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Category Symbol Source: HGNC EntrezGene Ensembl GeneCards RNA genes CroW21

GIFtS
-

PLAUR Gene

protein-coding GIFtS: 68
GCID: GC19M044150

plasminogen activator, urokinase receptor

Explore 156 diseases affiliated with
PLAUR via

MalaCards

our new
Human Malady Compendium

(According to ¹HGNC, ²Entrez Gene,
³UniProtKB/Swiss-Prot, ⁴UniProtKB/TrEMBL, ⁵OMIM, ⁶GeneLoc, ⁷Ensembl, ⁸DME, ⁹miRBase, and/or ¹⁰fRNAdb)
About This Section

Aliases
Plasminogen Activator, Urokinase Receptor¹ ²		U-Plasminogen Activator Receptor Form 2²
UPAR¹ ² ³		Urokinase Plasminogen Activator Surface Receptor²
URKR¹ ² ⁵		Urokinase-Type Plasminogen Activator (UPA) Receptor²
CD87¹ ²		MO3³
Monocyte Activation Antigen Mo3² ³		CD87 Antigen³
U-PAR² ³		UPAR¹ ² ³

External Ids:	HGNC: 9053¹	Entrez Gene: 5329²	Ensembl: ENSG00000011422⁷	OMIM: 173391⁵	UniProtKB: Q03405³

Export aliases for PLAUR gene to outside databases

Previous GC identifers: GC19M044796 GC19M044542 GC19M048826 GC19M048844 GC19M048842 GC19M040580

(According to Entrez Gene, Tocris Bioscience, Wikipedia's Gene Wiki, PharmGKB,
UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL)
About This Section

Entrez Gene summary for PLAUR:

This gene encodes the receptor for urokinase plasminogen activator and, given its role in localizing and promoting

plasmin formation, likely influences many normal and pathological processes related to cell-surface plasminogen

activation and localized degradation of the extracellular matrix. It binds both the proprotein and mature forms of

urokinase plasminogen activator and permits the activation of the receptor-bound pro-enzyme by plasmin. The protein

lacks transmembrane or cytoplasmic domains and may be anchored to the plasma membrane by a

glycosyl-phosphatidylinositol (GPI) moiety following cleavage of the nascent polypeptide near its carboxy-terminus.

However, a soluble protein is also produced in some cell types. Alternative splicing results in multiple transcript

variants encoding different isoforms. The proprotein experiences several post-translational cleavage reactions that

have not yet been fully defined. (provided by RefSeq, Jul 2008)

UniProtKB/Swiss-Prot: UPAR_HUMAN, Q03405

Function: Acts as a receptor for urokinase plasminogen activator. Plays a role in localizing and promoting plasmin

formation. Mediates the proteolysis-independent signal transduction activation effects of U-PA. It is subject to

negative-feedback regulation by U-PA which cleaves it into an inactive form

Gene Wiki entry for PLAUR (Urokinase receptor)

(According to GeneLoc and/or HGNC, and/or
Entrez Gene (NCBI build 37),
and/or miRBase,
Genomic Views according to UCSC (hg19) and Ensembl (release 69), Regulatory elements and Epigenetics data according to QIAGEN, SABiosciences, and/or SwitchGear Genomics)
About This Section

RefSeq DNA sequence:

NC_000019.9 NC_018930.1 NT_011109.16

Regulatory elements:

SABiosciences Regulatory transcription factor binding sites in the PLAUR gene promoter:
STAT1 Sp1 STAT3
Other transcription factors

SwitchGear Promoter luciferase reporter plasmids (see all 2): PLAUR promoter sequence

Search SABiosciences Chromatin IP Primers for PLAUR

Epigenetics:

QIAGEN PyroMark CpG Assay predesigned Pyrosequencing DNA Methylation assays in human, mouse, rat PLAUR

Genomic Location:
Genomic View: UCSC Golden Path with GeneCards custom track

Entrez Gene cytogenetic band: 19q13 Ensembl cytogenetic band: 19q13.31 HGNC cytogenetic band: 19q13

PLAUR Gene in genomic location: bands according to Ensembl, locations according to (and/or Entrez Gene and/or Ensembl if different)
PLAUR gene location

GeneLoc information about chromosome 19 GeneLoc Exon Structure

GeneLoc location for GC19M044150: view genomic region (about GC identifiers)

Start:	44,150,248 bp from pter	End:	44,174,502 bp from pter
Size:	24,255 bases	Orientation:	minus strand

