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QPCT Gene

protein-coding GIFtS: 59

GC02P037483

glutaminyl-peptide cyclotransferase
Symbol approved by the HUGO Gene Nomenclature Committee (HGNC) database

(According to ¹HGNC, ²Entrez Gene,
³UniProtKB/Swiss-Prot, ⁴UniProtKB/TrEMBL, ⁵OMIM, ⁶GeneLoc , and/or ⁷Ensembl, ⁸miRBase)
About This Section

Aliases
EC ^³
EC 2.3.2.5 ^³
GCT ^¹, ²
QC ^¹, ², ³, ⁵

Descriptions
Glutaminyl cyclase ^², ³
Glutaminyl-tRNA cyclotransferase ^³
Glutamyl cyclase ^³
glutaminyl-peptide cyclotransferase ^²

External Ids
HGNC: 9753¹
Entrez Gene: 25797²
UniProtKB: Q16769³
Ensembl: ENSG00000115828⁷

Search outside databases for aliases for QPCT gene

Previous GC identifers: GC02P037536 GC02P037718 GC02P037546

(According to Entrez Gene, Wikipedia's Gene Wiki,
UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL)
About This Section

EntrezGene summary for QPCT:

This gene encodes human pituitary glutaminyl cyclase, which is responsible for the presence of

pyroglutamyl residues in many neuroendocrine peptides. The amino acid sequence of this enzyme is

86% identical to that of bovine glutaminyl cyclase. [provided by RefSeq]

UniProtKB/Swiss-Prot: QPCT_HUMAN, Q16769

Function: Responsible for the biosynthesis of pyroglutamyl peptides. Has a bias against acidic and

tryptophan residues adjacent to the N-terminal glutaminyl residue and a lack of importance of

chain length after the second residue. Also catalyzes N-terminal pyroglutamate formation. In

vitro, catalyzes pyroglutamate formation of N-terminally truncated form of APP amyloid-beta

peptides [Glu-3]-beta-amyloid. May be involved in the N-terminal pyroglutamate formation of

several amyloid-related plaque-forming peptides

Gene Wiki entry for QPCT

(According to GeneLoc and/or HGNC, and/or
Entrez Gene (NCBI build 36),
and/or miRBase,
Genomic Views according to UCSC and Ensembl, Transcription factor binding sites according to SABiosciences)
About This Section

Genomic View:

UCSC Golden Path with GeneCards custom track

Transcription factor binding sites upstream to the QPCT gene

Entrez Gene cytogenetic band: 2p22.2 Ensembl cytogenetic band: 2p22.2 HGNC cytogenetic band: 2p22

QPCT Gene in genomic location: bands according to Ensembl, locations according to (and/or Entrez Gene and/or Ensembl if different)

GeneLoc gene densities for chromosome 2 GeneLoc Exon Structure

GeneLoc location for GC02P037483: (about GC identifiers)

Start:	37,425,241 bp from pter
End:	37,453,969 bp from pter
Size:	28,729 bases
Orientation:	plus strand

RefSeq DNA sequence:

NC_000002.10 NT_022184.14

(According to ¹UniProtKB, and/or Ensembl, Phosphorylation sites according to ²Phosphosite, RefSeq according to NCBI, PDB rendering according to OCA and/or Proteopedia, Recombinant Proteins from Invitrogen, Millipore, Sigma-Aldrich, R&D Systems, Enzo Life Sciences, Abnova, OriGene and/or, Abcam,
Biochemical Assays by Invitrogen, Millipore, R&D Systems, Cell Signaling Technology, and/or Enzo Life Sciences, Ontologies according to Gene Ontology Consortium 01 Apr 2009 and Entrez Gene, Antibodies by Invitrogen, Millipore, Sigma-Aldrich, R&D Systems, Cell Signaling Technology, Abcam, Abnova, and/or Novus Biologicals)
About This Section

UniProtKB/Swiss-Prot: QPCT_HUMAN, Q16769 (See protein sequence)

Recommended Name: Glutaminyl-peptide cyclotransferase precursor

Size: 361 amino acids; 40877 Da

Cofactor: Binds 1 zinc ion per subunit

PDB structures from and Proteopedia :

