densities for chromosome 11 GeneLoc Exon Structure

GeneLoc location for GC11M009117: (about GC identifiers)

Start:	9,116,948 bp from pter
End:	9,243,447 bp from pter
Size:	126,500 bases
Orientation:	minus strand

RefSeq DNA sequence:

NC_000011.8 NT_009237.17

(According to ¹UniProtKB, and/or Ensembl, Phosphorylation sites according to ²Phosphosite, RefSeq according to NCBI, PDB rendering according to OCA and/or Proteopedia, Recombinant Proteins from Invitrogen, Millipore, Sigma-Aldrich, R&D Systems, Enzo Life Sciences, Abnova, OriGene and/or, Abcam,
Biochemical Assays by Invitrogen, Millipore, R&D Systems, Cell Signaling Technology, and/or Enzo Life Sciences, Ontologies according to Gene Ontology Consortium 01 Apr 2009 and Entrez Gene, Antibodies by Invitrogen, Millipore, Sigma-Aldrich, R&D Systems, Cell Signaling Technology, Abcam, Abnova, and/or Novus Biologicals)
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UniProtKB/Swiss-Prot: DEN5A_HUMAN, Q6IQ26 (See protein sequence)

Recommended Name: DENN domain-containing protein 5A

Size: 1287 amino acids; 147095 Da

Subunit: Probably interacts with RAB6A bound to GTP (By similarity)

Secondary accessions: Q96GN3 Q9H6U7 Q9UFV0 Q9UPR1

Post-translational modifications:

View phosphorylation sites using PhosphoSite²

REFSEQ proteins: NP_056028.2

ENSEMBL proteins:

ENSP00000328524

Human Recombinant Proteins

	Browse Drug Discovery Central at Invitrogen for human recombinant proteins
	Browse Purified and Recombinant Proteins at Millipore
	Browse Human Recombinant Proteins at Sigma-Aldrich
	Browse R&D Systems for human recombinant proteins
	Browse recombinant and purified proteins available from Enzo Life Sciences
	Browse Abcam for Recombinant Proteins
	Human Recombinant Proteins from Abnova (RAB6IP1)

Browse Origene for full length recombinant human proteins expressed in human HEK293 cells

Antibodies for DENND5A:

	Browse Antibodies Central at Invitrogen
	Browse Millipore's Extensive Line of Mono- and Polyclonal Antibodies
	Browse Antibodies at Sigma-Aldrich
	Browse R&D Systems for Antibodies
	Search for antibodies at Abcam and the World's Antibody Gateway (free search engine of over 150 antibody companies)
	Monoclonal and Polyclonal Antibodies from Abnova (RAB6IP1)
	Novus Biologicals Antibody for DENND5A

Assays for DENND5A:

	Browse Invitrogen for biochemical assays
	Browse Kits and Assays available from Millipore
	Browse R&D Systems for biochemical assays
	Browse biochemical assays available from Enzo Life Sciences

(According to InterPro, ProtoNet, UniProtKB, and/or BLOCKS, Sets of similar genes according to GeneDecks)
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Graphical View of Domain Structure for InterPro Entry Q6IQ26

ProtoNet protein and cluster: Q6IQ26

1 Blocks protein family: IPB001024 Lipoxygenase

UniProtKB/Swiss-Prot: DEN5A_HUMAN, Q6IQ26

Similarity: Belongs to the RAB6IP1 family

Similarity: Contains 1 dDENN domain

Similarity: Contains 1 DENN domain

Similarity: Contains 1 PLAT domain

Similarity: Contains 2 RUN domains

Similarity: Contains 1 uDENN domain

(According to MGI Jun 06 2009, UniProtKB, IUBMB,and/or Genatlas,
shRNA from OriGene, Sigma-Aldrich, RNAi from Sigma-Aldrich,
RNAi Products, Clones, and Q-PCR Products from Invitrogen, Millipore, OriGene, and/or Abnova, siRNAs from Applied Biosystems, SYBR primers from OriGene, Cell-based Assays from Millipore, Ontologies according to Gene Ontology Consortium 01 Apr 2009 via Entrez Gene.)
About This Section