(According to ¹UniProtKB, HORDE, neXtProt, Ensembl, and/or Reactome, Modification sites according to ²PhosphoSitePlus, Specific Peptides from DME, Protein expression images according to data from SPIRE MOPED and PaxDb, RefSeq according to NCBI, PDB rendering according to OCA and/or Proteopedia, Recombinant Proteins from EMD Millipore, R&D Systems, GenScript, Enzo Life Sciences, OriGene, Novus Biologicals, Sino Biological, ProSpec, and/or Uscn,
Biochemical Assays by EMD Millipore, R&D Systems, OriGene, GenScript, Cell Signaling Technology, Enzo Life Sciences, and/or Uscn, Ontologies according to Gene Ontology Consortium 01 Mar 2013 and Entrez Gene, Antibodies by EMD Millipore, R&D Systems, GenScript, Cell Signaling Technology, OriGene, Novus Biologicals, Thermo Fisher Scientific, Abcam, and/or Uscn)
About This Section

UniProtKB/Swiss-Prot: UPAR_HUMAN, Q03405 (See protein sequence)

Recommended Name: Urokinase plasminogen activator surface receptor precursor

Size: 335 amino acids; 36978 Da

Subunit: Monomer (Probable). Interacts with MRC2. Interacts (via the UPAR/Ly6 domains) with SRPX2

Subcellular location: Isoform 1: Cell membrane; Lipid-anchor, GPI-anchor

Subcellular location: Isoform 2: Secreted (Probable)

6/7 PDB 3D structures from and Proteopedia for PLAUR (see all 7):

1YWH (3D)

2FD6 (3D)

2I9B (3D)

3BT1 (3D)

3BT2 (3D)

3U73 (3D)

Secondary accessions: A8K409 Q12876 Q15845 Q16887 Q6IB52 Q9BWT0 Q9NYC8 Q9UD69 Q9UEA6 Q9UM92 Q9UMV0

Alternative splicing: 3 isoforms: Q03405-1 Q03405-2 Q03405-3

Explore the universe of human proteins at neXtProt for PLAUR: NX_Q03405

Post-translational modifications:

View modification sites using PhosphoSitePlus²

View neXtProt modification sites for NX_Q03405

PLAUR Protein expression data from MOPED and PaxDb: About this image

REFSEQ proteins (3 alternative transcripts):

NP_001005376.1 NP_001005377.1 NP_002650.1

ENSEMBL proteins:

ENSP00000342049 ENSP00000339328 ENSP00000221264

Reactome Protein details: Q03405
Human Recombinant Protein Products for PLAUR:

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	Browse ProSpec Recombinant Proteins
	Uscn Proteins for PLAUR

Gene Ontology (GO): 5/9 cellular component terms (GO ID links to tree view) (see all 9): About this table

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0005576	extracellular region	IEA	--
GO:0005788	endoplasmic reticulum lumen	TAS	--
GO:0005789	endoplasmic reticulum membrane	TAS	--
GO:0005886	plasma membrane	TAS	--
GO:0005887	integral to plasma membrane	IEA	--

PLAUR for ontologies About GeneDecksing

PLAUR Antibody Products:

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Assay Products for PLAUR:

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	Uscn ELISAs and CLIAs for PLAUR

(According to InterPro, ProtoNet, UniProtKB, and/or BLOCKS, Sets of similar genes according to GeneDecks)
About This Section

PLAUR for domains About GeneDecksing

3 InterPro domains/families:

IPR018363 CD59_antigen_CS

IPR016054 LY6_UPA_recep-like

IPR001526 LY6_UPAR

Graphical View of Domain Structure for InterPro Entry Q03405

ProtoNet protein and cluster: Q03405

2 Blocks protein families:

IPB001526 CD59 antigen

IPB003631 Urokinase plasminogen activator surface receptor

UniProtKB/Swiss-Prot: UPAR_HUMAN, Q03405

Similarity: Contains 3 UPAR/Ly6 domains

(According to ¹UniProtKB, Genatlas, LifeMap Discovery™, IUBMB, and/or ²DME, Human phenotypes from GenomeRNAi, Animal models from MGI Mar 06 2013, inGenious Targeting Laboratory,
bound targets from SABiosciences, miRNA Gene Targets from miRTarBase, shRNA from OriGene, Sirion Biotech, RNAi from EMD Millipore, siRNAs from OriGene, QIAGEN, microRNA from QIAGEN, Gene Editing from DNA2.0, Sirion Biotech, Clones from EMD Millipore, OriGene, SwitchGear Genomics, GenScript, Sino Biological, DNA2.0, and Vector BioLabs, Cell Lines from GenScript, LifeMap BioReagents, Sirion Biotech, In Situ Hybridization Assays from Advanced Cell Diagnostics, Ontologies according to Gene Ontology Consortium 01 Mar 2013 via Entrez Gene.)
About This Section