1MOI (3D)

2AFM (3D)

2AFO (3D)

2AFS (3D)

2AFU (3D)

2AFW (3D)

2AFX (3D)

2AFZ (3D)

2ZED (3D)

2ZEE (3D)

2ZEF (3D)

2ZEG (3D)

2ZEH (3D)

2ZEL (3D)

2ZEM (3D)

2ZEN (3D)

2ZEO (3D)

2ZEP (3D)

Secondary accessions: Q16770 Q3KRG6

REFSEQ proteins: NP_036545.1

ENSEMBL proteins:

ENSP00000385391 ENSP00000344829

Human Recombinant Proteins

	Browse Drug Discovery Central at Invitrogen for human recombinant proteins
	Browse Purified and Recombinant Proteins at Millipore
	Browse Human Recombinant Proteins at Sigma-Aldrich
	Browse R&D Systems for human recombinant proteins
	Browse recombinant and purified proteins available from Enzo Life Sciences
	Recombinant Proteins from Abcam (QPCT)
	Human Recombinant Proteins from Abnova (QPCT)

OriGene Purified Recombinant Human Protein: QPCT

Antibodies for QPCT:

	Browse Antibodies Central at Invitrogen
	Browse Millipore's Extensive Line of Mono- and Polyclonal Antibodies
	Sigma-Aldrich Antibodies for QPCT
	Browse R&D Systems for Antibodies
	Antibodies from Abcam (QPCT), each with their Abpromise^SM
	Monoclonal and Polyclonal Antibodies from Abnova (QPCT)
	Novus Biologicals Antibodies for QPCT

Assays for QPCT:

	Browse Invitrogen for biochemical assays
	Browse Kits and Assays available from Millipore
	Browse R&D Systems for biochemical assays
	Browse biochemical assays available from Enzo Life Sciences

(According to InterPro, ProtoNet, UniProtKB, and/or BLOCKS, Sets of similar genes according to GeneDecks)
About This Section

Graphical View of Domain Structure for InterPro Entry Q16769

ProtoNet protein and cluster: Q16769

UniProtKB/Swiss-Prot: QPCT_HUMAN, Q16769

Similarity: Belongs to the glutaminyl-peptide cyclotransferase family

(According to MGI Jun 06 2009, UniProtKB, IUBMB,and/or Genatlas,
shRNA from OriGene, Sigma-Aldrich, RNAi from Sigma-Aldrich,
RNAi Products, Clones, and Q-PCR Products from Invitrogen, Millipore, OriGene, and/or Abnova, siRNAs from Applied Biosystems, SYBR primers from OriGene, Cell-based Assays from Millipore, Ontologies according to Gene Ontology Consortium 01 Apr 2009 via Entrez Gene.)
About This Section

Inhib. RNA:		Invitrogen RNAi Products for gene knock-down (QPCT)
		Browse for Gene Knock-down Tools from Millipore
		Abnova Chimera RNAi Products for Gene knock-down (QPCT)

OriGene 29mer shRNA kit in GFP-retroviral vector: NM_012413

Applied Biosystems Silencer® siRNAs for QPCT

Sigma-Aldrich siRNA and siRNA Panels for QPCT

Sigma-Aldrich shRNA for QPCT

Explore Sigma-Aldrich super-pooled esiRNAs

Clones:		Invitrogen Clones for QPCT
		Browse Clones for the Expression of Recombinant Proteins Available from Millipore

OriGene GFP tagged cDNA clone in CMV expression vector: NM_012413
Myc/DDK tagged cDNA clone in CMV expression vector: NM_012413
untagged cDNA clone in CMV expression vector: NM_012413

Primers:

Browse Quantitative PCR Central at Invitrogen for Q-PCR LUX™ Primers

OriGene genome-wide validated SYBR primer pairs: NM_012413

UniProtKB/Swiss-Prot: QPCT_HUMAN, Q16769

Function: Responsible for the biosynthesis of pyroglutamyl peptides. Has a bias against acidic and

tryptophan residues adjacent to the N-terminal glutaminyl residue and a lack of importance of

chain length after the second residue. Also catalyzes N-terminal pyroglutamate formation. In

vitro, catalyzes pyroglutamate formation of N-terminally truncated form of APP amyloid-beta

peptides [Glu-3]-beta-amyloid. May be involved in the N-terminal pyroglutamate formation of

several amyloid-related plaque-forming peptides

Catalytic activity: L-glutaminyl-peptide = 5-oxoprolyl-peptide + NH(3)

