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NACA2 Gene

protein-coding GIFtS: 41

GC17M057025

nascent polypeptide-associated complex alpha subunit 2
(Previous name: nascent-polypeptide-associated complex alpha polypeptide-like )
Symbol approved by the HUGO Gene Nomenclature Committee (HGNC) database
(Previous symbol: NACAL)

(According to ¹HGNC, ²Entrez Gene,
³UniProtKB/Swiss-Prot, ⁴UniProtKB/TrEMBL, ⁵OMIM, ⁶GeneLoc , and/or ⁷Ensembl, ⁸miRBase)
About This Section

Aliases
ANAC ^²
Alpha-NAC-like ^³
MGC71999 ^¹, ²
NAC-alpha-like ^³
NACAL ^², ³

Descriptions
Hom s 2.01 ^³
IgE autoantigen alpha-nascent polypeptide-associated complex ^²
Nascent polypeptide-associated complex subunit alpha-like ^³
alpha-NAC protein ^²
nascent polypeptide-associated complex alpha subunit 2 ^²
nascent polypeptide-associated complex, alpha polypeptide, 2 ^²
nascent-polypeptide-associated complex alpha polypeptide-like ^¹, ²

External Ids
HGNC: 23290¹
Entrez Gene: 342538²
UniProtKB: Q9H009³
Ensembl: ENSG00000196861⁷

Search outside databases for aliases for NACA2 gene

(According to Entrez Gene, Wikipedia's Gene Wiki,
UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL)
About This Section

UniProtKB/Swiss-Prot: NACA2_HUMAN, Q9H009

Function: Prevents inappropriate targeting of non-secretory polypeptides to the endoplasmic

reticulum (ER). Binds to nascent polypeptide chains as they emerge from the ribosome and blocks

their interaction with the signal recognition particle (SRP), which normally targets nascent

secretory peptides to the ER. Also reduces the inherent affinity of ribosomes for protein

translocation sites in the ER membrane (M sites) (By similarity)

(According to GeneLoc and/or HGNC, and/or
Entrez Gene (NCBI build 36),
and/or miRBase,
Genomic Views according to UCSC and Ensembl, Transcription factor binding sites according to SABiosciences)
About This Section

Genomic View:

UCSC Golden Path with GeneCards custom track

Entrez Gene cytogenetic band: 17q23.2 Ensembl cytogenetic band: 17q23.2 HGNC cytogenetic band: 17q23.3

NACA2 Gene in genomic location: bands according to Ensembl, locations according to (and/or Entrez Gene and/or Ensembl if different)

GeneLoc gene densities for chromosome 17 GeneLoc Exon Structure

GeneLoc location for GC17M057025: (about GC identifiers)

Start:	57,022,569 bp from pter
End:	57,023,345 bp from pter
Size:	777 bases
Orientation:	minus strand

RefSeq DNA sequence:

NC_000017.9 NT_010783.14

(According to ¹UniProtKB, and/or Ensembl, Phosphorylation sites according to ²Phosphosite, RefSeq according to NCBI, PDB rendering according to OCA and/or Proteopedia, Recombinant Proteins from Invitrogen, Millipore, Sigma-Aldrich, R&D Systems, Enzo Life Sciences, Abnova, OriGene and/or, Abcam,
Biochemical Assays by Invitrogen, Millipore, R&D Systems, Cell Signaling Technology, and/or Enzo Life Sciences, Ontologies according to Gene Ontology Consortium 01 Apr 2009 and Entrez Gene, Antibodies by Invitrogen, Millipore, Sigma-Aldrich, R&D Systems, Cell Signaling Technology, Abcam, Abnova, and/or Novus Biologicals)
About This Section

UniProtKB/Swiss-Prot: NACA2_HUMAN, Q9H009 (See protein sequence)

Recommended Name: Nascent polypeptide-associated complex subunit alpha-2

Size: 215 amino acids; 23223 Da

Subunit: Part of the nascent polypeptide-associated complex (NAC), consisting of NACA and BTF3. NAC

associates with ribosomes through the BTF3 subunit. Both subunits can contact nascent polypeptide

chains (By similarity)

Subcellular location: Cytoplasm (By similarity). Nucleus (By similarity)

