NOL9 Gene
protein-coding GIFtS: 45
GCID: GC01M006581
|
|
nucleolar protein 9
| |
Aliases
for NOL9 gene
(According to
1HGNC,
2Entrez Gene,
3UniProtKB/Swiss-Prot,
4UniProtKB/TrEMBL, 5OMIM, 6GeneLoc,
7Ensembl,
8DME,
9miRBase,
and/or 10fRNAdb)
About This Section
|
| Aliases |
|---|
| Nucleolar Protein 91 2 3 | | Polynucleotide 5'-Hydroxyl-Kinase NOL92 | | Grc31 2 | | Polynucleotide 5'-Kinase2 | | NET61 2 | | EC 2.7.1.-3 | | FLJ233231 | | |
Export aliases for NOL9 gene to outside databasesPrevious GC identifers: GC01M006520 GC01M005729 |
Summaries
for NOL9 gene(According to Entrez Gene,
Tocris Bioscience,
Wikipedia's
Gene Wiki,
PharmGKB,
UniProtKB/Swiss-Prot,
and/or
UniProtKB/TrEMBL)
About This Section
|
UniProtKB/Swiss-Prot: NOL9_HUMAN, Q5SY16Function: Polynucleotide 5'-kinase involved in rRNA processing. The kinase activity is required for the processing ofthe 32S precursor into 5.8S and 28S rRNAs, more specifically for the generation of the major 5.8S(S) form. In vitro,has both DNA and RNA 5'-kinase activities. Probably binds RNA
Gene Wiki entry for NOL9
|
Genomic Views
for NOL9 gene
(According to
GeneLoc and/or
HGNC, and/or
Entrez Gene (NCBI build 37),
and/or miRBase,
Genomic Views according to
UCSC (hg19) and
Ensembl (release 69),
Regulatory elements and Epigenetics data according to
QIAGEN,
SABiosciences, and/or
SwitchGear Genomics)
About This Section
| RefSeq DNA sequence:- NC_000001.10 NC_018912.1 NT_021937.19
Regulatory elements:
 SABiosciences Regulatory transcription factor binding sites in the NOL9 gene promoter:
MyoD FOXO3 FOXO3b AREB6 POU2F1 POU2F1a FOXO3a PPAR-gamma2 Hlf ATF6
Other transcription factors
Search SABiosciences Chromatin IP Primers for NOL9
Epigenetics:
| | QIAGEN PyroMark CpG Assay predesigned Pyrosequencing DNA Methylation assays in human, mouse, rat NOL9 |
Genomic Location:
Genomic View: UCSC Golden Path with GeneCards custom track
Entrez Gene cytogenetic band: 1p36.31 Ensembl cytogenetic band: 1p36.31 HGNC cytogenetic band: 1p36.31NOL9 Gene in genomic location: bands according to Ensembl, locations according to
 (and/or Entrez Gene and/or Ensembl if different)
GeneLoc information about chromosome 1 GeneLoc Exon Structure GeneLoc location for GC01M006581: view genomic region
(about GC identifiers)
Start:
|
6,581,407 bp from pter |
End:
|
6,614,658 bp from pter |
Size:
|
33,252 bases |
Orientation:
|
minus strand |
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Proteins
for NOL9 gene
(According to
1UniProtKB,
HORDE,
neXtProt,
Ensembl,
and/or Reactome,
Modification sites according to 2PhosphoSitePlus,
Specific Peptides from DME,
Protein expression images according to data from SPIRE MOPED and PaxDb,
RefSeq according to NCBI,
PDB rendering according to OCA and/or Proteopedia,
Recombinant Proteins
from
EMD Millipore,
R&D Systems,
GenScript,
Enzo Life Sciences,
OriGene,
Novus Biologicals,
Sino Biological,
ProSpec, and/or
Uscn,
Biochemical Assays by
EMD Millipore,
R&D Systems,
OriGene,
GenScript,
Cell Signaling Technology,
Enzo Life Sciences, and/or
Uscn,
Ontologies according to Gene
Ontology Consortium 01 Mar 2013 and
Entrez Gene,
Antibodies by
EMD Millipore,
R&D Systems,
GenScript,
Cell Signaling Technology,
OriGene,
Novus Biologicals,
Thermo Fisher Scientific,
Abcam, and/or
Uscn)
About This Section
|
UniProtKB/Swiss-Prot: NOL9_HUMAN, Q5SY16 (See
protein sequence)Recommended Name: Polynucleotide 5'-hydroxyl-kinase NOL9 Size: 702 amino acids; 79323 Da
Subunit: Interacts with PELP1, WDR18 and SENP3 (By similarity)
Subcellular location: Nucleus (By similarity). Nucleus, nucleolus. Note=Colocalizes with pre-60S rRNP particles
Sequence caution: Sequence=BAB15611.1; Type=Erroneous initiation; Note=Translation N-terminally extended;
Secondary accessions: Q2NL84 Q4VBY3 Q6P472 Q7L4D6 Q96EE0 Q9H5L4Explore the universe of human proteins at neXtProt for NOL9: NX_Q5SY16