Enzyme Number (IUBMB): EC 2.3.2.5

5/6 Gene Ontology (GO) molecular function terms (links to tree view) (see all 6 ):

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0008233	peptidase activity	IEA	--
GO:0008270	zinc ion binding	IEA	--
GO:0008415	acyltransferase activity	IEA	--
GO:0016603	glutaminyl-peptide cyclotransferase activity	IEA	--
GO:0016740	transferase activity	IEA	--

About this table

(Pathways according to Invitrogen (maps by GeneGo), Millipore, Cell Signaling Technology, Sigma-Aldrich, KEGG and/or UniProtKB,
Sets of similar genes according to GeneDecks, Proteins Network according to SABiosciences, Interactions according to ¹UniProtKB, ²MINT, and/or ³STRING, with links to IntAct and Ensembl, Ontologies according to Gene Ontology Consortium 01 Apr 2009 via Entrez Gene.)
About This Section

Gene Network Central^TM Interacting Genes and Proteins Network for QPCT

5/6 Interacting proteins for QPCT (ENSP00000344829³) via UniProtKB, MINT, and/or STRING (see all 6 )

Interactant		Interaction Details
GeneCard	External ID(s)	Interaction Details
DHPS	ENSP00000210060³	STRING (score=.846)
LINGO4	ENSP00000357810³	STRING (score=.844)
CCNDBP1	ENSP00000300213³	STRING (score=.776)
UBN1	ENSP00000262376³	STRING (score=.728)
TEAD1	ENSP00000334754³	STRING (score=.724)

About this table

2 Gene Ontology (GO) biological process terms (links to tree view):

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0006464	protein modification process	TAS	7999256
GO:0006508	proteolysis	IEA	--

About this table

(Chemical Compounds according to UniProtKB, Enzo Life Sciences, Sigma-Aldrich, Tocris Bioscience, and/or Novoseek and Drugs according to Enzo Life Sciences and/or PharmGKB)
About This Section

Browse drugs & compounds from Enzo Life Sciences

Browse Small Molecules at Sigma-Aldrich

Browse Tocris compounds for QPCT
3 Novoseek chemical compound relationships for QPCT gene

Compound	Score	Articles	PubMed IDs for Articles with Shared Sentences (# sentences)
pyroglutamic acid	92.05	14	17005457 (2), 16084398 (1), 1982622 (1), 16135565 (1) (see all 10)
zinc	43.60	6	15028118 (3), 16084398 (2), 11551224 (1)
histidine	33.05	4	11551224 (2), 16084398 (1), 10092499 (1)

About this table

(GenBank/EMBL/DDBJ Accessions according to
Unigene (Build 219 Homo sapiens; Jun 2 2009) or GenBank,
RefSeq according to Entrez Gene,
DOTS (version 10), and/or AceView,
non coding RNAs according to RNAdb,
ESTs according to GeneTide,
exon structure from GeneLoc, alternative splicing isoforms according to ASD and/or ECgene,
RNAi Products from Invitrogen, Millipore, and/or Abnova,
siRNAs from Applied Biosystems, Sigma-Aldrich,
shRNA from Sigma-Aldrich, OriGene,
Tagged/untagged cDNA clones from OriGene,
Expression Assays from Applied Biosystems)
About This Section

Inhib. RNA:		Invitrogen RNAi Products for gene knock-down (QPCT)
		Browse for Gene Knock-down Tools from Millipore
		Abnova Chimera RNAi Products for Gene knock-down (QPCT)

OriGene 29mer shRNA kit in GFP-retroviral vector: NM_012413

Sigma-Aldrich siRNA and siRNA Panels for QPCT

Sigma-Aldrich shRNA for QPCT

Explore Sigma-Aldrich super-pooled esiRNAs

Applied Biosystems Silencer® siRNAs:

NM_012413

REFSEQ mRNAs for QPCT gene:

NM_012413.3

Applied Biosystems TaqMan ® Gene Expression Assays:

NM_012413

              OriGene GFP tagged cDNA clone in CMV expression vector: NM_012413
                                 Myc/DDK tagged cDNA clone in CMV expression vector: NM_012413
                                 untagged cDNA clone in CMV expression vector: NM_012413

Additional cDNA sequence:

AB062487.1 AK290605.1 BC015460.1 BC020456.1 BC036721.1 BC047756.2 CR596088.1 CR608310.1

CR620857.1 CR623589.1 X67731.1 X71125.1

7 DOTS entries:

DT.92471540 DT.95101779 DT.95101775 DT.95101774 DT.100028509 DT.100782140 DT.91851733

24/159 AceView cDNA sequences (see all 159 ):

BM546830 CR620857 AV708489 BI859176 CR608310 BU076674 BQ676432 AI280994

BG104484 AA262267 CR596088 BU072918 BQ776785 BF971457 BX107292 BQ436787

BU179821 BF724762 AV705924 BF970452 AA489708 BF969845 CR623589 BF691241

highest scoring ESTs for QPCT:

AI955650 AA262878 AA282134 AA489708 AA745605 AA809678 AA915986 AA922311 AB062487 AI151392

Unigene Cluster for QPCT:

Glutaminyl-peptide cyclotransferase

Hs.79033 [show with all ESTs]

Unigene Representative Sequence: NM_012413

GeneLoc Exon Structure

5/6 Alternative Splicing Database (ASD) splice patterns (SP) for QPCT (see all 6 )

ExUns:	1	^	2a	·	2b	·	2c	^	3	^	4a	·	4b	^	5a	·	5b	·	5c	^	6a	·	6b	^	7	^	8	^	9a	·	9b
SP1:											-		-						-
SP2:									-		-		-						-
SP3:			-		-		-				-		-						-
SP4:											-
SP5:											-		-		-				-

About this scheme

ECgene alternative splicing isoforms for QPCT

2 Ensembl transcripts including schematic representations:

ENST00000404976 ENST00000338415

(Experimental results according to ¹GeneNote and GNF BioGPS,
probe sets-to-genes annotations according to ²GeneAnnot , ³GeneTide , Sets of similar genes according to GeneDecks, Electronic Northern calculations according to data from UniGene (Build 219 Homo sapiens), SAGE tags according to CGAP, plus additional links to SOURCE, and/or GNF BioGPS, and/or EXPOLDB, and/or UniProtKB,
Expression Assays from Applied Biosystems )
About This Section

QPCT expression in normal and diseased human tissues

Applied Biosystems TaqMan ® Gene Expression Assays for QPCT

¹ / ² / ³

4 probe-sets matching QPCT gene

Affymetrix probe-set	Array	# genes	Sensitivity	Specificity	Correlation	Length	Gb_Accession	Consensus	Uniqueness	Score	Rank
Affymetrix probe-set	Array	GeneAnnot data			GeneNote data		GeneTide data
91301_at²^{, 3}	U95-E	1	1.00	1.00	0.99	1.01	AI955650	1.00	1.00	1.00	1
35966_at²^{, 3}	U95-A	1	1.00	1.00	0.99	0.99	X71125	0.80	1.00	0.91	1
205174_s_at²^{, 3}	U133-A	1	1.00	1.00	--	--	NM_012413	0.60	1.00	0.82	1
205174_s_at²	U133Plus2	1	1.00	1.00	--	--	--	--	--	--	--

GeneDecks QPCT for binary patterns associated with the probe-sets selected above

About GeneDecksing

About this table

Data from

(Publications) and GNF BioGPS
About these images
About these images

CGAP SAGE TAG: TAGTTTGAAG

SOURCE GeneReport for Unigene cluster: Hs.79033
Expression variation in blood from EXPOLDB for QPCT

