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Category   Symbol Source: HGNC EntrezGene Ensembl GeneCards RNA genes CroW21
GIFtS

METRNL Gene

protein-coding   GIFtS: 41
GCID: GC17P081037

Meteorin, Glial Cell Differentiation Regulator-Like

  Search for METRNL
at  

(According to 1HGNC, 2Entrez Gene,
3UniProtKB/Swiss-Prot, 4UniProtKB/TrEMBL, 5OMIM, 6GeneLoc, 7Ensembl, 8DME, 9miRBase, 10fRNAdb, 12H-InvDB, 13NCBI, 14NONCODE, and/or 15RNAdb)
About This Section

Aliases
Meteorin, Glial Cell Differentiation Regulator-Like1 2
Meteorin-Like Protein2

External Ids:    HGNC: 275841   Entrez Gene: 2842072   Ensembl: ENSG000001768457   UniProtKB: Q641Q33   

Export aliases for METRNL gene to outside databases

Previous GC identifers: GC17P078631 GC17P076439


(According to Entrez Gene, Tocris Bioscience, Wikipedia's Gene Wiki, PharmGKB,
UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL)
About This Section

GeneCards Summary for METRNL Gene: 
METRNL (meteorin, glial cell differentiation regulator-like) is a protein-coding gene. An important paralog of this gene is METRN.


(According to GeneLoc and/or HGNC, and/or
Entrez Gene (NCBI build 37),
and/or miRBase,
Genomic Views according to UCSC (hg19) and Ensembl (release 73), Regulatory elements and Epigenetics data according to QIAGEN, SABiosciences, and/or SwitchGear Genomics)
About This Section
RefSeq DNA sequence:
NC_000017.10  NT_010663.15  NC_018928.2  
Regulatory elements:
   SABiosciences Regulatory transcription factor binding sites in the METRNL gene promoter:
         SRF   AML1a   SRF (504 AA)   SREBP-1c   LCR-F1   SREBP-1b   N-Myc   AREB6   SREBP-1a   
         Other transcription factors

SwitchGear Promoter luciferase reporter plasmidMETRNL promoter sequence
   Search SABiosciences Chromatin IP Primers for METRNL

Epigenetics:
QIAGEN PyroMark CpG Assay predesigned Pyrosequencing DNA Methylation assays in human, mouse, rat METRNL


Genomic Location:
Genomic View: UCSC Golden Path with GeneCards custom track

Entrez Gene cytogenetic band: 17q25.3   Ensembl cytogenetic band:  17q25.3   HGNC cytogenetic band: 17q25.3

METRNL Gene in genomic location: bands according to Ensembl, locations according to (and/or Entrez Gene and/or Ensembl if different)
METRNL gene location

GeneLoc information about chromosome 17         GeneLoc Exon Structure

GeneLoc location for GC17P081037:  view genomic region     (about GC identifiers)

Start:
81,037,567 bp from pter      End:
81,052,864 bp from pter
Size:
15,298 bases      Orientation:
plus strand

(According to UniProtKB, HORDE, neXtProt, Ensembl, and/or Reactome, Modification sites according to PhosphoSitePlus, Specific Peptides from DME, Protein expression images according to data from SPIRE 1MOPED, 2PaxDb, and 3MAXQB RefSeq according to NCBI, PDB rendering according to OCA and/or Proteopedia, Recombinant Proteins from EMD Millipore, R&D Systems, GenScript, Enzo Life Sciences, OriGene, Novus Biologicals, Sino Biological, ProSpec, and/or Cloud-Clone Corp.,
Biochemical Assays by EMD Millipore, R&D Systems, OriGene, GenScript, Cell Signaling Technology, Enzo Life Sciences, and/or Cloud-Clone Corp., Ontologies according to Gene Ontology Consortium 01 Oct 2013 and Entrez Gene, Antibodies by EMD Millipore, R&D Systems, GenScript, Cell Signaling Technology, OriGene, Novus Biologicals, Thermo Fisher Scientific, LSBio, Abcam, and/or Cloud-Clone Corp.)
About This Section

UniProtKB/Swiss-Prot: METRL_HUMAN, Q641Q3 (See protein sequence)
Recommended Name: Meteorin-like protein precursor  
Size: 311 amino acids; 34398 Da
Subcellular location: Secreted (Potential)
Sequence caution: Sequence=AAH50568.1; Type=Erroneous initiation;
Secondary accessions: Q86VM0