Post-translational modifications:
View modification sites using PhosphoSitePlus2View neXtProt modification sites for NX_Q5SY16
NOL9 Protein expression data from MOPED and PaxDb: About this image 
REFSEQ proteins: NP_078930.3
ENSEMBL proteins: ENSP00000366934
Human Recombinant Protein Products:
Gene Ontology (GO): 1 cellular component term (GO ID links to tree view): About this table | GO ID | Qualified GO term | Evidence | PubMed IDs |
|---|
| GO:0005730 | nucleolus |
TAS | -- |
NOL9 for ontologies About GeneDecksing
NOL9 Antibody Products:
Assay Products for NOL9: |
Protein
Domains /
Families
for NOL9 gene(According to InterPro, ProtoNet,
UniProtKB, and/or BLOCKS,
Sets of similar genes according to GeneDecks)
About This Section
|
NOL9 for domains About GeneDecksing
1 InterPro domain/family:Graphical View of Domain Structure for InterPro Entry Q5SY16ProtoNet protein and cluster: Q5SY16 UniProtKB/Swiss-Prot: NOL9_HUMAN, Q5SY16Similarity: Belongs to the Clp1 family. NOL9/GRC3 subfamily |
Function
for NOL9 gene
(According to 1UniProtKB,
Genatlas,
LifeMap Discovery™,
IUBMB, and/or
2DME,
Human phenotypes from GenomeRNAi,
Animal models from MGI Mar 06 2013,
bound targets from SABiosciences,
miRNA Gene Targets from miRTarBase
shRNA from
OriGene,
RNAi from
EMD Millipore,
siRNAs from
OriGene,
QIAGEN,
microRNA from QIAGEN,
Gene Editing from DNA2.0,
Clones from EMD Millipore,
OriGene,
SwitchGear Genomics,
GenScript,
Sino Biological,
DNA2.0,
and Vector BioLabs,
Cell Lines from GenScript,
LifeMap BioReagents,
In Situ Hybridization Assays from Advanced Cell Diagnostics,
Ontologies according to Gene Ontology Consortium 01 Mar 2013 via
Entrez Gene.)
About This Section
|
Function Summary:
UniProtKB/Swiss-Prot: NOL9_HUMAN, Q5SY16Function: Polynucleotide 5'-kinase involved in rRNA processing. The kinase activity is required for the processing ofthe 32S precursor into 5.8S and 28S rRNAs, more specifically for the generation of the major 5.8S(S) form. In vitro,has both DNA and RNA 5'-kinase activities. Probably binds RNAEnzyme Number (IUBMB): EC 2.7.1.-1
Clone
Products: |  | Browse Clones for the Expression of Recombinant Proteins Available from EMD Millipore | |  | OriGene Myc/DDK tagged cDNA clones in CMV expression vector in human, mouse, rat for NOL9 (see all 3)
OriGene untagged cDNA clones in CMV expression vector in human, mouse, rat for NOL9
OriGene custom cloning services – gene synthesis, subcloning, mutagenesis,
variant library, vector shuttling 
| |  | GenScript: all cDNA clones in your preferred vector: NOL9 (NM_024654) | |  | Browse Sino Biological Human cDNA Clones | |  | DNA2.0 Custom Codon Optimized Gene
Synthesis Service for NOL9 | |  | Vector BioLabs ready-to-use adenovirus/AAV for human, mouse, rat NOL9 |
In Situ Assay
Products: |
 | Advanced Cell Diagnostics RNAscope RNA in situ hybridization assays for NOL9 |
Gene Ontology (GO): 3 molecular function terms (GO ID links to tree view): About this table | GO ID | Qualified GO term | Evidence | PubMed IDs |
|---|
| GO:0003723 | RNA binding |
IEA | -- | | GO:0005524 | ATP binding |
TAS | -- | | GO:0051731 | polynucleotide 5'-hydroxyl-kinase activity |
IDA | -- |
NOL9 for ontologies About GeneDecksing
3 GenomeRNAi human phenotypes for NOL9:
|
Pathways & Interactions
for NOL9 gene
(Pathways according to
EMD Millipore,
R&D Systems,
Cell Signaling Technology,
KEGG,
PharmGKB,
BioSystems,
Reactome,
Tocris Bioscience,
GeneGo (Thomson Reuters),
QIAGEN,
and/or UniProtKB,
Sets of similar genes according to GeneDecks,
Interaction Networks according to
SABiosciences,
and/or STRING,
Interactions according to 1UniProtKB,
2MINT,
3I2D, and/or
4STRING,
with links to IntAct and
Ensembl,
Ontologies according to Gene Ontology Consortium 01 Mar 2013 via
Entrez Gene).
