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N4BP1 Gene

protein-coding GIFtS: 49

GC16M047131

NEDD4 binding protein 1
Symbol approved by the HUGO Gene Nomenclature Committee (HGNC) database

(According to ¹HGNC, ²Entrez Gene,
³UniProtKB/Swiss-Prot, ⁴UniProtKB/TrEMBL, ⁵OMIM, ⁶GeneLoc , and/or ⁷Ensembl, ⁸miRBase)
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Aliases
FLJ31821 ^²
KIAA0615 ^², ³
MGC176730 ^²

Descriptions
NEDD4 binding protein 1 ^²

External Ids
HGNC: 29850¹
Entrez Gene: 9683²
UniProtKB: O75113³
Ensembl: ENSG00000102921⁷

Search outside databases for aliases for N4BP1 gene

Previous GC identifers: GC16M038991 GC16M048624 GC16M048311 GC16M048354

(According to Entrez Gene, Wikipedia's Gene Wiki,
UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL)
About This Section

(According to GeneLoc and/or HGNC, and/or
Entrez Gene (NCBI build 36),
and/or miRBase,
Genomic Views according to UCSC and Ensembl, Transcription factor binding sites according to SABiosciences)
About This Section

Genomic View:

UCSC Golden Path with GeneCards custom track

Transcription factor binding sites upstream to the N4BP1 gene

Entrez Gene cytogenetic band: 16q12.1 Ensembl cytogenetic band: 16q12.1 HGNC cytogenetic band: 16q12.1

N4BP1 Gene in genomic location: bands according to Ensembl, locations according to (and/or Entrez Gene and/or Ensembl if different)

GeneLoc gene densities for chromosome 16 GeneLoc Exon Structure

GeneLoc location for GC16M047131: (about GC identifiers)

Start:	47,130,138 bp from pter
End:	47,201,621 bp from pter
Size:	71,484 bases
Orientation:	minus strand

RefSeq DNA sequence:

NC_000016.8 NT_010498.15

(According to ¹UniProtKB, and/or Ensembl, Phosphorylation sites according to ²Phosphosite, RefSeq according to NCBI, PDB rendering according to OCA and/or Proteopedia, Recombinant Proteins from Invitrogen, Millipore, Sigma-Aldrich, R&D Systems, Enzo Life Sciences, Abnova, OriGene and/or, Abcam,
Biochemical Assays by Invitrogen, Millipore, R&D Systems, Cell Signaling Technology, and/or Enzo Life Sciences, Ontologies according to Gene Ontology Consortium 01 Apr 2009 and Entrez Gene, Antibodies by Invitrogen, Millipore, Sigma-Aldrich, R&D Systems, Cell Signaling Technology, Abcam, Abnova, and/or Novus Biologicals)
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UniProtKB/Swiss-Prot: N4BP1_HUMAN, O75113 (See protein sequence)

Recommended Name: NEDD4-binding protein 1

Size: 896 amino acids; 100409 Da

Subunit: Binds NEDD4 (By similarity)

Subcellular location: Nucleus. Note=In discrete circular structures (By similarity)

Secondary accessions: A7MD49 Q2YDX1

Post-translational modifications:

Monoubiquitinated (By similarity)¹

Phosphorylated upon DNA damage, probably by ATM or ATR¹

View phosphorylation sites using PhosphoSite²

REFSEQ proteins: NP_694574.3

ENSEMBL proteins:

ENSP00000262384

Human Recombinant Proteins

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	Recombinant Proteins from Abcam (N4BP1)
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Browse Origene for full length recombinant human proteins expressed in human HEK293 cells

1 Gene Ontology (GO) cellular component term (links to tree view):

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0005634	nucleus	IEA	--

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Antibodies for N4BP1:

	Browse Antibodies Central at Invitrogen
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	Browse R&D Systems for Antibodies
	Antibodies from Abcam (N4BP1), each with their Abpromise^SM
	Browse Abnova for Monoclonal and Polyclonal Antibodies
	Search Novus for antibodies for N4BP1