Miscellaneous: NACAL arose from a recent (40-63 million year old), anthropoid primate-specific

retroduplication of NACA

Secondary accessions: Q2VIR9

REFSEQ proteins: NP_954984.1

ENSEMBL proteins:

ENSP00000352719

Human Recombinant Proteins

	Browse Drug Discovery Central at Invitrogen for human recombinant proteins
	Browse Purified and Recombinant Proteins at Millipore
	Browse Human Recombinant Proteins at Sigma-Aldrich
	Browse R&D Systems for human recombinant proteins
	Browse recombinant and purified proteins available from Enzo Life Sciences
	Recombinant Proteins from Abcam (NACA2)
	Human Recombinant Proteins from Abnova (NACAL)

Browse Origene for full length recombinant human proteins expressed in human HEK293 cells

2 Gene Ontology (GO) cellular component terms (links to tree view):

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0005634	nucleus	IEA	--
GO:0005737	cytoplasm	IEA	--

About this table

Antibodies for NACA2:

	Invitrogen Antibodies for NACA2
	Browse Millipore's Extensive Line of Mono- and Polyclonal Antibodies
	Browse Antibodies at Sigma-Aldrich
	Browse R&D Systems for Antibodies
	Antibodies from Abcam (NACA2), each with their Abpromise^SM
	Monoclonal and Polyclonal Antibodies from Abnova (NACAL)
	Novus Biologicals Antibodies for NACA2

Assays for NACA2:

	Browse Invitrogen for biochemical assays
	Browse Kits and Assays available from Millipore
	Browse R&D Systems for biochemical assays
	Browse biochemical assays available from Enzo Life Sciences

(According to InterPro, ProtoNet, UniProtKB, and/or BLOCKS, Sets of similar genes according to GeneDecks)
About This Section

Graphical View of Domain Structure for InterPro Entry Q9H009

ProtoNet protein and cluster: Q9H009

1 Blocks protein family: IPB000449 Ubiquitin-associated domain

UniProtKB/Swiss-Prot: NACA2_HUMAN, Q9H009

Similarity: Belongs to the NAC-alpha family

Similarity: Contains 1 NAC-A/B (NAC-alpha/beta) domain

Similarity: Contains 1 UBA domain

(According to MGI Jun 06 2009, UniProtKB, IUBMB,and/or Genatlas,
shRNA from OriGene, Sigma-Aldrich, RNAi from Sigma-Aldrich,
RNAi Products, Clones, and Q-PCR Products from Invitrogen, Millipore, OriGene, and/or Abnova, siRNAs from Applied Biosystems, SYBR primers from OriGene, Cell-based Assays from Millipore, Ontologies according to Gene Ontology Consortium 01 Apr 2009 via Entrez Gene.)
About This Section

Inhib. RNA:		Invitrogen RNAi Products for gene knock-down (NACA2)
		Browse for Gene Knock-down Tools from Millipore
		Abnova Chimera RNAi Products for Gene knock-down (NACAL)

OriGene 29mer shRNA kit in GFP-retroviral vectors (see all 2): XM_292598

Applied Biosystems Silencer® siRNAs for NACA2

Sigma-Aldrich siRNA and siRNA Panels for NACA2

Sigma-Aldrich shRNA for NACA2

Explore Sigma-Aldrich super-pooled esiRNAs

Clones:		Invitrogen Clones for NACA2
		Browse Clones for the Expression of Recombinant Proteins Available from Millipore

OriGene GFP tagged cDNA clone in CMV expression vector: NM_199290
Myc/DDK tagged cDNA clone in CMV expression vector: NM_199290
untagged cDNA clone in CMV expression vector: NM_199290

Primers:

Browse Quantitative PCR Central at Invitrogen for Q-PCR LUX™ Primers

OriGene genome-wide validated SYBR primer pairs: NM_199290

UniProtKB/Swiss-Prot: NACA2_HUMAN, Q9H009

Function: Prevents inappropriate targeting of non-secretory polypeptides to the endoplasmic

reticulum (ER). Binds to nascent polypeptide chains as they emerge from the ribosome and blocks

their interaction with the signal recognition particle (SRP), which normally targets nascent

secretory peptides to the ER. Also reduces the inherent affinity of ribosomes for protein

translocation sites in the ER membrane (M sites) (By similarity)