Inhib. RNA:		Invitrogen RNAi Products for gene knock-down (DENND5A)
		Browse for Gene Knock-down Tools from Millipore
		Abnova Chimera RNAi Products for Gene knock-down (RAB6IP1)

OriGene 29mer shRNA kit in GFP-retroviral vectors (see all 2): NM_015213

Applied Biosystems Silencer® siRNAs for DENND5A

Sigma-Aldrich siRNA for DENND5A

Sigma-Aldrich shRNA for DENND5A

Explore Sigma-Aldrich super-pooled esiRNAs

Clones:		Invitrogen Clones for DENND5A
		Browse Clones for the Expression of Recombinant Proteins Available from Millipore

OriGene GFP tagged cDNA clone in CMV expression vector: NM_015213
Myc/DDK tagged cDNA clone in CMV expression vector: NM_015213
untagged cDNA clone in CMV expression vector: NM_015213

Primers:

Browse Quantitative PCR Central at Invitrogen for Q-PCR LUX™ Primers

OriGene genome-wide validated SYBR primer pairs: NM_015213

UniProtKB/Swiss-Prot: DEN5A_HUMAN, Q6IQ26

Function: May be involved in Rab6-mediated GTPase signaling

(Pathways according to Invitrogen (maps by GeneGo), Millipore, Cell Signaling Technology, Sigma-Aldrich, KEGG and/or UniProtKB,
Sets of similar genes according to GeneDecks, Proteins Network according to SABiosciences, Interactions according to ¹UniProtKB, ²MINT, and/or ³STRING, with links to IntAct and Ensembl, Ontologies according to Gene Ontology Consortium 01 Apr 2009 via Entrez Gene.)
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(Chemical Compounds according to UniProtKB, Enzo Life Sciences, Sigma-Aldrich, Tocris Bioscience, and/or Novoseek and Drugs according to Enzo Life Sciences and/or PharmGKB)
About This Section

Browse drugs & compounds from Enzo Life Sciences

Browse Small Molecules at Sigma-Aldrich

Browse Tocris compounds for DENND5A

(GenBank/EMBL/DDBJ Accessions according to
Unigene (Build 219 Homo sapiens; Jun 2 2009) or GenBank,
RefSeq according to Entrez Gene,
DOTS (version 10), and/or AceView,
non coding RNAs according to RNAdb,
ESTs according to GeneTide,
exon structure from GeneLoc, alternative splicing isoforms according to ASD and/or ECgene,
RNAi Products from Invitrogen, Millipore, and/or Abnova,
siRNAs from Applied Biosystems, Sigma-Aldrich,
shRNA from Sigma-Aldrich, OriGene,
Tagged/untagged cDNA clones from OriGene,
Expression Assays from Applied Biosystems)
About This Section

Inhib. RNA:		Invitrogen RNAi Products for gene knock-down (DENND5A)
		Browse for Gene Knock-down Tools from Millipore
		Abnova Chimera RNAi Products for Gene knock-down (RAB6IP1)

OriGene 29mer shRNA kit in GFP-retroviral vectors (see all 2): NM_015213

Sigma-Aldrich siRNA for DENND5A

Sigma-Aldrich shRNA for DENND5A

Explore Sigma-Aldrich super-pooled esiRNAs

Applied Biosystems Silencer® siRNAs:

NM_015213

REFSEQ mRNAs for DENND5A gene:

NM_015213.2

Applied Biosystems TaqMan ® Gene Expression Assays:

NM_015213

              OriGene GFP tagged cDNA clone in CMV expression vector: NM_015213
                                 Myc/DDK tagged cDNA clone in CMV expression vector: NM_015213
                                 untagged cDNA clone in CMV expression vector: NM_015213