Molecular Function:

UniProtKB/Swiss-Prot Summary: UPAR_HUMAN, Q03405

Function: Acts as a receptor for urokinase plasminogen activator. Plays a role in localizing and promoting plasmin

formation. Mediates the proteolysis-independent signal transduction activation effects of U-PA. It is subject to

negative-feedback regulation by U-PA which cleaves it into an inactive form

Genatlas biochemistry entry for PLAUR:

plasminogen activator urokinase receptor,significantly diminished in diffuse type gastric carcinoma

Gene Ontology (GO): 4 molecular function terms (GO ID links to tree view): About this table

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0004872	receptor activity	NAS	1689240
GO:0005515	protein binding	IPI	18718938
GO:0019899	enzyme binding	IPI	14688365
GO:0030377	U-plasminogen activator receptor activity	NAS	1689240

PLAUR for ontologies About GeneDecksing

Phenotypes:
6 GenomeRNAi human phenotypes for PLAUR:

Decreased G3BP1 protein expres	G0/1 arrest	Increased G1 DNA content	Increased cell death HMECs cel
Synthetic lethal with Ras	Upregulation of Wnt/beta-caten

12 MGI mutant phenotypes (inferred from 2 alleles) (MGI details for Plaur):

behavior/neurological	cardiovascular system	cellular	growth/size	hematopoietic system
homeostasis/metabolism	immune system	liver/biliary system	muscle	nervous system
renal/urinary system	respiratory system

PLAUR for phenotypes About GeneDecksing

Animal Models:
Mouse knock-outs for PLAUR: Plaur^tm1Jld Plaur^tm1Mlg

inGenious Targeting Laboratory - Customizable classic, inducible, and humanized mouse model solutions for PLAUR

miRNA Products:		miRTarBase miRNAs that target PLAUR: hsa-mir-204 (MIRT005839)

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		hsa-miR-3163 hsa-miR-4283 hsa-miR-544b hsa-miR-377 hsa-miR-3120-3p
		SwitchGear 3'UTR luciferase reporter plasmid: PLAUR 3' UTR sequence

Inhib. RNA Products:		Browse for Gene Knock-down Tools from EMD Millipore
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In Situ Assay
Products:

Advanced Cell Diagnostics RNAscope RNA in situ hybridization assays for PLAUR

(Pathways according to EMD Millipore, R&D Systems, Cell Signaling Technology, KEGG, PharmGKB, BioSystems, Reactome, Tocris Bioscience, GeneGo (Thomson Reuters), QIAGEN, and/or UniProtKB, Sets of similar genes according to GeneDecks, Interaction Networks according to SABiosciences, and/or STRING, Interactions according to ¹UniProtKB, ²MINT, ³I2D, and/or ⁴STRING, with links to IntAct and Ensembl, Ontologies according to Gene Ontology Consortium 01 Mar 2013 via Entrez Gene).
About This Section

Unified GeneCards pathways - 5/17 super-pathways (see all 17) About this table

See pathways by source

Super-pathway

contained gene-specific pathways

Cell adhesion_ECM remodeling

Cell adhesion ECM remodeling

1.00

Cell adhesion_ECM remodeling

1.00

Complement and coagulation cascades

Complement and coagulation cascades

1.00

Complement and Coagulation Cascades

0.72

Development_CNTF receptor signaling

Cell adhesion PLAU signaling

0.35

Cell adhesion_PLAU signaling

0.35

Asparagine N-linked glycosylation

Post-translational protein modification

0.44

Metabolism of proteins

0.15

Adhesion

Adhesion

1.00

Pathway sources
See GeneCards unified pathways
Show all pathways

2 EMD Millipore Pathways for PLAUR

	Cell adhesion PLAU signaling
	Cell adhesion ECM remodeling

3 Downloadable PowerPoint Slides of QIAGEN Pathway Central Maps for PLAUR

	Glioma Invasiveness
	UPA-UPAR Pathway
	WNT Signaling

1 Cell Signaling Technology (CST) Pathway for PLAUR

Adhesion

3 GeneGo (Thomson Reuters) Pathways for PLAUR

	Development VEGF signaling via VEGFR2 - generic cascades
	Cell adhesion PLAU signaling
	Cell adhesion ECM remodeling