(Pathways according to Invitrogen (maps by GeneGo), Millipore, Cell Signaling Technology, Sigma-Aldrich, KEGG and/or UniProtKB,
Sets of similar genes according to GeneDecks, Proteins Network according to SABiosciences, Interactions according to ¹UniProtKB, ²MINT, and/or ³STRING, with links to IntAct and Ensembl, Ontologies according to Gene Ontology Consortium 01 Apr 2009 via Entrez Gene.)
About This Section

1 Gene Ontology (GO) biological process term (links to tree view):

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0015031	protein transport	IEA	--

About this table

(Chemical Compounds according to UniProtKB, Enzo Life Sciences, Sigma-Aldrich, Tocris Bioscience, and/or Novoseek and Drugs according to Enzo Life Sciences and/or PharmGKB)
About This Section

Browse drugs & compounds from Enzo Life Sciences

Browse Small Molecules at Sigma-Aldrich

Browse Tocris compounds for NACA2

(GenBank/EMBL/DDBJ Accessions according to
Unigene (Build 219 Homo sapiens; Jun 2 2009) or GenBank,
RefSeq according to Entrez Gene,
DOTS (version 10), and/or AceView,
non coding RNAs according to RNAdb,
ESTs according to GeneTide,
exon structure from GeneLoc, alternative splicing isoforms according to ASD and/or ECgene,
RNAi Products from Invitrogen, Millipore, and/or Abnova,
siRNAs from Applied Biosystems, Sigma-Aldrich,
shRNA from Sigma-Aldrich, OriGene,
Tagged/untagged cDNA clones from OriGene,
Expression Assays from Applied Biosystems)
About This Section

Inhib. RNA:		Invitrogen RNAi Products for gene knock-down (NACA2)
		Browse for Gene Knock-down Tools from Millipore
		Abnova Chimera RNAi Products for Gene knock-down (NACAL)

OriGene 29mer shRNA kit in GFP-retroviral vectors (see all 2): XM_292598

Sigma-Aldrich siRNA and siRNA Panels for NACA2

Sigma-Aldrich shRNA for NACA2

Explore Sigma-Aldrich super-pooled esiRNAs

Applied Biosystems Silencer® siRNAs:

NM_199290

REFSEQ mRNAs for NACA2 gene:

NM_199290.2

Applied Biosystems TaqMan ® Gene Expression Assays:

NM_199290

              OriGene GFP tagged cDNA clone in CMV expression vector: NM_199290
                                 Myc/DDK tagged cDNA clone in CMV expression vector: NM_199290
                                 untagged cDNA clone in CMV expression vector: NM_199290

Additional cDNA sequence: BC062710.1

15 DOTS entries:

DT.97868164 DT.97868229 DT.121209295 DT.91733258 DT.91783886 DT.452984 DT.100889874 DT.121209326

DT.95228035 DT.121209132 DT.121470534 DT.85101137 DT.86993099 DT.91718355 DT.91783876

16 AceView cDNA sequences:

NM_199290 BC062710 BU567791 T61059 AA450295 BU740309 BU949208 AA485916

CA942667 BM662462 AI422625 AI434582 N89716 BQ346430 AW868519 AW863307

Unigene Cluster for NACA2:

Nascent polypeptide-associated complex alpha subunit 2

Hs.591178 [show with all ESTs]

Unigene Representative Sequence: BU567791

GeneLoc Exon Structure

1 Ensembl transcript including schematic representation:

ENST00000359690

(Experimental results according to ¹GeneNote and GNF BioGPS,
probe sets-to-genes annotations according to ²GeneAnnot , ³GeneTide , Sets of similar genes according to GeneDecks, Electronic Northern calculations according to data from UniGene (Build 219 Homo sapiens), SAGE tags according to CGAP, plus additional links to SOURCE, and/or GNF BioGPS, and/or EXPOLDB, and/or UniProtKB,
Expression Assays from Applied Biosystems )
About This Section