Additional cDNA sequence:

AB029014.1 AK025499.1 AK026007.1 AK091148.1 AK125444.1 AK294016.1 AK295881.1 AK302478.1

AL117448.1 BC005100.1 BC009354.2 BC018797.1 BC071596.1

15 DOTS entries:

DT.87017134 DT.101956321 DT.101986352 DT.95329667 DT.100784277 DT.100784278 DT.91823780 DT.95329646

DT.97844609 DT.100784279 DT.102829477 DT.112569 DT.92024766 DT.92437017 DT.97844610

24/290 AceView cDNA sequences (see all 290 ):

BF982271 CB131470 AI003545 AI927774 BF115019 AB029014 BI861576 AI494213

AI375155 AW615313 BQ670079 BG540403 AI093534 CA440880 BU155999 BQ672442

AI372726 BU553540 AA349621 AA365828 AK025499 CF136888 BM721990 BE296081

Unigene Cluster for DENND5A:

DENN/MADD domain containing 5A

Hs.501857 [show with all ESTs]

Unigene Representative Sequence: NM_015213

GeneLoc Exon Structure

5/10 Alternative Splicing Database (ASD) splice patterns (SP) for DENND5A (see all 10 )

ExUns:	1a	·	1b	^	2	^	3	^	4	^	5a	·	5b	^	6	^	7a	·	7b	^	8a	·	8b	^	9a	·	9b	^	10	^	11	^	12	^	13	^	14	^	15a	·	15b	^	16	^	17a	·	17b	^	18a	·	18b	^
SP1:							-																						-		-
SP2:																																															-		-		-
SP3:
SP4:
SP5:

ExUns:	19	^	20	^	21a	·	21b	^	22	^	23a	·	23b	^	24	^	25	^	26	^	27a	·	27b	·	27c	·	27d	·	27e
SP1:					-														-
SP2:	-		-		-		-		-		-								-
SP3:	-				-
SP4:					-						-		-		-				-
SP5:																			-

About this scheme

ECgene alternative splicing isoforms for DENND5A

1 Ensembl transcript including schematic representation:

ENST00000328194

(Experimental results according to ¹GeneNote and GNF BioGPS,
probe sets-to-genes annotations according to ²GeneAnnot , ³GeneTide , Sets of similar genes according to GeneDecks, Electronic Northern calculations according to data from UniGene (Build 219 Homo sapiens), SAGE tags according to CGAP, plus additional links to SOURCE, and/or GNF BioGPS, and/or EXPOLDB, and/or UniProtKB,
Expression Assays from Applied Biosystems )
About This Section

DENND5A expression in normal and diseased human tissues

Applied Biosystems TaqMan ® Gene Expression Assays for DENND5A

¹ / ²

3 probe-sets matching DENND5A gene

Affymetrix probe-set	Array	# genes	Sensitivity	Specificity	Correlation	Length	Gb_Accession	Consensus	Uniqueness	Score	Rank
Affymetrix probe-set	Array	GeneAnnot data			GeneNote data		GeneTide data
33924_at²	U95-A	1	1.00	1.00	1.00	1.00	--	--	--	--	--
212561_at²	U133-A	1	1.00	1.00	--	--	--	--	--	--	--
212561_at²	U133Plus2	1	1.00	1.00	--	--	--	--	--	--	--

GeneDecks DENND5A for binary patterns associated with the probe-sets selected above

About GeneDecksing

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Data from

(Publications) and GNF BioGPS
About these images
About these images

CGAP SAGE TAG: TTTTAACAGG

SOURCE GeneReport for Unigene cluster: Hs.501857

(Orthologs according to ^1,2HomoloGene (²older version, for species not in ¹newer version), ³euGenes, ⁴SGD and/or ⁵MGI Jun 06 2009, with possible further links to Flybase and/or WormBase, Gene Trees according to Ensembl)
About This Section