5 BioSystems Pathways for PLAUR

	Complement and Coagulation Cascades
	amb2 Integrin signaling
	FGF signaling pathway
	Arf6 downstream pathway
	Validated transcriptional targets of AP1 family members Fra1 and Fra2

5/6 Reactome Pathways for PLAUR (see all 6)

	Hemostasis
	Attachment of GPI anchor to uPAR
	Dissolution of Fibrin Clot
	Metabolism of proteins
	Post-translational modification: synthesis of GPI-anchored proteins

1 Kegg Pathway (Kegg details for PLAUR):

Complement and coagulation cascades

PLAUR for pathways About GeneDecksing

Interactions:

SABiosciences Gene Network Central^TM Interacting Genes and Proteins Network for PLAUR

STRING Interaction Network Preview (showing 5 interactants - click image to see 25)

5/69 Interacting proteins for PLAUR (Q03405^{2, 3} ENSP00000339328⁴) via UniProtKB, MINT, STRING, and/or I2D (see all 69)

Interactant		Interaction Details
GeneCard	External ID(s)	Interaction Details
PFN2	P35080²^,³	MINT-64050 I2D: score=5
SURF2	Q15527²^,³, ENSP00000361032⁴	MINT-64051 I2D: score=4 STRING: ENSP00000361032
TYK2	P29597³, ENSP00000264818⁴	I2D: score=4 STRING: ENSP00000264818
IL6ST	P40189³, ENSP00000338799⁴	I2D: score=3 STRING: ENSP00000338799
LRP1	Q07954³, ENSP00000243077⁴	I2D: score=3 STRING: ENSP00000243077

About this table

Gene Ontology (GO): 5/16 biological process terms (GO ID links to tree view) (see all 16): About this table

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0006501	C-terminal protein lipidation	TAS	--
GO:0006928	cellular component movement	NAS	1689240
GO:0006935	chemotaxis	TAS	10749881
GO:0007165	signal transduction	TAS	10749881
GO:0007596	blood coagulation	TAS	--

PLAUR for ontologies About GeneDecksing

(Chemical Compounds according to UniProtKB, Enzo Life Sciences, EMD Millipore, Tocris Bioscience HMDB, BitterDB, and/or Novoseek, and Drugs according to DrugBank, Enzo Life Sciences, and/or PharmGKB, with drugs/clinical trials/news search links to CenterWatch)
About This Section

PLAUR for compounds About GeneDecksing

Browse Small Molecules at EMD Millipore

Browse drugs & compounds from Enzo Life Sciences

Browse Tocris compounds for PLAUR

5 DrugBank Compounds for PLAUR About this table

Compound	Synonyms	CAS #	Type	Actions	PubMed Ids
Urokinase	U-plasminogen activator (see all 3)	9039-53-6	target	--	14688365 12817472 17327908 12615913 14679304 14720519
Tenecteplase	t- PA (see all 4)	191588-94-0	target	--	12210068 11447759 15231724
Alteplase	t-PA (see all 4)	105857-23-6	target	--	17139284 17016423
Anistreplase	t- PA (see all 4)	81669-57-0	target	--	17139284 17016423
Reteplase	t- PA (see all 4)	133652-38-7	target	--	17139284 17016423

10/103 Novoseek chemical compound relationships for PLAUR gene (see all 103) About this table

Compound	-log (P-Val)	Hits	PubMed IDs for Articles with Shared Sentences (# sentences)
matrigel	77.5	40	10599442 (5), 15302576 (2), 8187097 (2), 10591631 (1) (see all 25)
kininogen	73.6	18	14764580 (2), 11414365 (1), 12944405 (1), 15044324 (1) (see all 13)
serine	63.4	81	12790317 (2), 18508598 (2), 9191470 (1), 9198168 (1) (see all 62)
aprotinin	51.5	11	11422178 (2), 7962113 (1), 8601593 (1), 11169951 (1) (see all 7)
glycolipid	49	23	1325906 (3), 7615819 (1), 8392005 (1), 7548000 (1) (see all 10)
p-amino benzamidine	47.9	3	9872599 (1), 11804183 (1)
mannose 6-phosphate	47.8	17	12665524 (2), 18220730 (1), 15085180 (1), 9566979 (1) (see all 9)
pimonidazole	46.3	4	14729600 (1), 15781654 (1), 11912164 (1)
vegf	44.1	102	16810742 (7), 12708473 (5), 7730348 (4), 16088182 (4) (see all 35)
epsilon aminocaproic acid	39	3	7962113 (1), 11169951 (1), 10090848 (1)