About This Section
|
Interactions:
Search SABiosciences Gene Network CentralTM Interacting Genes and Proteins Networks for NOL9
STRING Interaction
Network Preview (showing 4 interactants - click image to see more details)
4 Interacting proteins for NOL9 (Q5SY163 ENSP000003669344) via UniProtKB, MINT, STRING, and/or I2DAbout this table
Gene Ontology (GO): 1 biological process term (GO ID links to tree view): About this table | GO ID | Qualified GO term | Evidence | PubMed IDs |
|---|
| GO:0000460 | maturation of 5.8S rRNA |
IMP | -- |
NOL9 for ontologies About GeneDecksing
|
Drugs & Compounds
for NOL9 gene(Chemical Compounds according to UniProtKB, Enzo Life Sciences,
EMD Millipore, Tocris Bioscience
HMDB,
BitterDB, and/or
Novoseek, and Drugs according to
DrugBank,
Enzo Life Sciences, and/or
PharmGKB, with drugs/clinical trials/news
search links to CenterWatch)
About This Section
|
 Browse Tocris compounds for NOL9
Search CenterWatch for drugs/clinical trials and news about NOL9
|
Transcripts
for NOL9 gene(Secondary structures according to
fRNAdb,
GenBank/EMBL/DDBJ Accessions according to
Unigene
(Build 235 Homo sapiens; Mar 10 2013) or GenBank,
RefSeq according to Entrez Gene,
DOTS (version 10), and/or
AceView,
transcript ids from Ensembl
with links to UCSC,
exon structure from GeneLoc,
alternative splicing isoforms according to ASD and/or
ECgene,
RNAi Products from
EMD Millipore,
siRNAs from
OriGene,
QIAGEN,
shRNA from
OriGene,
microRNA from QIAGEN,
Tagged/untagged cDNA clones from
OriGene,
SwitchGear Genomics,
GenScript,
DNA2.0,
Vector BioLabs,
Primers from
OriGene,
SABiosciences, and/or
QIAGEN
)
About This Section
|
REFSEQ mRNAs for NOL9 gene: NM_024654.4 Unigene Cluster for NOL9: Nucleolar protein 9 Hs.59425 [show with all ESTs]Unigene Representative Sequence: NM_0246545 Ensembl transcripts including schematic representations, and UCSC links where relevant: ENST00000377705(uc001ans.3 uc010nzs.2) ENST00000464383 ENST00000462798
ENST00000460777 ENST00000464665
Clone
Products: | | OriGene Myc/DDK tagged cDNA clones in CMV expression vector in human, mouse, rat for NOL9 (see all 3)
OriGene untagged cDNA clones in CMV expression vector in human, mouse, rat for NOL9
OriGene custom cloning services – gene synthesis, subcloning, mutagenesis,
variant library, vector shuttling
| | | GenScript: all cDNA clones in your preferred vector: NOL9 (NM_024654) | | | DNA2.0 Custom Codon Optimized Gene
Synthesis Service for NOL9 | | | Vector BioLabs ready-to-use adenovirus/AAV for human, mouse, rat NOL9 |