Orthologs for DENND5A gene from 5/9 species (see all 9 )

Organism	Gene	Locus	Description	Human Similarity	NCBI accessions
dog (Canis familiaris)	DENND5A¹	--	DENN/MADD domain containing 5A	94.29(n) 98.6(a)	476844 XM_534048.2 XP_534048.2
chimpanzee (Pan troglodytes)	DENND5A¹	--	DENN/MADD domain containing 5A	99.64(n) 99.77(a)	451734 XM_508914.2 XP_508914.2
cow (Bos taurus)	DENND5A¹	--	DENN/MADD domain containing 5A	93(n) 97.9(a)	784484 XM_001252217.1 XP_001252218.1
rat (Rattus norvegicus)	Dennd5a¹	--	DENN/MADD domain containing 5A	90.7(n) 97.59(a)	308942 XM_219270.3 XP_219270.3
mouse (Mus musculus)	Dennd5a^{1, 5}	7⁵	DENN/MADD domain containing 5A^{1, 5}	90.99(n)¹ 97.75(a)¹	19347¹ NM_021494.1¹ NP_067469.1¹ AJ245569⁵ AK005224⁵ (see all 10)

About this table Species with no ortholog for DENND5A

ENSEMBL Gene Tree for DENND5A

(Paralogs according to ¹HomoloGene
and ²Ensembl, Pseudogenes according to ³Pseudogene.org)
About This Section

Paralogs for DENND5A gene

DENND4B² DENND5B²

(According to the ¹NCBI SNP Database, ²Ensembl, ³PupaSUITE, and UniProtKB, Linkage Disequilibrium by HapMap, Genotyping Reagents from Applied Biosystems)
About This Section

10/653 NCBI SNPs in DENND5A are shown (see all 653 )
(Click for Applied Biosystems TaqMan ® Genotyping Assay) (see all 147)

AB	SNP ID	Valid	Chr 11 pos	Sequence	Recs	AA Chg	Type	More	Recs	Allele freq	Pop	Total sample	More
AB	Genomic Data				Transcription Data				Allele Frequencies
				--	--	--		--	--				--
	rs1353730^1,2	A,C,F,H,O	9116646(+)	`GACAAC/TCTCTG`	1	--	ng5¹		13		MN EA NA EU WA	1240
	rs2016375^1,2	A,C,F,H	9245029(-)	`GAGACG/AATGGC`	1	--	ng3¹		8		NA EA MN EU WA	852
	rs952374^1,2	C,F,H,O	9182062(+)	`TTGCTC/GCAGCA`	1	E/Q	mis¹		9		MN EU EA WA	1022
--	rs1809855^1,2	A,C	9245350(-)	`accacA/Gccctg`	1	--	ng3¹		0	--	--	--	--
	rs16906742^1,2	F,H	9117342(+)	`CCCGCG/ACAGCA`	1	--	ut3¹ ese³		6		NA EU EA	404
	rs11602070^1,2	C,H	9244924(+)	`AGGCAG/TCTGGC`	1	--	ng3¹ tfbs³		4		EU EA WA	418
	rs10466509^1,2	F,H	9117777(+)	`GCTGCT/CGGCTG`	1	--	ut3¹ ese³		4		EU EA WA	420
--	rs11558096^1,2	C	9120091(+)	`CTGAAA/GCTTGC`	1	F/L	mis¹		0	--	--	--	--
	rs2133217^1,2	A,C,F,H,O	9133588(-)	`TGTAAA/TCATTG`	1	--	int¹		7		EU EA WA NA	558
	rs2173406^1,2	A,C,F,H,O	9133659(-)	`GAACTA/GGAAAG`	1	--	int¹		11		EU EA WA NA	980

About this table

HapMap Linkage Disequilibrium images for DENND5A (up to first 250kb)