Search CenterWatch for drugs/clinical trials and news about PLAUR / UPAR

(Secondary structures according to fRNAdb,
GenBank/EMBL/DDBJ Accessions according to
Unigene (Build 235 Homo sapiens; Mar 10 2013) or GenBank,
RefSeq according to Entrez Gene,
DOTS (version 10), and/or AceView, transcript ids from Ensembl with links to UCSC,
exon structure from GeneLoc, alternative splicing isoforms according to ASD and/or ECgene,
RNAi Products from EMD Millipore,
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Tagged/untagged cDNA clones from OriGene, SwitchGear Genomics, GenScript, DNA2.0, Vector BioLabs, Primers from OriGene, SABiosciences, and/or QIAGEN )
About This Section

REFSEQ mRNAs for PLAUR gene (3 alternative transcripts):

NM_001005376.2 NM_001005377.2 NM_002659.3

Unigene Cluster for PLAUR:

Plasminogen activator, urokinase receptor

Hs.466871 [show with all ESTs]

Unigene Representative Sequence: BM554045

3 Ensembl transcripts including schematic representations, and UCSC links where relevant:

ENST00000339082(uc002oxd.2) ENST00000340093(uc002oxf.2) ENST00000221264(uc002oxg.2)

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		hsa-miR-3163 hsa-miR-4283 hsa-miR-544b hsa-miR-377 hsa-miR-3120-3p
		SwitchGear 3'UTR luciferase reporter plasmid: PLAUR 3' UTR sequence

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		GenScript: all cDNA clones in your preferred vector (see all 3): PLAUR (NM_002659)
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		QIAGEN QuantiTect SYBR Green Assays in human, mouse, rat PLAUR
		QIAGEN QuantiFast Probe-based Assays in human, mouse, rat PLAUR

Additional cDNA sequence:

AF257789.1 AK290774.1 AY029180.1 BC002788.2 CR456952.1 M83246.1 U08839.1 U09347.1

X51675.1 X74039.1

14 DOTS entries:

DT.454959 DT.99968496 DT.100797888 DT.97803353 DT.75101350 DT.95276935 DT.99930521 DT.121442407

DT.40228927 DT.91940708 DT.95257090 DT.75148085 DT.92431704 DT.95276939

24/289 AceView cDNA sequences (see all 289):

CR601067 BM767461 CA426359 CD364814 BM914922 CR606767 CK821016 F05423

BM673224 BF433323 CD370941 U09347 BM719074 CB148651 CD368062 AA379455

CF126247 AI570820 BQ647008 CA307585 BI793096 AL710258 CA307155 CD672247

GeneLoc Exon Structure

5/11 Alternative Splicing Database (ASD) splice patterns (SP) for PLAUR (see all 11) About this scheme

ExUns:	1a	·	1b	·	1c	·	1d	·	1e	^	2a	·	2b	^	3a	·	3b	·	3c	^	4	^	5a	·	5b	·	5c	^	6	^	7a	·	7b	^	8	^	9a	·	9b	^	10a	·	10b	·	10c	^	11a	·	11b
SP1:											-		-		-		-		-								-		-
SP2:											-		-		-		-		-								-		-												-		-		-
SP3:											-		-		-		-		-								-		-						-
SP4:											-		-		-		-		-								-		-								-		-
SP5:											-		-		-		-		-				-		-		-		-												-		-		-

ECgene alternative splicing isoforms for PLAUR

(RNA expression data according to H-InvDB, NONCODE, miRBase, and RNAdb, Expression images according to data from BioGPS, Illumina Human BodyMap, and CGAP SAGE, Sets of similar genes according to GeneDecks, in vivo and in vitro expression data from LifeMap Discovery™, plus additional links to Genevestigator, and/or SOURCE, and/or BioGPS, and/or UniProtKB,
PCR Arrays from SABiosciences, Primers from OriGene, SABiosciences, and/or QIAGEN, In Situ Hybridization Assays from Advanced Cell Diagnostics)
About This Section

PLAUR expression in normal human tissues (normalized intensities)
See probesets specificity/sensitivity at GeneAnnot
About this imageBioGPS
CGAP TAG: AACCACATTG