Explore the universe of human proteins at neXtProt for METRNL: NX_Q641Q3

Explore proteomics data for METRNL at MOPED 

Post-translational modifications:

  • View modification sites using PhosphoSitePlus
  • View neXtProt modification sites for NX_Q641Q3

  • METRNL Protein expression data from MOPED1, PaxDb2 and MAXQB3 :    About this image 

    METRNL Protein Expression
    REFSEQ proteins: NP_001004431.1  
    ENSEMBL proteins: 
     ENSP00000315731   ENSP00000458566   ENSP00000460798   ENSP00000458275  

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    Gene Ontology (GO): 1 cellular component term (GO ID links to tree view):    About this table

    GO IDQualified GO termEvidencePubMed IDs
    GO:0005576extracellular region IEA--

    METRNL for ontologies           About GeneDecksing



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    (According to HGNC, IUPHAR, InterPro, ProtoNet, UniProtKB, and/or BLOCKS, Sets of similar genes according to GeneDecks)
    About This Section

    ProtoNet protein and cluster: Q641Q3

    UniProtKB/Swiss-Prot: METRL_HUMAN, Q641Q3
    Similarity: Belongs to the meteorin family


    (According to 1UniProtKB, Genatlas, LifeMap Discovery™, IUBMB, and/or 2DME, Human phenotypes from GenomeRNAi, Animal models from MGI Mar 06 2013, inGenious Targeting Laboratory, genOway,
    bound targets from SABiosciences, miRNA Gene Targets from miRTarBase, shRNA from OriGene, RNAi from EMD Millipore, siRNAs from OriGene, QIAGEN, microRNA from QIAGEN, Gene Editing from DNA2.0, Sirion Biotech, Clones from EMD Millipore, OriGene, SwitchGear Genomics, GenScript, Sino Biological, DNA2.0, Vector BioLabs, and Sirion Biotech, Cell Lines from GenScript, LifeMap BioReagents, In Situ Hybridization Assays from Advanced Cell Diagnostics, Ontologies according to Gene Ontology Consortium 01 Oct 2013 via Entrez Gene.)
    About This Section

    Phenotypes:
         1 GenomeRNAi human phenotype for METRNL:
     Synthetic lethal with Ras 

    Animal Models:
       inGenious Targeting Laboratory - Custom generated mouse model solutions for METRNL 
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    (Pathways according to EMD Millipore, R&D Systems, Cell Signaling Technology, KEGG, PharmGKB, BioSystems, Sino Biological, Reactome, Tocris Bioscience, GeneGo (Thomson Reuters), QIAGEN, and/or UniProtKB, Sets of similar genes according to GeneDecks, Interaction Networks according to SABiosciences, and/or STRING, Interactions according to 1UniProtKB, 2MINT, 3I2D, and/or 4STRING, with links to IntAct and Ensembl, Ontologies according to Gene Ontology Consortium 01 Oct 2013 via Entrez Gene).
    About This Section




    Interactions:

        Search SABiosciences Gene Network CentralTM Interacting Genes and Proteins Networks for METRNL

    (Chemical Compounds according to UniProtKB, Enzo Life Sciences, EMD Millipore, Tocris Bioscience HMDB, BitterDB, and/or Novoseek, Ligands according to IUPHAR, and Drugs according to DrugBank, Enzo Life Sciences, and/or PharmGKB, with drugs/clinical trials/news search links to CenterWatch)
    About This Section
    Browse Small Molecules at EMD Millipore
    Browse drugs & compounds from Enzo Life Sciences

    Browse Tocris compounds for METRNL (METRL)

    Search CenterWatch for drugs/clinical trials and news about METRNL / METRL

    (Secondary structures according to fRNAdb,
    GenBank/EMBL/DDBJ Accessions according to
    Unigene (Build 236 Homo sapiens; Apr 25 2013) or GenBank,
    RefSeq according to Entrez Gene,
    DOTS (version 10), and/or AceView, transcript ids from Ensembl with links to UCSC,
    Conferences by KenesGroup, exon structure from GeneLoc, alternative splicing isoforms according to ASD and/or ECgene,
    RNAi Products from EMD Millipore,
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    About This Section