(in which this Gene is Involved, According to OMIM, UniProtKB, PharmGKB, Genatlas, GeneTests, Blood group antigen gene mutations by BGMUT, HGMD, GAD, HuGE Navigator, BCGD, and/or TGDB.)
About This Section

(Possibly Related Articles in Doctor's Guide)
About This Section

(in PubMed. Associations of this gene to articles via ¹Novoseek, ²HGNC, ³Entrez Gene, ⁴UniProtKB/Swiss-Prot, ⁵UniProtKB/TrEMBL, ⁶GAD, and/or ⁷PharmGKB)
About This Section

9 PubMed articles for DENND5A gene:

Prediction of the coding sequences of unidentified human genes. XIV. The complete sequences of 100 new cDNA clones from brain which code for large proteins in vitro. (PubMed id 10470851)^{2, 3, 4} Kikuno R....Ohara O. (1999)
The status, quality, and expansion of the NIH full-length cDNA project: the Mammalian Gene Collection (MGC). (PubMed id 15489334)^{3, 4} Gerhard D.S....Malek J. (2004)
Complete sequencing and characterization of 21,243 full-length human cDNAs. (PubMed id 14702039)^{3, 4} Ota T....Sugano S. (2004)
Structural basis for recruitment of Rab6-interacting protein 1 to Golgi via a RUN domain. (PubMed id 19141279)³ Recacha R....Khan A.R. (2009)
A quantitative atlas of mitotic phosphorylation. (PubMed id 18669648)⁴ Dephoure N.... Gygi S.P. (2008)
The full-ORF clone resource of the German cDNA Consortium. (PubMed id 17974005)⁴ Bechtel S....Schupp I. (2007)
Generation and initial analysis of more than 15,000 full-length human and mouse cDNA sequences. (PubMed id 12477932)³ Strausberg R.L....Marra M.A. (2002)
Two-hybrid system screen with the small GTP-binding protein Rab6. Identification of a novel mouse GDP dissociation inhibitor isoform and two other potential partners of Rab6. (PubMed id 7782346)³ Janoueix-Lerosey I....Goud B. (1995)
uDENN, DENN, and dDENN: indissociable domains in Rab and MAP kinases signaling pathways. (PubMed id 11563850)¹ Levivier E....Callebaut I. (2001)

(in PubMed, OMIM, and NCBI Bookshelf)
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		AND	OR
Aliases
Free Text

Query String		PubMed OMIM NCBI Bookshelf
	(Note: In FireFox, select the above section and copy using Ctrl-C)

(According to Entrez Gene, HGNC, AceView, euGenes, Ensembl, miRBase, ECgene, and/or H-InvDB)
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Entrez Gene: 23258	HGNC: 19344	AceView: RAB6IP1	Ensembl:ENSG00000184014	euGenes: HUgn23258
ECgene: DENND5A	H-InvDB: DENND5A

(According to HUGE)
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HUGE: KIAA1091

(According to ATLAS, HORDE, IMGT, MTDB, LEIDEN, UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL,
Wikipedia and/or GeneReviews via UniProtKB/Swiss-Prot)
About This Section

Licensable Technologies for DENND5A
(Available from WIS Yeda, Salk, Tufts)

About This Section

(Reagents available from Applied Biosystems, Antibodies and assays by Cell Signaling Technology, Abcam, Novus Biologicals,
Sigma-Aldrich, R&D Systems, Millipore, Abnova, and/or Invitrogen, Clones available from OriGene,and/or Invitrogen, Drugs and/or compounds by Sigma-Aldrich,
Enzo Life Sciences, and/or Tocris Bioscience)
About This Section

Products for DENND5A:

TaqMan ® Gene Expression Assays

TaqMan ® Genotyping Assays

Free SNP selection tool

Invitrogen iPath Pathways	Invitrogen BLOCK-iT™ RNAi
Invitrogen Antibodies	Invitrogen Assays
Invitrogen Clones	Invitrogen Q-PCR Products
Invitrogen Human Recombinant Kinases	Invitrogen Custom Antibody and Peptide Service
Invitrogen Proteins / Assays / Screening Services	Search Invitrogen catalog for DENND5A-related products