Assays for N4BP1:

	Browse Invitrogen for biochemical assays
	Browse Kits and Assays available from Millipore
	Browse R&D Systems for biochemical assays
	Browse biochemical assays available from Enzo Life Sciences

(According to InterPro, ProtoNet, UniProtKB, and/or BLOCKS, Sets of similar genes according to GeneDecks)
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ProtoNet protein and cluster: O75113

UniProtKB/Swiss-Prot: N4BP1_HUMAN, O75113

Similarity: Belongs to the N4BP1 family

(According to MGI Jun 06 2009, UniProtKB, IUBMB,and/or Genatlas,
shRNA from OriGene, Sigma-Aldrich, RNAi from Sigma-Aldrich,
RNAi Products, Clones, and Q-PCR Products from Invitrogen, Millipore, OriGene, and/or Abnova, siRNAs from Applied Biosystems, SYBR primers from OriGene, Cell-based Assays from Millipore, Ontologies according to Gene Ontology Consortium 01 Apr 2009 via Entrez Gene.)
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Inhib. RNA:		Invitrogen RNAi Products for gene knock-down (N4BP1)
		Browse for Gene Knock-down Tools from Millipore
		Browse Abnova for Chimera RNAi Products

OriGene 29mer shRNA kit in GFP-retroviral vectors (see all 2): NM_014664

Applied Biosystems Silencer® siRNAs for N4BP1

Sigma-Aldrich siRNA for N4BP1

Sigma-Aldrich shRNA for N4BP1

Explore Sigma-Aldrich super-pooled esiRNAs

Clones:		Browse Clone Ranger at Invitrogen for clones
		Browse Clones for the Expression of Recombinant Proteins Available from Millipore

OriGene GFP tagged cDNA clone in CMV expression vector: NM_153029
Myc/DDK tagged cDNA clone in CMV expression vector: NM_153029
untagged cDNA clones in CMV expression vector (see all 3): NM_153029

Primers:

Browse Quantitative PCR Central at Invitrogen for Q-PCR LUX™ Primers

OriGene genome-wide validated SYBR primer pairs: NM_153029

1 MGI mutant phenotype (inferred from 1 allele) (MGI details for N4bp1):

no phenotypic analysis

(Pathways according to Invitrogen (maps by GeneGo), Millipore, Cell Signaling Technology, Sigma-Aldrich, KEGG and/or UniProtKB,
Sets of similar genes according to GeneDecks, Proteins Network according to SABiosciences, Interactions according to ¹UniProtKB, ²MINT, and/or ³STRING, with links to IntAct and Ensembl, Ontologies according to Gene Ontology Consortium 01 Apr 2009 via Entrez Gene.)
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1 Interacting protein for N4BP1 (O75113²) via UniProtKB, MINT, and/or STRING
Interactant Interaction Details
GeneCard External ID(s)
-- P46935² MINT-14958 MINT-14957
About this table

(Chemical Compounds according to UniProtKB, Enzo Life Sciences, Sigma-Aldrich, Tocris Bioscience, and/or Novoseek and Drugs according to Enzo Life Sciences and/or PharmGKB)
About This Section

Browse drugs & compounds from Enzo Life Sciences

Browse Small Molecules at Sigma-Aldrich

Browse Tocris compounds for N4BP1

(GenBank/EMBL/DDBJ Accessions according to
Unigene (Build 219 Homo sapiens; Jun 2 2009) or GenBank,
RefSeq according to Entrez Gene,
DOTS (version 10), and/or AceView,
non coding RNAs according to RNAdb,
ESTs according to GeneTide,
exon structure from GeneLoc, alternative splicing isoforms according to ASD and/or ECgene,
RNAi Products from Invitrogen, Millipore, and/or Abnova,
siRNAs from Applied Biosystems, Sigma-Aldrich,
shRNA from Sigma-Aldrich, OriGene,
Tagged/untagged cDNA clones from OriGene,
Expression Assays from Applied Biosystems)
About This Section