About this image

PLAUR expression in embryonic tissues and stem cells
Expression by the Database of Embryonic development, Stem cell research, and Regenerative medicine About this table

1 LifeMap In Vivo Development Anatomical Compartment/Cell

Tissue

Anatomical Compartment

Cell

Category (developmental path)

Adipose

Visceral White Adipose

Adipose

Expression:

Positive

Negative

Selective marker

Experimental details:

Curated

Microarrays

In-situ hybridization

Stem Cell Differentiation: 1 LifeMap Cell

Name

Category

PureStem™ progenitor EN13 (Embryonic Progenitor Cell)

See PLAUR Protein Expression from SPIRE MOPED and PaxDB
Genevestigator expression for PLAUR

SOURCE GeneReport for Unigene cluster: Hs.466871

UniProtKB/Swiss-Prot: UPAR_HUMAN, Q03405

Tissue specificity: Expressed in neurons of the rolandic area of the brain (at protein level). Expressed in the brain

SABiosciences Expression via Pathway-Focused PCR Arrays including PLAUR:

WNT Signaling Targets in human mouse rat

Wound Healing in human mouse rat

Tumor Metastasis in human mouse rat

Cell Motility in human mouse rat

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In Situ
Assay Products:

Advanced Cell Diagnostics RNAscope RNA in situ hybridization assays for PLAUR

(Orthologs according to ^1,2HomoloGene (²older version, for species not in ¹newer version), ³euGenes, ⁴SGD , ⁵MGI Mar 06 2013, with possible further links to Flybase and/or WormBase, and/or ⁶Ensembl pan taxonomic compara , Gene Trees according to Ensembl and TreeFam)
About This Section

This gene was present in the common ancestor of chordates.

Orthologs for PLAUR gene from 3/8 species (see all 8) About this table

Organism	Taxonomic classification	Gene	Description	Human Similarity	Orthology Type	Details
mouse (Mus musculus)	Mammalia	Plaur^{1 , 5}	plasminogen activator, urokinase receptor^{1, 5}	69.75(n)¹ 63.27(a)¹		7 (10.92 cM)⁵ 18793¹ NM_011113.3¹ NP_035243.1¹ 24462500⁵
lizard (Anolis carolinensis)	Reptilia	PLAUR⁶	--	33(a)	1 ↔ 1	LGf(966826-972159)
zebrafish (Danio rerio)	Actinopterygii	CABZ01063789.1⁶ BX927369.4⁶ (see all 9)	--	25(a) 25(a) (see all 9)	many → 1 many → 1 (see all 9)	17(49629200-49636494) 17(49721512-49724024)

ENSEMBL Gene Tree for PLAUR (if available)
TreeFam Gene Tree for PLAUR (if available)

(Paralogs according to ¹HomoloGene,
²Ensembl, and ³SIMAP, Pseudogenes according to ⁴Pseudogene.org Build 68)
About This Section

(SNPs/Variants according to the ¹NCBI SNP Database, ²Ensembl, ³PupaSUITE, UniProtKB, and DNA2.0, Linkage Disequilibrium by HapMap, Structural Variations(CNVs/InDels/Inversions) from the Database of Genomic Variants, Mutations from the Human Gene Mutation Database (HGMD) and the Locus Specific Mutation Databases (LSDB), Blood group antigen gene mutations by BGMUT, Resequencing Primers from QIAGEN, Cancer Mutation PCR Arrays and Assays and Copy Number PCR Arrays from SABiosciences)
About This Section

SNP ID	Valid	Clinical significance	Chr 19 pos	Sequence	#	AA Chg	Type	More	#	Allele freq	Pop	Total sample	More
Genomic Data					Transcription Related Data				Allele Frequencies
				--	--	--		--	--
rs11672362^1,2	C,H	--	40580485(+)	`tgatcC/Tgcccg`	1	--	ds500¹		1		NA	2
rs112271549^1,2	C	--	40580577(+)	`TGTAAA/GATTTG`	1	--	ds500¹		0	--	--	--	--
rs112395073^1,2	F	--	40581419(+)	`ACAGAC/TCTCAC`	1	--	int¹		3		CSA WA	122
rs75913969^1,2	C,F	--	40581449(+)	`GACAGG/TTGGTG`	1	--	int¹		4		CSA WA NA	242
rs79119737^1,2	C	--	40581769(+)	`CACCTT/CGGCCT`	1	--	int¹		2		WA CSA	3
rs113668131^1,2		--	40581818(+)	`CAGCCG/AAAGAT`	1	--	int¹		1		CSA	2
rs113246301^1,2		--	40581916(+)	`CATGAG/ATTTTT`	1	--	int¹		1		CSA	2
rs4251953^1,2	C,F,H	--	40582111(-)	`GAGACG/AGATAG`	1	--	int¹		21		NS EA NA WA	2286
rs4251937^1,2	C,F	--	40582273(-)	`GCCACC/TGTGCC`	1	--	int¹		2		NS	92
rs4251952^1,2	C,F	--	40582284(-)	`tacagG/Acatga`	1	--	int¹		2		NS	92