	Millipore Custom Antibody & Bulk Services
	Millipore Preclinical / Clinical Development Services
	Millipore Immunoassay Services
	Millipore Target Screening & Profiling Services

	Predesigned and custom siRNAs for DENND5A		Browse antibodies at Sigma-Aldrich
	Explore super-pooled esiRNAs		Browse proteins at Sigma-Aldrich
	Lentivirus-delivered shRNAs for DENND5A		Browse small molecules at Sigma-Aldrich
	"Your Favorite Gene" Pathways		feedback

	Antibodies		Primer Pairs
	Cell Culture Products		ELISAs
	Flow Cytometry Kits		Protease Activity Assays & Reagents
	Cell Selection/Detection Kits & Reagents		ELISA/Assay Development Kits & Reagents
	Multiplex/Array Assay Kits & Reagents		ELISpot Kits & Development Modules
	Recombinant & Natural Proteins

	Recombinant Proteins (RAB6IP1)
	Antibodies (RAB6IP1)
	Chimera RNAi (RAB6IP1)
	Custom Service for Mouse Mab
	Custom Service for Rabbit Pab from Full-length Protein

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DENND5A Gene at Home site.
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DENND5A Gene

protein-coding GIFtS: 46

Aliases &
Descriptions
for DENND5A

Summaries
for DENND5A

Genomic Location
for DENND5A

Proteins
for DENND5A

Protein Domains/
Families
for DENND5A

Gene Function
for DENND5A

Pathways & Interactions
for DENND5A

Drugs & Compounds
for DENND5A

Transcripts
for DENND5A

Expression
for DENND5A

Orthologs
for DENND5A

Paralogs
for DENND5A

SNPs/Variants
for DENND5A

Disorders & Mutations
for DENND5A

Medical News
for DENND5A

Publications
for DENND5A

Search for DENND5A

Genome Databases showing DENND5A

Other Databases showing DENND5A

Specialized Databases showing DENND5A

Licensable Technologies
for DENND5A

Services
for DENND5A

Gene Index: 3 5 A B C D E F G H I J K L M N O P Q R S T U V W X Y Z

DENND5A Gene

protein-coding GIFtS: 46

Aliases & Descriptionsfor DENND5A

Summariesfor DENND5A

Genomic Locationfor DENND5A

Proteinsfor DENND5A

Protein Domains/Familiesfor DENND5A

Gene Functionfor DENND5A

Pathways & Interactionsfor DENND5A

Drugs & Compoundsfor DENND5A

Transcriptsfor DENND5A

Expression for DENND5A

Orthologsfor DENND5A

Paralogsfor DENND5A

SNPs/Variantsfor DENND5A

Disorders & Mutationsfor DENND5A

Medical Newsfor DENND5A

Publicationsfor DENND5A

Search for DENND5A

Genome Databases showing DENND5A

Other Databases showing DENND5A

Specialized Databases showing DENND5A

Licensable Technologiesfor DENND5A

Servicesfor DENND5A

Gene Index: 3 5 A B C D E F G H I J K L M N O P Q R S T U V W X Y Z

Aliases &
Descriptions
for DENND5A

Summaries
for DENND5A

Genomic Location
for DENND5A

Proteins
for DENND5A

Protein Domains/
Families
for DENND5A

Gene Function
for DENND5A

Pathways & Interactions
for DENND5A

Drugs & Compounds
for DENND5A

Transcripts
for DENND5A

Expression
for DENND5A

Orthologs
for DENND5A

Paralogs
for DENND5A

SNPs/Variants
for DENND5A

Disorders & Mutations
for DENND5A

Medical News
for DENND5A

Publications
for DENND5A

Licensable Technologies
for DENND5A

Services
for DENND5A