NACA2 expression in normal and diseased human tissues

Applied Biosystems TaqMan ® Gene Expression Assays for NACA2

¹ / ²

3 probe-sets matching NACA2 gene

Affymetrix probe-set	Array	# genes	Sensitivity	Specificity	Correlation	Length	Gb_Accession	Consensus	Uniqueness	Score	Rank
Affymetrix probe-set	Array	GeneAnnot data			GeneNote data		GeneTide data
39739_at²	U95-A	4	0.94	0.29	1.00	1.00	--	--	--	--	--
222224_at²	U133-A	1	1.00	1.00	--	--	--	--	--	--	--
222224_at²	U133Plus2	1	1.00	1.00	--	--	--	--	--	--	--

About this table

Data from

(Publications) and GNF BioGPS
About these images
About these images

CGAP SAGE TAG: --

SOURCE GeneReport for Unigene cluster: Hs.591178
UniProtKB/Swiss-Prot: NACA2_HUMAN, Q9H009
Tissue specificity: Expressed specifically in testis and skeletal muscle

(Orthologs according to ^1,2HomoloGene (²older version, for species not in ¹newer version), ³euGenes, ⁴SGD and/or ⁵MGI Jun 06 2009, with possible further links to Flybase and/or WormBase, Gene Trees according to Ensembl)
About This Section

Orthologs for NACA2 gene from 2 species

Organism	Gene	Locus	Description	Human Similarity	NCBI accessions
chimpanzee (Pan troglodytes)	NACA2¹	--	nascent polypeptide-associated complex alpha subunit more	98.91(n) 97.21(a)	454797 XM_511608.2 XP_511608.1
thale cress (Arabidopsis thaliana)	AT4G10480¹	--	nascent polypeptide associated complex alpha chain more	60.59(n) 58.78(a)	826640 NM_001084900.1 NP_001078369.1

About this table Species with no ortholog for NACA2

ENSEMBL Gene Tree for NACA2

(Paralogs according to ¹HomoloGene
and ²Ensembl, Pseudogenes according to ³Pseudogene.org)
About This Section

Paralogs for NACA2 gene

NACA3P² NACAD² NACA²

Pseudogenes

2 related pseudogenes

(According to the ¹NCBI SNP Database, ²Ensembl, ³PupaSUITE, and UniProtKB, Linkage Disequilibrium by HapMap, Genotyping Reagents from Applied Biosystems)
About This Section

10 NCBI SNPs in NACA2 are shown
(Click for Applied Biosystems TaqMan ® Genotyping Assay) (only AB snps)

AB	SNP ID	Valid	Chr 17 pos	Sequence	Recs	AA Chg	Type	More	Recs	Allele freq	Pop	Total sample	More
AB	Genomic Data				Transcription Data				Allele Frequencies
				--	--	--		--	--				--
	rs2188729^1,2	A,C,F,H,O	57023916(-)	`TTAAAC/TTGAAT`	1	--	ng3¹		11		NA EU EA WA	980
	rs7219527^1,2	A,C,F,H	57024671(+)	`TTTGGC/TAGCAC`	1	--	ng3¹		8		EU EA WA	820
--	rs17531723^1,2	C,F,H	57023134(+)	`ACCGAC/TTGGTT`	1	V/I	mis¹ ese³		11		NA EU EA WA	968
--	rs17610181^1,2	C,F,H	57022735(+)	`AGCTCG/AGACTG`	1	X/R	non¹		11		NA EU EA WA	978
	rs4141531^1,2	C,F,H	57023859(-)	`AGATCG/AAGGGA`	1	--	ng3¹		4		EU EA WA	420
--	rs61748811^1,2	--	57023308(+)	`TTCTGA/GGGCTT`	2	T/P T/S	mis¹		0	--	--	--	--
--	rs61748812^1,2	--	57023299(+)	`AGGGAG/TGGTTT`	2	V/I V/L	mis¹		0	--	--	--	--
--	rs61739272^1,2	--	57022834(+)	`CTCCTA/GTTGTA`	2	E/Q E/X	mis¹ non¹		0	--	--	--	--
--	rs61739273^1,2	--	57022803(+)	`CACCTC/GTTTCA`	1	T/R	mis¹		0	--	--	--	--
--	rs61739271^1,2	--	57023057(+)	`ACTCCA/TGTAAC`	1	T/T	syn¹		0	--	--	--	--