HapMap Linkage Disequilibrium report for PLAUR (44150248 - 44174502 bp)
Structural Variations (Copy Number Variations, Insertions/Deletions, Inversions)
Database of Genomic Variants (DGV): 1 variation for PLAUR
1 Indel: 102848
Human Gene Mutation Database (HGMD): PLAUR

SABiosciences Cancer Mutation PCR Assays

QIAGEN SeqTarget long-range PCR primers in human, mouse, rat for resequencing PLAUR

DNA2.0 Custom Variant and Variant Library Synthesis for PLAUR

(in which this Gene is Involved, According to MalaCards, OMIM, UniProtKB, the University of Copenhagen DISEASES database, Novoseek, Genatlas, GeneTests, GAD, HuGE Navigator, and/or TGDB.)
About This Section

PLAUR for disorders           About GeneDecksing

OMIM gene information: 173391    OMIM disorders: --

20/156 diseases for PLAUR (see all 156):    About MalaCards

paroxysmal nocturnal hemoglobinuria hepatocellular carcinoma (fibrolamellar variant) aids dementia complex pemphigus foliaceus

pemphigus vulgaris non-small cell lung carcinoma ureter carcinoma paranasal sinus disease

central nervous system disease pemphigus common variable immunodeficiency heart block

nervous system disease congenital heart block malignant pleural mesothelioma squamous cell carcinoma

giant cell tumor soft tissue sarcoma hemoglobinuria severe combined immunodeficiency

2 diseases from the University of Copenhagen DISEASES database for PLAUR:

Breast cancer Carcinoma

10/97 Novoseek disease relationships for PLAUR gene (see all 97) About this table

Disease	-log (P-Val)	Hits	PubMed IDs for Articles with Shared Sentences (# sentences)
metastasis	73.6	353	19050704 (5), 18399120 (5), 17264329 (4), 11912164 (4) (see all 99)
tumors	66.8	802	15172186 (7), 17549400 (6), 9102229 (6), 10090848 (6) (see all 99)
cancer	66.3	398	14595671 (5), 9137501 (4), 11223863 (3), 15279614 (3) (see all 99)
ureter carcinoma	63.3	2	15191676 (1), 14981816 (1)
breast cancer	55.1	180	14527916 (5), 10934293 (5), 12967140 (5), 7768629 (4) (see all 86)
glioblastoma	54.5	72	7747809 (5), 18819934 (5), 9219733 (4), 11234878 (3) (see all 23)
colon cancer	48.5	65	14977859 (6), 9191056 (4), 15004225 (3), 19782465 (3) (see all 23)
ovarian cancer	47.7	56	9581823 (3), 17891393 (3), 14563447 (3), 19395996 (3) (see all 24)
cancer invasiveness	47	5	14977859 (2), 11079728 (1), 9794194 (1)
paroxysmal hemoglobinuria nocturnal	45.1	12	9114600 (2), 1325906 (1), 15182554 (1), 18954937 (1) (see all 6)

Human Genome Epidemiology (HuGE) Navigator: PLAUR (19 documents)

Export disorders for PLAUR gene to outside databases

(in PubMed. Associations of this gene to articles via ¹Entrez Gene, ²UniProtKB/Swiss-Prot, ³HGNC, ⁴GAD, ⁵PharmGKB, ⁶HMDB, ⁷DrugBank, ⁸UniProtKB/TrEMBL, ⁹ Novoseek, and/or ¹⁰fRNAdb)
About This Section

PubMed articles for PLAUR gene, integrated from 9 sources (see all 1271):
(articles sorted by number of sources associating them with PLAUR)