Inhib. RNA:		Invitrogen RNAi Products for gene knock-down (N4BP1)
		Browse for Gene Knock-down Tools from Millipore
		Browse Abnova for Chimera RNAi Products

OriGene 29mer shRNA kit in GFP-retroviral vectors (see all 2): NM_014664

Sigma-Aldrich siRNA for N4BP1

Sigma-Aldrich shRNA for N4BP1

Explore Sigma-Aldrich super-pooled esiRNAs

Applied Biosystems Silencer® siRNAs:

NM_153029

REFSEQ mRNAs for N4BP1 gene:

NM_153029.3

Applied Biosystems TaqMan ® Gene Expression Assays:

NM_153029

              OriGene GFP tagged cDNA clone in CMV expression vector: NM_153029
                                 Myc/DDK tagged cDNA clone in CMV expression vector: NM_153029
                                 untagged cDNA clones in CMV expression vector (see all 3): NM_153029

Additional cDNA sequence:

AB014515.1 AK001464.1 AK026937.1 AK056383.1 AK094234.1 AK124782.1 AL831882.1 BC051897.1

BC108288.1 BC152472.1

8 DOTS entries:

DT.213542 DT.75103885 DT.112202 DT.209678 DT.86854338 DT.95369347 DT.40119828 DT.91713557

24/179 AceView cDNA sequences (see all 179 ):

BQ000756 AK001464 BU160325 AB014515 BU521373 AL523704 AI858244 AA232451

AI859000 BF509350 F13180 BM707079 AL046130 AK056383 BX477474 NM_153029

AA911171 AI379495 BQ645844 F13173 AA919039 BQ962044 AI199182 CA393960

highest scoring ESTs for N4BP1:

AB014515 AA225490 BC051897 BU160325 CA393960 AA225384 AA225442 AA312219 AA316977 AA320069

Unigene Cluster for N4BP1:

NEDD4 binding protein 1

Hs.511839 [show with all ESTs]

Unigene Representative Sequence: NM_153029

GeneLoc Exon Structure

3 Alternative Splicing Database (ASD) splice patterns (SP) for N4BP1

ExUns:	1	^	2	^	3a	·	3b	^	4	^	5a	·	5b	^	6	^	7	^	8a	·	8b
SP1:			-
SP2:																	-
SP3:

About this scheme

ECgene alternative splicing isoforms for N4BP1

1 Ensembl transcript including schematic representation:

ENST00000262384

(Experimental results according to ¹GeneNote and GNF BioGPS,
probe sets-to-genes annotations according to ²GeneAnnot , ³GeneTide , Sets of similar genes according to GeneDecks, Electronic Northern calculations according to data from UniGene (Build 219 Homo sapiens), SAGE tags according to CGAP, plus additional links to SOURCE, and/or GNF BioGPS, and/or EXPOLDB, and/or UniProtKB,
Expression Assays from Applied Biosystems )
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N4BP1 expression in normal and diseased human tissues

Applied Biosystems TaqMan ® Gene Expression Assays for N4BP1

¹ / ² / ³

10 probe-sets matching N4BP1 gene

Affymetrix probe-set	Array	# genes	Sensitivity	Specificity	Correlation	Length	Gb_Accession	Consensus	Uniqueness	Score	Rank
Affymetrix probe-set	Array	GeneAnnot data			GeneNote data		GeneTide data
32069_at²^{, 3}	U133-A	1	1.00	1.00	0.96	0.72	AB014515	1.00	1.00	1.00	1
48612_at²^{, 3}	U133-C	1	1.00	1.00	0.37	1.28	AA225490	0.80	1.00	0.91	1
204601_at²^{, 3}	U133-A	1	1.00	1.00	--	--	NM_014664	0.20	1.00	0.72	1
32069_at²^{, 3}	U133-A	1	1.00	1.00	--	--	AB014515	1.00	1.00	1.00	1
48612_at²^{, 3}	U133-A	1	1.00	1.00	--	--	AA225490	0.80	1.00	0.91	1
221867_at²^{, 3}	U133-A	1	1.00	1.00	--	--	BF436315	0.60	1.00	0.82	1
204601_at²	U133Plus2	1	1.00	1.00	--	--	--	--	--	--	--
32069_at²	U133Plus2	1	1.00	1.00	--	--	--	--	--	--	--
48612_at²	U133Plus2	1	1.00	1.00	--	--	--	--	--	--	--
221867_at²	U133Plus2	1	1.00	1.00	--	--	--	--	--	--	--