About this table
The following additional 37 SNP(s) have been found in Applied Biosystems data source:

(Click for Applied Biosystems TaqMan ® Genotyping Assay)

rs9899863	rs10445326	rs9893569	rs725900	rs12051660	rs728368	rs728367
rs12602207	rs7225708	rs9901535	rs9910461	rs16945466	rs12951015	rs8076873
rs7207425	rs725899	rs4968576	rs9910311	rs11079440	rs9303446	rs9909556
rs739579	rs7226057	rs12453545	rs10445324	rs9303447	rs4968575	rs4968442
rs10445327	rs9914047	rs2188730	rs12947026	rs16945455	rs16945472	rs2188728
rs7214809	rs10515203

HapMap Linkage Disequilibrium images for NACA2 (up to first 250kb)

(in which this Gene is Involved, According to OMIM, UniProtKB, PharmGKB, Genatlas, GeneTests, Blood group antigen gene mutations by BGMUT, HGMD, GAD, HuGE Navigator, BCGD, and/or TGDB.)
About This Section

OMIM: 609274

(Possibly Related Articles in Doctor's Guide)
About This Section

(in PubMed. Associations of this gene to articles via ¹Novoseek, ²HGNC, ³Entrez Gene, ⁴UniProtKB/Swiss-Prot, ⁵UniProtKB/TrEMBL, ⁶GAD, and/or ⁷PharmGKB)
About This Section

7 PubMed articles for NACA2 gene:

Characterization of a novel isoform of alpha-nascent polypeptide- associated complex as IgE-defined autoantigen. (PubMed id 12406326)^{2, 3, 4} Mossabeb R.... Valenta R. (2002)
Emergence of young human genes after a burst of retroposition in primates. (PubMed id 16201836)^{3, 4} Marques A.C.... Kaessmann H. (2005)
The status, quality, and expansion of the NIH full-length cDNA project: the Mammalian Gene Collection (MGC). (PubMed id 15489334)^{3, 4} Gerhard D.S....Malek J. (2004)
Many sequence variants affecting diversity of adult human height. (PubMed id 18391951)³ Gudbjartsson D.F....Stefansson K. (2008)
Generation and initial analysis of more than 15,000 full-length human and mouse cDNA sequences. (PubMed id 12477932)³ Strausberg R.L....Marra M.A. (2002)
The alpha and beta subunit of the nascent polypeptide-associated complex have distinct functions. (PubMed id 10982809)³ Beatrix B.... Wiedmann M. (2000)
NAC covers ribosome-associated nascent chains thereby forming a protective environment for regions of nascent chains just emerging from the peptidyl transferase center. (PubMed id 7622554)³ Wang S....Wiedmann M. (1995)

(in PubMed, OMIM, and NCBI Bookshelf)
About This Section

		AND	OR
Aliases
Free Text

Query String		PubMed OMIM NCBI Bookshelf
	(Note: In FireFox, select the above section and copy using Ctrl-C)

(According to Entrez Gene, HGNC, AceView, euGenes, Ensembl, miRBase, ECgene, and/or H-InvDB)
About This Section

Entrez Gene: 342538	HGNC: 23290	AceView: MGC71999	Ensembl:ENSG00000196861	euGenes: HUgn342538
ECgene: NACA2	H-InvDB: NACA2

(According to HUGE)
About This Section

(According to ATLAS, HORDE, IMGT, MTDB, LEIDEN, UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL,
Wikipedia and/or GeneReviews via UniProtKB/Swiss-Prot)
About This Section

Licensable Technologies for NACA2
(Available from WIS Yeda, Salk, Tufts)

About This Section

(Reagents available from Applied Biosystems, Antibodies and assays by Cell Signaling Technology, Abcam, Novus Biologicals,
Sigma-Aldrich, R&D Systems, Millipore, Abnova, and/or Invitrogen, Clones available from OriGene,and/or Invitrogen, Drugs and/or compounds by Sigma-Aldrich,
Enzo Life Sciences, and/or Tocris Bioscience)
About This Section