(Orthologs according to ^1,2HomoloGene (²older version, for species not in ¹newer version), ³euGenes, ⁴SGD and/or ⁵MGI Jun 06 2009, with possible further links to Flybase and/or WormBase, Gene Trees according to Ensembl)
About This Section

Orthologs for QPCT gene from 5/14 species (see all 14 )

Organism	Gene	Locus	Description	Human Similarity	NCBI accessions
dog (Canis familiaris)	QPCT¹	--	glutaminyl-peptide cyclotransferase	88.75(n) 89.06(a)	403861 XM_532934.2 XP_532934.2
chimpanzee (Pan troglodytes)	QPCT¹	--	glutaminyl-peptide cyclotransferase	99.26(n) 99.17(a)	744549 XM_001167364.1 XP_001167364.1
cow (Bos taurus)	QPCT¹	--	glutaminyl-peptide cyclotransferase	89.01(n) 86.43(a)	281437 NM_177506.2 NP_803472.1
rat (Rattus norvegicus)	RGD1562284¹	--	similar to Glutaminyl-peptide cyclotransferase precursor more	83.1(n) 83.38(a)	313837 XM_233812.4 XP_233812.2
mouse (Mus musculus)	Qpct^{1, 5}	17⁵	glutaminyl-peptide cyclotransferase (glutaminyl cyclase)^{1, 5}	82.83(n)¹ 82.27(a)¹	70536¹ NM_027455.2¹ NP_081731.1¹ AK017598⁵ AK037197⁵ (see all 12)

About this table Species with no ortholog for QPCT

ENSEMBL Gene Tree for QPCT

(Paralogs according to ¹HomoloGene
and ²Ensembl, Pseudogenes according to ³Pseudogene.org)
About This Section

Paralogs for QPCT gene

QPCTL²

(According to the ¹NCBI SNP Database, ²Ensembl, ³PupaSUITE, and UniProtKB, Linkage Disequilibrium by HapMap, Genotyping Reagents from Applied Biosystems)
About This Section

10/256 NCBI SNPs in QPCT are shown (see all 256 )
(Click for Applied Biosystems TaqMan ® Genotyping Assay) (see all 87)

AB	SNP ID	Valid	Chr 2 pos	Sequence	Recs	AA Chg	Type	More	Recs	Allele freq	Pop	Total sample	More
AB	Genomic Data				Transcription Data				Allele Frequencies
				--	--	--		--	--				--
--	rs6544071^1,2	A,C,F,H	37454297(+)	`cctacG/Aagtga`	1	--	ng3¹		8		EU EA WA	800
	rs4670697^1,2	A,C,F,H	37453492(+)	`GTTTAG/TGATAA`	1	--	ut3¹		8		EU EA WA	824
--	rs6708310^1,2	A,C,F,H	37453931(+)	`TGTAGA/GGTTAA`	1	--	ut3¹ ese³		4		EU EA WA	408
	rs2255991^1,2	A,C,F,H	37433475(-)	`TTGCCG/AAAGAG`	1	W/R	mis¹ ese³		13		EU EA WA NA	1334
--	rs6753973^1,2	A,F,H	37423536(+)	`TTGACC/GGTCTT`	1	--	ng5¹		4		EU EA WA	418
	rs6544068^1,2	A,C,F	37424801(+)	`AAGATT/CCACTG`	1	--	ng5¹		6		EA NA MN	1468
--	rs3755192^1,2	C,F,H	37423535(-)	`AGACGG/ATCAAA`	1	--	ng5¹		7		EU EA WA NA	558
	rs4670696^1,2	C,F,H	37453467(+)	`TCTTCA/CTTTGT`	1	H/P	mis¹		5		EU EA WA NA	480
--	rs6544070^1,2	A,C	37454258(+)	`tcttgA/Gcctcc`	1	--	ng3¹		0	--	--	--	--
	rs10170761^1,2	A,C	37424042(+)	`GAGCTG/TTATTG`	1	--	ng5¹		0	--	--	--	--

About this table

HapMap Linkage Disequilibrium images for QPCT (up to first 250kb)

(in which this Gene is Involved, According to OMIM, UniProtKB, Novoseek, PharmGKB, Genatlas, GeneTests, Blood group antigen gene mutations by BGMUT, HGMD, GAD, HuGE Navigator, BCGD, and/or TGDB.)
About This Section