Additional cDNA sequence: AK026976.1 AK091284.1 AK289653.1 BC009257.2 BC012439.1 BC063639.1 BC073860.1 BC094836.1 BC105095.1 BC110849.1 BC112278.1 BC143646.1 10 DOTS entries: DT.445929 DT.97842025 DT.91940181 DT.100761804 DT.91875388 DT.100725257 DT.91954283 DT.95110332 DT.95258026 DT.439414 24/93 AceView cDNA sequences (see all 93): BM852027 AA504370 BC063639 AI253686 T70351 BX282410 NM_024654 AI075717 CD632184 BC012439 AA343037 CA424414 AA504112 CB109356 AX746895 CA435061 BM856367 AK091284 BF223494 AA219564 BQ428156 CD369212 CB305425 BE396475
GeneLoc Exon Structure
4 Alternative Splicing Database (ASD) splice patterns (SP) for NOL9 About this scheme
| ExUns: | 1 | ^ | 2a | · | 2b | ^ | 3a | · | 3b | ^ | 4 | ^ | 5a | · | 5b | · | 5c | ^ | 6 | ^ | 7a | · | 7b | ^ | 8 | ^ | 9 | ^ | 10 | ^ | 11 | ^ | 12 |
|---|
|
| SP1: | | | | | | | | | - | | | | - | | - | | | | | | | | | | | | | | | | | | | | |
| SP2: | | | | | | | | | | | | | | | | | | | | | | | | | | | | | | | | | | | |
| SP3: | | | | | | | | | | | | | | | | | | | - | | | | | | | | | | | | | | | | |
| SP4: | | | | | | | | | | | | | | | | | | | | | | | | | | | | | | | | | | |
ECgene alternative splicing isoforms for NOL9
|
Expression
for NOL9 gene
(RNA expression data according to
H-InvDB,
NONCODE,
miRBase, and
RNAdb,
Expression images according to data from
BioGPS,
Illumina Human BodyMap, and
CGAP
SAGE,
Sets of similar genes according to GeneDecks,
in vivo and in vitro expression data from LifeMap Discovery™,
plus additional links to
Genevestigator, and/or
SOURCE, and/or
BioGPS, and/or
UniProtKB,
PCR Arrays from
SABiosciences,
Primers from
OriGene,
SABiosciences, and/or
QIAGEN,
In Situ Hybridization Assays from Advanced Cell Diagnostics)
About This Section
| NOL9 expression in normal human tissues (normalized intensities)
See probesets specificity/sensitivity at GeneAnnot
About this imageBioGPS
CGAP TAG: --
About this image
See NOL9 Protein Expression from SPIRE MOPED and PaxDB
Genevestigator expression for NOL9
SOURCE GeneReport for Unigene cluster: Hs.59425
SABiosciences Custom PCR Arrays for NOL9
Primer
Products: | | OriGene genome-wide validated SYBR primer pairs in human, mouse, rat for NOL9
Browse OriGene validated miRNA SYBR primer pairs
| | | SABiosciences RT2 qPCR Primer Assay in human, mouse, rat NOL9 | | | QIAGEN QuantiTect SYBR Green Assays in human, mouse, rat NOL9 | | | QIAGEN QuantiFast Probe-based Assays in human, mouse, rat NOL9 | In Situ
Assay Products: |
| Advanced Cell Diagnostics RNAscope RNA in situ hybridization assays for NOL9 |
Orthologs
for NOL9 gene
(Orthologs according to
1,2HomoloGene (2older version, for species not in 1newer version),
3euGenes,
4SGD
,
5MGI Mar 06 2013,
with possible further links to
Flybase
and/or
WormBase,
and/or
6Ensembl pan taxonomic compara ,
Gene Trees according to Ensembl and
TreeFam)
About This Section
|
This gene was present in the common ancestor of eukaryotes.