Products for NACA2:

TaqMan ® Gene Expression Assays

TaqMan ® Genotyping Assays

Free SNP selection tool

Invitrogen iPath Pathways	Invitrogen BLOCK-iT™ RNAi
Invitrogen Antibodies	Invitrogen Assays
Invitrogen Clones	Invitrogen Q-PCR Products
Invitrogen Human Recombinant Kinases	Invitrogen Custom Antibody and Peptide Service
Invitrogen Proteins / Assays / Screening Services	Search Invitrogen catalog for NACA2-related products

	Millipore Custom Antibody & Bulk Services
	Millipore Preclinical / Clinical Development Services
	Millipore Immunoassay Services
	Millipore Target Screening & Profiling Services

	Predesigned and custom siRNAs for NACA2		Browse antibodies at Sigma-Aldrich
	Explore super-pooled esiRNAs		Browse proteins at Sigma-Aldrich
	Lentivirus-delivered shRNAs for NACA2		Browse small molecules at Sigma-Aldrich
	"Your Favorite Gene" Pathways		feedback

	Antibodies		Primer Pairs
	Cell Culture Products		ELISAs
	Flow Cytometry Kits		Protease Activity Assays & Reagents
	Cell Selection/Detection Kits & Reagents		ELISA/Assay Development Kits & Reagents
	Multiplex/Array Assay Kits & Reagents		ELISpot Kits & Development Modules
	Recombinant & Natural Proteins

	Recombinant Proteins (NACAL)
	Antibodies (NACAL)
	Chimera RNAi (NACAL)
	Custom Service for Mouse Mab
	Custom Service for Rabbit Pab from Full-length Protein

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	Antibodies (NACA2)

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Search Tocris compounds for NACA2

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Antibodies for NACA2

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NACA2 Gene

protein-coding GIFtS: 41

Aliases &
Descriptions
for NACA2

Summaries
for NACA2

Genomic Location
for NACA2

Proteins
for NACA2

Protein Domains/
Families
for NACA2

Gene Function
for NACA2

Pathways & Interactions
for NACA2

Drugs & Compounds
for NACA2

Transcripts
for NACA2

Expression
for NACA2

Orthologs
for NACA2

Paralogs
for NACA2

SNPs/Variants
for NACA2

Disorders & Mutations
for NACA2

Medical News
for NACA2

Publications
for NACA2

Search for NACA2

Genome Databases showing NACA2

Other Databases showing NACA2

Specialized Databases showing NACA2

Licensable Technologies
for NACA2

Services
for NACA2

Gene Index: 3 5 A B C D E F G H I J K L M N O P Q R S T U V W X Y Z

NACA2 Gene

protein-coding GIFtS: 41

Aliases & Descriptionsfor NACA2

Summariesfor NACA2

Genomic Locationfor NACA2

Proteinsfor NACA2

Protein Domains/Familiesfor NACA2

Gene Functionfor NACA2

Pathways & Interactionsfor NACA2

Drugs & Compoundsfor NACA2

Transcriptsfor NACA2

Expression for NACA2

Orthologsfor NACA2

Paralogsfor NACA2

SNPs/Variantsfor NACA2

Disorders & Mutationsfor NACA2

Medical Newsfor NACA2

Publicationsfor NACA2

Search for NACA2

Genome Databases showing NACA2

Other Databases showing NACA2

Specialized Databases showing NACA2

Licensable Technologiesfor NACA2

Servicesfor NACA2

Gene Index: 3 5 A B C D E F G H I J K L M N O P Q R S T U V W X Y Z

Aliases &
Descriptions
for NACA2

Summaries
for NACA2

Genomic Location
for NACA2

Proteins
for NACA2

Protein Domains/
Families
for NACA2

Gene Function
for NACA2

Pathways & Interactions
for NACA2

Drugs & Compounds
for NACA2

Transcripts
for NACA2

Expression
for NACA2

Orthologs
for NACA2

Paralogs
for NACA2

SNPs/Variants
for NACA2

Disorders & Mutations
for NACA2

Medical News
for NACA2

Publications
for NACA2

Licensable Technologies
for NACA2

Services
for NACA2