OMIM: 607065

2 Novoseek disease relationships for QPCT gene

Disease	Score	Articles	PubMed IDs for Articles with Shared Sentences (# sentences)
osteoporosis	23.01	1	15231017 (1)
tumors	0.00	2	16820060 (2)

About this table

Genetic Association Database: QPCT
Human Genome Epidemiology Navigator: QPCT (3 documents)

(Possibly Related Articles in Doctor's Guide)
About This Section

(in PubMed. Associations of this gene to articles via ¹Novoseek, ²HGNC, ³Entrez Gene, ⁴UniProtKB/Swiss-Prot, ⁵UniProtKB/TrEMBL, ⁶GAD, and/or ⁷PharmGKB)
About This Section

10/36 PubMed articles for QPCT gene (see all 36 ):

Molecular cloning, sequence analysis and expression of human pituitary glutaminyl cyclase. (PubMed id 7999256)^{1, 2, 3, 4} Song I.... Bateman R.C. Jr. (1994)
Association of multiple nucleotide variations in the pituitary glutaminyl cyclase gene (QPCT) with low radial BMD in adult women. (PubMed id 15231017)^{1, 3, 6} Ezura Y....Emi M. (2004)
Crystal structures of human glutaminyl cyclase, an enzyme responsible for protein N-terminal pyroglutamate formation. (PubMed id 16135565)^{1, 3, 4} Huang K.-F.... Wang A.H.-J. (2005)
The status, quality, and expansion of the NIH full-length cDNA project: the Mammalian Gene Collection (MGC). (PubMed id 15489334)^{3, 4} Gerhard D.S....Malek J. (2004)
Heterologous expression and characterization of human glutaminyl cyclase: evidence for a disulfide bond with importance for catalytic activity. (PubMed id 12196024)^{1, 3} Schilling S....Demuth H.U. (2002)
Identification of human glutaminyl cyclase as a metalloenzyme. Potent inhibition by imidazole derivatives and heterocyclic chelators. (PubMed id 14522962)^{1, 3} Schilling S....Demuth H.U. (2003)
Isolation of an isoenzyme of human glutaminyl cyclase: retention in the Golgi complex suggests involvement in the protein maturation machinery. (PubMed id 18486145)^{1, 3} Cynis H....Schilling S. (2008)
Glutaminyl cyclases unfold glutamyl cyclase activity under mild acid conditions. (PubMed id 15063747)^{1, 4} Schilling S.... Demuth H.U. (2004)
Evidence for essential histidines in human pituitary glutaminyl cyclase. (PubMed id 11551224)^{1, 3} Bateman R.C....Booth R.E. (2001)
A conserved hydrogen-bond network in the catalytic centre of animal glutaminyl cyclases is critical for catalysis. (PubMed id 18072935)^{1, 3} Huang K.F....Wang A.H. (2008)

(in PubMed, OMIM, and NCBI Bookshelf)
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		AND	OR
Aliases
Free Text

Query String		PubMed OMIM NCBI Bookshelf
	(Note: In FireFox, select the above section and copy using Ctrl-C)

(According to Entrez Gene, HGNC, AceView, euGenes, Ensembl, miRBase, ECgene, and/or H-InvDB)
About This Section

Entrez Gene: 25797	HGNC: 9753	AceView: QPCT	Ensembl:ENSG00000115828	euGenes: HUgn25797
ECgene: QPCT	H-InvDB: QPCT

(According to HUGE)
About This Section

(According to ATLAS, HORDE, IMGT, MTDB, LEIDEN, UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL,
Wikipedia and/or GeneReviews via UniProtKB/Swiss-Prot)
About This Section

Licensable Technologies for QPCT
(Available from WIS Yeda, Salk, Tufts)

About This Section

(Reagents available from Applied Biosystems, Antibodies and assays by Cell Signaling Technology, Abcam, Novus Biologicals,
Sigma-Aldrich, R&D Systems, Millipore, Abnova, and/or Invitrogen, Clones available from OriGene,and/or Invitrogen, Drugs and/or compounds by Sigma-Aldrich,
Enzo Life Sciences, and/or Tocris Bioscience)
About This Section