    REFSEQ mRNAs for METRNL gene: 
    NM_001004431.1  

    Unigene Cluster for METRNL:

    Meteorin, glial cell differentiation regulator-like
    Hs.378821  [show with all ESTs]
    Unigene Representative Sequence: AK093748
    5 Ensembl transcripts including schematic representations, and UCSC links where relevant:
    ENST00000320095(uc002kgh.3) ENST00000570778 ENST00000571814(uc002kgi.3)
    ENST00000571940 ENST00000574053
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    Browse SwitchGear 3'UTR luciferase reporter plasmids
    Inhib. RNA
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      QIAGEN QuantiTect SYBR Green Assays in human, mouse, rat METRNL
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    Additional mRNA sequence: 

    AF086149.1 AK093748.1 BC050568.1 BC082252.1 BC118558.1 BC118638.1 

    5 DOTS entries:

    DT.40120570  DT.100691512  DT.120935027  DT.100060301  DT.100783269 

    24/111 AceView cDNA sequences (see all 111):

    AA454519 BM709698 CF272400 BQ014457 BM710375 BQ772108 BM671791 BU733639 
    BM710224 CD369635 BQ027062 CF264891 BU681631 AI798485 CB850650 BM671640 
    CF264755 BC082252 BQ575901 NM_001004431 AI813302 BU734204 AI879700 BM756962 

    GeneLoc Exon Structure


    (RNA expression data according to H-InvDB, NONCODE, miRBase, and RNAdb, Expression images according to data from BioGPS, Illumina Human BodyMap, and CGAP SAGE, Sets of similar genes according to GeneDecks, in vivo and in vitro expression data from LifeMap Discovery™, plus additional links to Genevestigator, and/or SOURCE, and/or BioGPS, and/or UniProtKB,
    PCR Arrays from SABiosciences, Primers from OriGene, SABiosciences, and/or QIAGEN, In Situ Hybridization Assays from Advanced Cell Diagnostics)
    About This Section

    METRNL expression in normal human tissues (normalized intensities)      METRNL embryonic expression: see
    See probesets specificity/sensitivity at GeneAnnot
    About this imageBioGPS <intensity>2/3
    CGAP TAG: AGCTTCCAGC
    METRNL Expression
    About this image


    METRNL expression in embryonic tissues and stem cells    About this table
    Data from LifeMap, the Embryonic Development and Stem Cells Database 
     5/9 selected tissues (see all 9) fully expand
     
     Neural Tube (Nervous System)    fully expand to see all 5 entries
             Floor Plate Cells Spinal Floor Plate
             Metencephalon
     
     Bone (Muscoskeletal System)    fully expand to see all 4 entries
             Zeugopod Long Bone
             skeleton/cranium/viscerocranium   
     
     Brain (Nervous System)    fully expand to see all 2 entries
             Medulla Oblongata
             brain/hindbrain   
     
     Somite (Muscoskeletal System)    fully expand to see all 2 entries
             skeleton/axial skeleton/cervical region   
     
     Paraxial Mesoderm (Muscoskeletal System)    fully expand to see all 2 entries
             skeleton/axial skeleton/cervical region   

    See METRNL Protein Expression from SPIRE MOPED and PaxDB
    Genevestigator expression for METRNL

    SOURCE GeneReport for Unigene cluster: Hs.378821
        SABiosciences Custom PCR Arrays for METRNL
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    QIAGEN QuantiFast Probe-based Assays in human, mouse, rat METRNL
    In Situ
    Assay Products:
     

     
    Advanced Cell Diagnostics RNAscope RNA in situ hybridization assays for METRNL

    (Orthologs according to 1,2HomoloGene (2older version, for species not in 1newer version), 3euGenes, 4SGD , 5MGI Mar 06 2013, with possible further links to Flybase and/or WormBase, and/or 6Ensembl pan taxonomic compara , Gene Trees according to Ensembl and TreeFam)
    About This Section

    This gene was present in the common ancestor of chordates.