Orthologs for NOL9 gene from 9/25 species (see all 25) About this table
ENSEMBL Gene Tree for NOL9 (if available)
TreeFam Gene Tree for NOL9 (if available) |
Paralogs
for NOL9 gene(Paralogs according to
1HomoloGene,
2Ensembl, and 3SIMAP, Pseudogenes according to 4Pseudogene.org Build 68)
About This Section
| Paralogs for NOL9 gene
NOL9 for paralogs About GeneDecksing
1 Pseudogenes.org Pseudogene for NOL9
PGOHUM00000241094
|
Genomic Variants
for NOL9 gene(SNPs/Variants according to the
1NCBI SNP Database,
2Ensembl,
3PupaSUITE,
UniProtKB, and
DNA2.0,
Linkage Disequilibrium by HapMap,
Structural Variations(CNVs/InDels/Inversions) from the Database of Genomic Variants, Mutations from the Human Gene
Mutation Database (HGMD) and the Locus Specific Mutation
Databases (LSDB), Blood group antigen gene mutations by BGMUT,
Resequencing Primers from QIAGEN,
Cancer Mutation PCR Arrays and Assays and Copy Number PCR Arrays from SABiosciences)
About This Section
|
| Genomic Data | Transcription Related Data | Allele Frequencies | | SNP ID | Valid | Clinical
significance | Chr 1 pos | Sequence | # | AA
Chg | Type | More | # | Allele
freq | Pop | Total
sample | More |
|---|
HapMap Linkage Disequilibrium report for NOL9 (6581407 - 6614658 bp)
Structural Variations (Copy Number Variations, Insertions/Deletions, Inversions)
Database of Genomic Variants (DGV): 3 variations for NOL9
3 CNVs: 4202 30364 4201 | SABiosciences Cancer Mutation PCR Assays |
| | QIAGEN SeqTarget long-range PCR primers in human, mouse, rat for resequencing NOL9 |
|
Disorders
/ Diseases
for NOL9 gene
(in which this Gene is Involved, According to MalaCards,
OMIM, UniProtKB,
the University of Copenhagen DISEASES
database,
Genatlas, GeneTests,
GAD,
HuGE Navigator,
and/or TGDB.)
About This Section
|
NOL9 for disorders About GeneDecksing
1  disease for NOL9: About MalaCardschoroiditis
Export disorders for NOL9 gene to outside databases
|
Publications
for NOL9 gene (in
PubMed.
Associations of this gene to articles via
1Entrez Gene,
2UniProtKB/Swiss-Prot,
3HGNC,
4GAD,
5PharmGKB,
6HMDB,
7DrugBank,
8UniProtKB/TrEMBL,
9 Novoseek, and/or
10fRNAdb)
About This Section
|
PubMed articles for NOL9 gene, integrated from 9 sources (see all 19):
(articles sorted by number of sources associating them with NOL9) | |  | Utopia: connect your pdf to the dynamic
world of online information |
- Nol9 is a novel polynucleotide 5'-kinase involved in ribosomal RNA processing. (PubMed id 21063389)1, 2, 3 Heindl K. and Martinez J. (2010)
- The DNA sequence and biological annotation of human chromosome 1. (PubMed id 16710414)1, 2 Gregory S.G.... Bentley D.R. (2006)
- The status, quality, and expansion of the NIH full-length cDNA project: the Mammalian Gene Collection (MGC). (PubMed id 15489334)1, 2 Gerhard D.S....Malek J. (2004)
- Complete sequencing and characterization of 21,243 full-length human cDNAs. (PubMed id 14702039)1, 2 Ota T.... Sugano S. (2004)
- Functional proteomic analysis of human nucleolus. (PubMed id 12429849)1, 2 Scherl A.... Diaz J.-J. (2002)
- Charting the landscape of tandem BRCT domain-mediated protein interactions. (PubMed id 22990118)1 Woods N.T....Monteiro A.N. (2012)
- Functional proteomics establishes the interaction of S IRT7 with chromatin remodeling complexes and expands its role in regulation of R NA polymerase I transcription. (PubMed id 22586326)1 Tsai Y.C....Cristea I.M. (2012)
- A proteome-wide, quantitative survey of in vivo ubiqui tylation sites reveals widespread regulatory roles. (PubMed id 21890473)1 Wagner S.A....Choudhary C. (2011)
- Systematic and quantitative assessment of the ubiquiti n-modified proteome. (PubMed id 21906983)1 Kim W....Gygi S.P. (2011)
- System-wide temporal characterization of the proteome and phosphoproteome of human embryonic stem cell differentiation. (PubMed id 21406692)2 Rigbolt K.T....Blagoev B. (2011)