GeneDecks N4BP1 for binary patterns associated with the probe-sets selected above

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Data from

(Publications) and GNF BioGPS
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CGAP SAGE TAG: ATGTTGTGAC

SOURCE GeneReport for Unigene cluster: Hs.511839
Expression variation in blood from EXPOLDB for N4BP1
UniProtKB/Swiss-Prot: N4BP1_HUMAN, O75113
Tissue specificity: Detected in heart, lung, brain, liver, skeletal muscle, pancreas, kidney,
spleen, testis and ovary

(Orthologs according to ^1,2HomoloGene (²older version, for species not in ¹newer version), ³euGenes, ⁴SGD and/or ⁵MGI Jun 06 2009, with possible further links to Flybase and/or WormBase, Gene Trees according to Ensembl)
About This Section

Orthologs for N4BP1 gene from 5/9 species (see all 9 )

Organism	Gene	Locus	Description	Human Similarity	NCBI accessions
dog (Canis familiaris)	N4BP1¹	--	NEDD4 binding protein 1	90.69(n) 89.65(a)	478135 XM_535311.2 XP_535311.2
chimpanzee (Pan troglodytes)	N4BP1¹	--	Nedd4 binding protein 1	99.74(n) 99.55(a)	465156 XM_520627.2 XP_520627.2
cow (Bos taurus)	N4BP1¹	--	similar to Nedd4 binding protein 1	90.55(n) 88.58(a)	540948 XM_613819.3 XP_613819.3
rat (Rattus norvegicus)	N4bp1¹	--	Nedd4 binding protein 1	83(n) 81.73(a)	291921 XM_214648.4 XP_214648.4
mouse (Mus musculus)	N4bp1^{1, 5}	8⁵	NEDD4 binding protein 1^{1, 5}	83.28(n)¹ 81.41(a)¹	80750¹ XM_894155.2¹ XP_899248.2¹ AA414154⁵ AA683726⁵ (see all 19)

About this table Species with no ortholog for N4BP1

ENSEMBL Gene Tree for N4BP1

(Paralogs according to ¹HomoloGene
and ²Ensembl, Pseudogenes according to ³Pseudogene.org)
About This Section

Paralogs for N4BP1 gene

KIAA0323²

(According to the ¹NCBI SNP Database, ²Ensembl, ³PupaSUITE, and UniProtKB, Linkage Disequilibrium by HapMap, Genotyping Reagents from Applied Biosystems)
About This Section

10/305 NCBI SNPs in N4BP1 are shown (see all 305 )
(Click for Applied Biosystems TaqMan ® Genotyping Assay) (see all 98)