Products for QPCT:

TaqMan ® Gene Expression Assays

TaqMan ® Genotyping Assays

Free SNP selection tool

Invitrogen iPath Pathways	Invitrogen BLOCK-iT™ RNAi
Invitrogen Antibodies	Invitrogen Assays
Invitrogen Clones	Invitrogen Q-PCR Products
Invitrogen Human Recombinant Kinases	Invitrogen Custom Antibody and Peptide Service
Invitrogen Proteins / Assays / Screening Services	Search Invitrogen catalog for QPCT-related products

	Millipore Custom Antibody & Bulk Services
	Millipore Preclinical / Clinical Development Services
	Millipore Immunoassay Services
	Millipore Target Screening & Profiling Services

	Predesigned and custom siRNAs for QPCT		Antibodies for QPCT
	Explore super-pooled esiRNAs		Browse proteins at Sigma-Aldrich
	Lentivirus-delivered shRNAs for QPCT		Browse small molecules at Sigma-Aldrich
	"Your Favorite Gene" Pathways		feedback

	Antibodies		Primer Pairs
	Cell Culture Products		ELISAs
	Flow Cytometry Kits		Protease Activity Assays & Reagents
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	Multiplex/Array Assay Kits & Reagents		ELISpot Kits & Development Modules
	Recombinant & Natural Proteins

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	Tagged/untagged cDNA clones
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	Purified proteins from HEK293T cells
	Western blot validated antibodies
	shRNA in GFP-retroviral vector

Search Tocris compounds for QPCT

Search www.enzolifesicences.com for proteins, assays, substrates, inhibitors & antibodies

Antibodies for QPCT

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QPCT Gene

protein-coding GIFtS: 59

Aliases &
Descriptions
for QPCT

Summaries
for QPCT

Genomic Location
for QPCT

Proteins
for QPCT

Protein Domains/
Families
for QPCT

Gene Function
for QPCT

Pathways & Interactions
for QPCT

Drugs & Compounds
for QPCT

Transcripts
for QPCT

Expression
for QPCT

Orthologs
for QPCT

Paralogs
for QPCT

SNPs/Variants
for QPCT

Disorders & Mutations
for QPCT

Medical News
for QPCT

Publications
for QPCT

Search for QPCT

Genome Databases showing QPCT

Other Databases showing QPCT

Specialized Databases showing QPCT

Licensable Technologies
for QPCT

Services
for QPCT

Gene Index: 3 5 A B C D E F G H I J K L M N O P Q R S T U V W X Y Z

QPCT Gene

protein-coding GIFtS: 59

Aliases & Descriptionsfor QPCT

Summariesfor QPCT

Genomic Locationfor QPCT

Proteinsfor QPCT

Protein Domains/Familiesfor QPCT

Gene Functionfor QPCT

Pathways & Interactionsfor QPCT

Drugs & Compoundsfor QPCT

Transcriptsfor QPCT

Expression for QPCT

Orthologsfor QPCT

Paralogsfor QPCT

SNPs/Variantsfor QPCT

Disorders & Mutationsfor QPCT

Medical Newsfor QPCT

Publicationsfor QPCT

Search for QPCT

Genome Databases showing QPCT

Other Databases showing QPCT

Specialized Databases showing QPCT

Licensable Technologiesfor QPCT

Servicesfor QPCT

Gene Index: 3 5 A B C D E F G H I J K L M N O P Q R S T U V W X Y Z

Aliases &
Descriptions
for QPCT

Summaries
for QPCT

Genomic Location
for QPCT

Proteins
for QPCT

Protein Domains/
Families
for QPCT

Gene Function
for QPCT

Pathways & Interactions
for QPCT

Drugs & Compounds
for QPCT

Transcripts
for QPCT

Expression
for QPCT

Orthologs
for QPCT

Paralogs
for QPCT

SNPs/Variants
for QPCT

Disorders & Mutations
for QPCT

Medical News
for QPCT

Publications
for QPCT

Licensable Technologies
for QPCT

Services
for QPCT