|
External Searches for NOL9 gene
(in PubMed,
OMIM, and NCBI Bookshelf)
About This Section
|
|
Genome Databases showing NOL9 gene
(According to
Entrez Gene,
HGNC,
AceView,
euGenes,
Ensembl,
miRBase,
ECgene,
Kegg,
and/or
H-InvDB)
About This Section
|
|
Other Databases showing NOL9 gene
(According to HUGE)
About This Section
| -- |
Specialized Databases showing NOL9 gene(According to PharmGKB,
ATLAS, HORDE, IMGT, LEIDEN, UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL,
Wikipedia and/or GeneReviews via UniProtKB/Swiss-Prot)
About This Section
|
| Name | Description |
| PharmGKB entry for NOL9 | Pharmacogenomics, SNPs, Pathways |
|
| | |
About This Section
| Patent Information for NOL9 gene:
Search GeneIP for patents involving NOL9
GeneCards and IP:
Japan Patent Office Licenses GeneCards European Patent Office Licenses GeneCards Improving the IP Search
|
Products
for NOL9 gene(Antibodies, recombinant proteins, and assays from EMD Millipore, R&D Systems, OriGene, QIAGEN, GenScript, Cell Signaling Technology, SABiosciences, Novus Biologicals, Sino Biological, Enzo Life Sciences, Abcam, ProSpec, Uscn, Thermo Fisher Scientific, Gene Editing from DNA2.0, Clones from EMD Millipore, OriGene, GenScript, Sino Biological, DNA2.0, SwitchGear Genomics, Vector BioLabs, Cell lines from GenScript and LifeMap BioReagents, PCR Arrays from SABiosciences, Drugs and/or compounds from EMD Millipore, Tocris Bioscience, and/or
Enzo Life Sciences),
In Situ Hybridization Assays from
Advanced Cell Diagnostics
About This Section
|
 | |
 | |
 |
| | |  | Browse OriGene Antibodies | | OriGene shRNA RFP for NOL9 | | | OriGene 29mer shRNA kits in GFP-retroviral vector in human, mouse, rat for NOL9 | | OriGene genome-wide validated SYBR primer pairs in human, mouse, rat for NOL9 | | | Browse OriGene Protein Over-expression Lysates | | Browse OriGene Fluorogenic Cell Assay Kits | | | OriGene siRNA for NOL9 | | OriGene 3'-UTR Clone for NOL9 | | | OriGene untagged cDNA clones in CMV expression vector in human, mouse, rat for NOL9 | | OriGene Myc/DDK tagged cDNA clones in CMV expression vector in human, mouse, rat for NOL9 | | | Browse OriGene GFP tagged cDNA clones in CMV expression vector | | Browse OriGene MicroRNA Expression Plasmids | | | Browse OriGene basic RS shRNAs | | Browse OriGene validated miRNA SYBR primer pairs | | | Browse OriGene full length recombinant human proteins expressed in human HEK293 cells | | OriGene custom cloning services - gene synthesis, subcloning, mutagenesis,
variant library, vector shuttling | | | OriGene Custom Antibody Services for NOL9 | | OriGene Custom Protein Services for NOL9 | | | OriGene Custom Immunoassay Development | | |
| |
 |
 |
| | | QIAGEN Custom miScript Target Protector blocks miRNA-binding site of in human, mouse, rat NOL9 | | | QIAGEN SeqTarget long-range PCR primers in human, mouse, rat for resequencing NOL9 | | | QIAGEN PyroMark CpG Assay predesigned Pyrosequencing DNA Methylation assays in human, mouse, rat NOL9 | | | QIAGEN FlexiTube/FlexiPlate siRNA for gene silencing in human, mouse, rat NOL9 | | | QIAGEN QuantiFast Probe-based Assays in human, mouse, rat NOL9 | | | QIAGEN QuantiTect SYBR Green Assays in human, mouse, rat NOL9 |
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| | | | Search Tocris compounds for NOL9 |
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 | | | NOL9 Proteins, Antibodies, CLIAs, and ELISAs |
| | | | | Advanced Cell Diagnostics RNAscope RNA in situ hybridization assays for NOL9 |
|  |  |  | | | | ThermoFisher Antibodies for NOL9 |
| | | Vector BioLabs ready-to-use adenovirus/AAV for human, mouse, rat NOL9 |
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