AB	SNP ID	Valid	Chr 16 pos	Sequence	Recs	AA Chg	Type	More	Recs	Allele freq	Pop	Total sample	More
AB	Genomic Data				Transcription Data				Allele Frequencies
				--	--	--		--	--				--
	rs1039340^1,2	A,C,F,H,O	47132138(+)	`GGAAGA/CGGCGA`	1	--	ut3¹ ese³ trp³		8		MN EU EA WA NA	736
	rs1224^1,2	A,C,F,H,O	47130318(+)	`AAAGCA/GAGACA`	1	--	ut3¹ ese³		12		MN NA EU EA WA	939
	rs3743779^1,2	A,C,F,H	47131541(-)	`CCAGGA/CGGTGT`	1	--	ut3¹ ese³		8		EA NA EU WA	2034
	rs9941275^1,2	A,C,F,H	47130929(+)	`CAGCGA/GGGAGA`	1	--	ut3¹ ese³ trp³		4		EU EA WA	412
--	rs7197732^1,2	A,C,F	47133379(+)	`AGGGCG/ATCTCC`	1	--	ut3¹ ese³		3		NA	142
	rs1039341^1,2	A,C,F	47132370(+)	`ATAGCC/TGAGAA`	1	--	ut3¹ ese³		4		MN NA	324
	rs9936446^1,2	A,C,F	47203308(+)	`ATGACG/CTGGGA`	1	--	ng3¹ tfbs³		4		NA EA	362
--	rs8058362^1,2	A,C	47202817(+)	`TAATTC/TTTCAT`	1	--	ng3¹ tfbs³		0	--	--	--	--
--	rs12709084^1,2	A,C	47202402(+)	`aggcaA/Gaggtt`	1	--	ng3¹		0	--	--	--	--
	rs3743778^1,2	C,F	47131760(+)	`AAGCCC/GCATCG`	1	--	ut3¹ ese³		4		EA NA	886

About this table

HapMap Linkage Disequilibrium images for N4BP1 (up to first 250kb)

(in which this Gene is Involved, According to OMIM, UniProtKB, PharmGKB, Genatlas, GeneTests, Blood group antigen gene mutations by BGMUT, HGMD, GAD, HuGE Navigator, BCGD, and/or TGDB.)
About This Section

Human Genome Epidemiology Navigator: N4BP1 (1 document)

(Possibly Related Articles in Doctor's Guide)
About This Section

(in PubMed. Associations of this gene to articles via ¹Novoseek, ²HGNC, ³Entrez Gene, ⁴UniProtKB/Swiss-Prot, ⁵UniProtKB/TrEMBL, ⁶GAD, and/or ⁷PharmGKB)
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9 PubMed articles for N4BP1 gene:

Prediction of the coding sequences of unidentified human genes. X. The complete sequences of 100 new cDNA clones from brain which can code for large proteins in vitro. (PubMed id 9734811)^{2, 3, 4} Ishikawa K....Ohara O. (1998)
Identification of developmentally expressed proteins that functionally interact with Nedd4 ubiquitin ligase. (PubMed id 11717310)^{2, 3} Murillas R....Kuehn M.R. (2002)
Genetic variants in apoptosis and immunoregulation-related genes are associated with risk of chronic lymphocytic leukemia. (PubMed id 19074885)³ Enjuanes A....Campo E. (2008)
A quantitative atlas of mitotic phosphorylation. (PubMed id 18669648)⁴ Dephoure N.... Gygi S.P. (2008)
ATM and ATR substrate analysis reveals extensive protein networks responsive to DNA damage. (PubMed id 17525332)⁴ Matsuoka S.... Elledge S.J. (2007)
A protein interaction framework for human mRNA degradation. (PubMed id 15231747)³ Lehner B. and Sanderson C.M. (2004)
The status, quality, and expansion of the NIH full-length cDNA project: the Mammalian Gene Collection (MGC). (PubMed id 15489334)⁴ Gerhard D.S....Malek J. (2004)
Complete sequencing and characterization of 21,243 full-length human cDNAs. (PubMed id 14702039)³ Ota T....Sugano S. (2004)
Generation and initial analysis of more than 15,000 full-length human and mouse cDNA sequences. (PubMed id 12477932)³ Strausberg R.L....Marra M.A. (2002)

(in PubMed, OMIM, and NCBI Bookshelf)
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		AND	OR
Aliases
Free Text

Query String		PubMed OMIM NCBI Bookshelf
	(Note: In FireFox, select the above section and copy using Ctrl-C)

(According to Entrez Gene, HGNC, AceView, euGenes, Ensembl, miRBase, ECgene, and/or H-InvDB)
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Entrez Gene: 9683	HGNC: 29850	AceView: N4BP1	Ensembl:ENSG00000102921	euGenes: HUgn9683
ECgene: N4BP1	H-InvDB: N4BP1