    Orthologs for METRNL gene from 5/12 species (see all 12)    About this table
    Organism Taxonomic
    classification
    Gene Description Human
    Similarity
    Orthology
    Type
    Details
    mouse
    (Mus musculus)
    Mammalia Metrnl1 , 5 meteorin, glial cell differentiation regulator-like1, 5 78.14(n)1
    77.49(a)1
      11 (86.89 cM)5
    2100291  NM_144797.31  NP_659046.11 
     1217015445 
    chicken
    (Gallus gallus)
    Aves METRNL1 meteorin, glial cell differentiation regulator-like 72.49(n)
    75(a)
      417332  XM_415598.3  XP_415598.1 
    lizard
    (Anolis carolinensis)
    Reptilia METRNL6
    --
    Uncharacterized protein
    68(a)
    44(a)
    1 ↔ 1
    possible ortholog
    2(107923143-107957013)
    GL343198.1(638917-649124)
    African clawed frog
    (Xenopus laevis)
    Amphibia 480501722   -- 75.42(n)    48050172 
    zebrafish
    (Danio rerio)
    Actinopterygii Dr.172002 Danio rerio mRNA similar to cDNA sequence BC019776 more 76.3(n)    BC046006.1 


    ENSEMBL Gene Tree for METRNL (if available)
    TreeFam Gene Tree for METRNL (if available)

    (Paralogs according to 1HomoloGene,
    2Ensembl, and 3SIMAP, Pseudogenes according to 4Pseudogene.org Build 68)
    About This Section
    Paralogs for METRNL gene
    METRN2  
    1 SIMAP similar gene for METRNL using alignment to 4 protein entries:     METRL_HUMAN (see all proteins):
    METRN

    METRNL for paralogs           About GeneDecksing



    (SNPs/Variants according to the 1NCBI SNP Database, 2Ensembl, 3PupaSUITE, 4UniProtKB, and DNA2.0, Linkage Disequilibrium by HapMap, Structural Variations(CNVs/InDels/Inversions) from the Database of Genomic Variants, Mutations from the Human Gene Mutation Database (HGMD) and the Locus Specific Mutation Databases (LSDB), Blood group antigen gene mutations by BGMUT, Resequencing Primers from QIAGEN, Cancer Mutation PCR Arrays and Assays and Copy Number PCR Arrays from SABiosciences)
    About This Section

    10/478 SNPs in METRNL are shown (see all 478)    About this table     
    Genomic DataTranscription Related DataAllele Frequencies
    SNP IDValidClinical
    significance
    Chr 17 posSequence#AA
    Chg
    TypeMore#Allele
    freq
    PopTotal
    sample
    More
    ----------
    rs64168561,2
    C,F,H--81035574(+) AGAGGT/CGGCCT 1 -- us2k122Minor allele frequency- C:0.20NS EA NA WA CSA 2344
    rs713765481,2
    C,F--81035656(+) ATGTCC/TGCATC 1 -- us2k13Minor allele frequency- T:0.05NA 124
    rs1388933261,2
    --81035703(+) CCGTGA/GTGTGA 1 -- us2k10--------
    rs75016721,2
    C,F,A,H--81035823(+) AAGCCG/ACAGGT 1 -- us2k118Minor allele frequency- A:0.39NS EA NA WA 2342
    rs1900314581,2
    --81035834(+) CCCTTC/TCCACA 1 -- us2k10--------
    rs75018521,2
    C,F--81035930(+) GCAGAC/GGGGGG 1 -- us2k17Minor allele frequency- G:0.41NA WA EA 366
    rs609757251,2
    C,F--81035979(+) CCAGAC/TCTTGA 1 -- us2k15Minor allele frequency- T:0.42WA NA EA 362
    rs586339061,2
    C,F--81036061(+) GAACAG/CATCCT 1 -- us2k15Minor allele frequency- C:0.42WA NA EA 362
    rs1823198461,2
    --81036096(+) CACTGC/TCATCG 1 -- us2k10--------
    rs1173258781,2
    F--81036175(+) TTTGCC/TGGTGA 1 -- us2k11Minor allele frequency- T:0.02EA 120

    HapMap Linkage Disequilibrium report for METRNL (81037567 - 81052864 bp)

    Structural Variations
         Database of Genomic Variants (DGV) 10/12 variations for METRNL (see all 12):    About this table     
    Variant IDTypeSubtypePubMed ID
    dgv3374n71CNV Loss21882294
    dgv3372n71CNV Loss21882294
    nsv522768CNV Loss19592680
    dgv3373n71CNV Loss21882294
    nsv909283CNV Loss21882294
    nsv828147CNV Loss20364138
    nsv529045CNV Loss19592680
    nsv828149CNV Gain20364138
    nsv819941CNV Gain19587683
    nsv828150CNV Gain20364138