Utopia: connect your pdf to the dynamic
world of online information

cDNA for Mo3, a monocyte activation antigen, encodes the human receptor for urokinase plasminogen activator. (PubMed id 1316922)^{1, 2, 9} Min H.Y.... Liu D.Y. (1992)
Crystal structure of the human urokinase plasminogen activator receptor bound to an antagonist peptide. (PubMed id 15861141)^{1, 2, 9} Llinas P.... Menez A. (2005)
Proteolytic regulation of the urokinase receptor/CD87 on monocytic cells by neutrophil elastase and cathepsin G. (PubMed id 14688365)^{1, 7, 9} Beaufort N....Pidard D. (2004)
Cell-surface acceleration of urokinase-catalyzed receptor cleavage. (PubMed id 9030717)^{1, 2, 9} Hoeyer-Hansen G.... Danoe K. (1997)
The structure of the urokinase-type plasminogen activator receptor gene. (PubMed id 8049431)^{1, 2, 9} Casey J.R.... Rosse W.F. (1994)
An alternatively spliced variant of mRNA for the human receptor for urokinase plasminogen activator. (PubMed id 8392005)^{1, 2, 9} Pyke C.... Danoe K. (1993)
Localization of the disulfide bonds in the NH2-terminal domain of the cellular receptor for human urokinase-type plasminogen activator. A domain structure belonging to a novel superfamily of glycolipid- anchored membrane proteins. (PubMed id 8394346)^{1, 2, 9} Ploug M.... Danoe K. (1993)
Epileptic and developmental disorders of the speech cortex: ligand/receptor interaction of wild-type and mutant SRPX2 with the plasminogen activator receptor uPAR. (PubMed id 18718938)^{1, 2} Royer-Zemmour B....Szepetowski P. (2008)
Identification of uPAR-positive chemoresistant cells in small cell lung cancer. (PubMed id 17327908)^{1, 7} Gutova M....Aboody K.S. (2007)
Modification-specific proteomics of plasma membrane proteins: identification and characterization of glycosylphosphatidylinositol- anchored proteins released upon phospholipase D treatment. (PubMed id 16602701)^{1, 2} Elortza F.... Jensen O.N. (2006)

(in PubMed, OMIM, and NCBI Bookshelf)
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		AND	OR
Aliases
Disorders
Free Text

Query String		PubMed OMIM NCBI Bookshelf
	(Note: In FireFox, select the above section and copy using Ctrl-C)

(According to Entrez Gene, HGNC, AceView, euGenes, Ensembl, miRBase, ECgene, Kegg, and/or H-InvDB)
About This Section

Entrez Gene: 5329	HGNC: 9053	AceView: PLAUR	Ensembl:ENSG00000011422	euGenes: HUgn5329
ECgene: PLAUR	Kegg: 5329	H-InvDB: PLAUR

(According to HUGE)
About This Section

(According to PharmGKB, ATLAS, HORDE, IMGT, LEIDEN, UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL,
Wikipedia and/or GeneReviews via UniProtKB/Swiss-Prot)
About This Section

Name	Description
PharmGKB entry for PLAUR	Pharmacogenomics, SNPs, Pathways
ATLAS Chromosomes in Cancer entry for PLAUR	Genetics and Cytogenetics in Oncology and Haematology
SeattleSNPs	http://pga.gs.washington.edu/data/plaur/

Intellectual Property for PLAUR gene
(Patent information from GeneIP, Licensable technologies from WIS Yeda, Salk, Tufts, IP news from LifeMap Sciences, Inc.)

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Patent Information for PLAUR gene:
Search GeneIP for patents involving PLAUR

GeneCards and IP:
Japan Patent Office Licenses GeneCards European Patent Office Licenses GeneCards Improving the IP Search

(Antibodies, recombinant proteins, and assays from EMD Millipore, R&D Systems, OriGene, QIAGEN, GenScript, Cell Signaling Technology, SABiosciences, Novus Biologicals, Sino Biological, Enzo Life Sciences, Abcam, ProSpec, Uscn, Thermo Fisher Scientific, Gene Editing from DNA2.0 and Sirion Biotech, Clones from EMD Millipore, OriGene, GenScript, Sino Biological, DNA2.0, SwitchGear Genomics, Vector BioLabs, Cell lines from GenScript, LifeMap BioReagents, and Sirion Biotech, PCR Arrays from SABiosciences, Drugs and/or compounds from EMD Millipore, Tocris Bioscience, and/or Enzo Life Sciences, In Situ Hybridization Assays from
Advanced Cell Diagnostics, Animal models from inGenious Targeting Laboratory)
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PLAUR Gene

plasminogen activator, urokinase receptor

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