(According to HUGE)
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HUGE: KIAA0615

(According to ATLAS, HORDE, IMGT, MTDB, LEIDEN, UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL,
Wikipedia and/or GeneReviews via UniProtKB/Swiss-Prot)
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Licensable Technologies for N4BP1
(Available from WIS Yeda, Salk, Tufts)

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(Reagents available from Applied Biosystems, Antibodies and assays by Cell Signaling Technology, Abcam, Novus Biologicals,
Sigma-Aldrich, R&D Systems, Millipore, Abnova, and/or Invitrogen, Clones available from OriGene,and/or Invitrogen, Drugs and/or compounds by Sigma-Aldrich,
Enzo Life Sciences, and/or Tocris Bioscience)
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Products for N4BP1:

TaqMan ® Gene Expression Assays

TaqMan ® Genotyping Assays

Free SNP selection tool

Invitrogen iPath Pathways	Invitrogen BLOCK-iT™ RNAi
Invitrogen Antibodies	Invitrogen Assays
Invitrogen Clones	Invitrogen Q-PCR Products
Invitrogen Human Recombinant Kinases	Invitrogen Custom Antibody and Peptide Service
Invitrogen Proteins / Assays / Screening Services	Search Invitrogen catalog for N4BP1-related products

	Millipore Custom Antibody & Bulk Services
	Millipore Preclinical / Clinical Development Services
	Millipore Immunoassay Services
	Millipore Target Screening & Profiling Services

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N4BP1 Gene

protein-coding GIFtS: 49

Aliases &
Descriptions
for N4BP1

Summaries
for N4BP1

Genomic Location
for N4BP1

Proteins
for N4BP1

Protein Domains/
Families
for N4BP1

Gene Function
for N4BP1

Pathways & Interactions
for N4BP1

Drugs & Compounds
for N4BP1

Transcripts
for N4BP1

Expression
for N4BP1

Orthologs
for N4BP1

Paralogs
for N4BP1

SNPs/Variants
for N4BP1

Disorders & Mutations
for N4BP1

Medical News
for N4BP1

Publications
for N4BP1

Search for N4BP1

Genome Databases showing N4BP1

Other Databases showing N4BP1

Specialized Databases showing N4BP1

Licensable Technologies
for N4BP1

Services
for N4BP1

Gene Index: 3 5 A B C D E F G H I J K L M N O P Q R S T U V W X Y Z

N4BP1 Gene

protein-coding GIFtS: 49

Aliases & Descriptionsfor N4BP1

Summariesfor N4BP1

Genomic Locationfor N4BP1

Proteinsfor N4BP1

Protein Domains/Familiesfor N4BP1

Gene Functionfor N4BP1

Pathways & Interactionsfor N4BP1

Drugs & Compoundsfor N4BP1

Transcriptsfor N4BP1

Expression for N4BP1

Orthologsfor N4BP1

Paralogsfor N4BP1

SNPs/Variantsfor N4BP1

Disorders & Mutationsfor N4BP1

Medical Newsfor N4BP1

Publicationsfor N4BP1

Search for N4BP1

Genome Databases showing N4BP1

Other Databases showing N4BP1

Specialized Databases showing N4BP1

Licensable Technologiesfor N4BP1

Servicesfor N4BP1

Gene Index: 3 5 A B C D E F G H I J K L M N O P Q R S T U V W X Y Z

Aliases &
Descriptions
for N4BP1

Summaries
for N4BP1

Genomic Location
for N4BP1

Proteins
for N4BP1

Protein Domains/
Families
for N4BP1

Gene Function
for N4BP1

Pathways & Interactions
for N4BP1

Drugs & Compounds
for N4BP1

Transcripts
for N4BP1

Expression
for N4BP1

Orthologs
for N4BP1

Paralogs
for N4BP1

SNPs/Variants
for N4BP1

Disorders & Mutations
for N4BP1

Medical News
for N4BP1

Publications
for N4BP1

Licensable Technologies
for N4BP1

Services
for N4BP1