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    (in which this Gene is Involved, According to MalaCards, OMIM, UniProtKB, the University of Copenhagen DISEASES database, Conferences by KenesGroup, Genatlas, GeneTests, GAD, HuGE Navigator, and/or TGDB.)
    About This Section
      --

    (in PubMed. Associations of this gene to articles via 1Entrez Gene, 2UniProtKB/Swiss-Prot, 3HGNC, 4GAD, 5PharmGKB, 6HMDB, 7DrugBank, 8UniProtKB/TrEMBL, 9 Novoseek, and/or 10fRNAdb)
    About This Section

    PubMed articles for METRNL gene integrated from 9 sources:
    (articles sorted by number of sources associating them with METRNL)
        Utopia: connect your pdf to the dynamic
    world of online information

    1. The status, quality, and expansion of the NIH full-length cDNA project: the Mammalian Gene Collection (MGC). (PubMed id 15489334)2 Gerhard D.S....Malek J. (2004)
    2. Complete sequencing and characterization of 21,243 full-length human cDNAs. (PubMed id 14702039)1 Ota T.... Sugano S. (2004)
    3. Generation and initial analysis of more than 15,000 full-length human and mouse cDNA sequences. (PubMed id 12477932)1 Strausberg R.L....Marra M.A. (2002)

    (in PubMed, OMIM, and NCBI Bookshelf)
    About This Section
     ANDOR
    Aliases
    Free Text  

      Query String
    PubMed
    OMIM
    NCBI Bookshelf
      (Note: In FireFox, select the above section and copy using Ctrl-C)

    (According to Entrez Gene, HGNC, AceView, euGenes, Ensembl, miRBase, ECgene, Kegg, and/or H-InvDB)
    About This Section
    Entrez Gene: 284207 HGNC: 27584 AceView: LOC284207 Ensembl:ENSG00000176845 euGenes: HUgn284207
    ECgene: METRNL H-InvDB: METRNL

    (According to HUGE)
    About This Section
      --

    (According to PharmGKB, ATLAS, HORDE, IMGT, LEIDEN, UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL,
    Wikipedia and/or GeneReviews via UniProtKB/Swiss-Prot)
    About This Section
    NameDescription
    PharmGKB entry for METRNL Pharmacogenomics, SNPs, Pathways

    (Patent information from GeneIP,
    Licensable technologies from WIS Yeda, Salk, Tufts,
    IP news from LifeMap Sciences, Inc.)
    About This Section
    Patent Information for METRNL gene:
    Search GeneIP for patents involving METRNL

    GeneCards and IP:
    Japan Patent Office Licenses GeneCards     European Patent Office Licenses GeneCards     Improving the IP Search



    (Antibodies, recombinant proteins, and assays from EMD Millipore, R&D Systems, OriGene, QIAGEN, GenScript, Cell Signaling Technology, SABiosciences, Novus Biologicals, Sino Biological, Enzo Life Sciences, Abcam, ProSpec, Cloud-Clone Corp., Thermo Fisher Scientific, LSBio, Gene Editing from DNA2.0 and Sirion Biotech, Clones from EMD Millipore, OriGene, GenScript, Sino Biological, DNA2.0, SwitchGear Genomics, Vector BioLabs, Sirion Biotech, Cell lines from GenScript, and LifeMap BioReagents, PCR Arrays from SABiosciences, Drugs and/or compounds from EMD Millipore, Tocris Bioscience, and/or Enzo Life Sciences, In Situ Hybridization Assays from
    Advanced Cell Diagnostics, Animal models from inGenious Targeting Laboratory, genOway)
    About This Section

     
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    GeneCards Homepage - Last full update: 23 Oct 2013 - Incrementals: 4 Nov 2013 , 7 Nov 2013 , 23 Jan 2014

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    Category
    (GIFtS: 73)
    transforming growth factor, beta 1
    GIFtS Group
    The GeneCards human gene database gene index: 1 3 5 6 A B C D E F G H I J K L M N O P Q R S T U V W X Y Z 


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