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Category Symbol Source: HGNC EntrezGene Ensembl GeneCards RNA genes CroW21

GIFtS
-

METRNL Gene

protein-coding GIFtS: 39
GCID: GC17P081037

meteorin, glial cell differentiation regulator-like

Search for METRNL
in

MalaCards

our new
Human Malady Compendium

(According to ¹HGNC, ²Entrez Gene,
³UniProtKB/Swiss-Prot, ⁴UniProtKB/TrEMBL, ⁵OMIM, ⁶GeneLoc, ⁷Ensembl, ⁸DME, ⁹miRBase, and/or ¹⁰fRNAdb)
About This Section

Aliases
Meteorin, Glial Cell Differentiation Regulator-Like¹ ²
Meteorin-Like Protein²

External Ids:	HGNC: 27584¹	Entrez Gene: 284207²	Ensembl: ENSG00000176845⁷	UniProtKB: Q641Q3³

Export aliases for METRNL gene to outside databases

Previous GC identifers: GC17P078631 GC17P076439

(According to Entrez Gene, Tocris Bioscience, Wikipedia's Gene Wiki, PharmGKB,
UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL)
About This Section

(According to GeneLoc and/or HGNC, and/or
Entrez Gene (NCBI build 37),
and/or miRBase,
Genomic Views according to UCSC (hg19) and Ensembl (release 69), Regulatory elements and Epigenetics data according to QIAGEN, SABiosciences, and/or SwitchGear Genomics)
About This Section

RefSeq DNA sequence:

NC_000017.10 NC_018928.1 NT_010663.15

Regulatory elements:

SABiosciences Regulatory transcription factor binding sites in the METRNL gene promoter:
SRF AML1a SRF (504 AA) SREBP-1c LCR-F1 SREBP-1b N-Myc AREB6 SREBP-1a
Other transcription factors

SwitchGear Promoter luciferase reporter plasmid: METRNL promoter sequence

Search SABiosciences Chromatin IP Primers for METRNL

Epigenetics:

QIAGEN PyroMark CpG Assay predesigned Pyrosequencing DNA Methylation assays in human, mouse, rat METRNL

Genomic Location:
Genomic View: UCSC Golden Path with GeneCards custom track

Entrez Gene cytogenetic band: 17q25.3 Ensembl cytogenetic band: 17q25.3 HGNC cytogenetic band: 17q25.3

METRNL Gene in genomic location: bands according to Ensembl, locations according to (and/or Entrez Gene and/or Ensembl if different)
METRNL gene location

GeneLoc information about chromosome 17 GeneLoc Exon Structure

GeneLoc location for GC17P081037: view genomic region (about GC identifiers)

Start:	81,037,567 bp from pter	End:	81,052,864 bp from pter
Size:	15,298 bases	Orientation:	plus strand

(According to ¹UniProtKB, HORDE, neXtProt, Ensembl, and/or Reactome, Modification sites according to ²PhosphoSitePlus, Specific Peptides from DME, Protein expression images according to data from SPIRE MOPED and PaxDb, RefSeq according to NCBI, PDB rendering according to OCA and/or Proteopedia, Recombinant Proteins from EMD Millipore, R&D Systems, GenScript, Enzo Life Sciences, OriGene, Novus Biologicals, Sino Biological, ProSpec, and/or Uscn,
Biochemical Assays by EMD Millipore, R&D Systems, OriGene, GenScript, Cell Signaling Technology, Enzo Life Sciences, and/or Uscn, Ontologies according to Gene Ontology Consortium 01 Mar 2013 and Entrez Gene, Antibodies by EMD Millipore, R&D Systems, GenScript, Cell Signaling Technology, OriGene, Novus Biologicals, Thermo Fisher Scientific, Abcam, and/or Uscn)
About This Section

UniProtKB/Swiss-Prot: METRL_HUMAN, Q641Q3 (See protein sequence)

Recommended Name: Meteorin-like protein precursor

Size: 311 amino acids; 34398 Da

Subcellular location: Secreted (Potential)

Sequence caution: Sequence=AAH50568.1; Type=Erroneous initiation;

Secondary accessions: Q86VM0

Explore the universe of human proteins at neXtProt for METRNL: NX_Q641Q3

Post-translational modifications:

View modification sites using PhosphoSitePlus²

View neXtProt modification sites for NX_Q641Q3

METRNL Protein expression data from MOPED and PaxDb: About this image

REFSEQ proteins: NP_001004431.1

ENSEMBL proteins:

ENSP00000315731 ENSP00000458566 ENSP00000460798 ENSP00000458275

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Gene Ontology (GO): 1 cellular component term (GO ID links to tree view): About this table

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0005576	extracellular region	IEA	--

METRNL for ontologies About GeneDecksing

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(According to InterPro, ProtoNet, UniProtKB, and/or BLOCKS, Sets of similar genes according to GeneDecks)
About This Section

ProtoNet protein and cluster: Q641Q3

UniProtKB/Swiss-Prot: METRL_HUMAN, Q641Q3

Similarity: Belongs to the meteorin family

(According to ¹UniProtKB, Genatlas, LifeMap Discovery™, IUBMB, and/or ²DME, Human phenotypes from GenomeRNAi, Animal models from MGI Mar 06 2013,
bound targets from SABiosciences, miRNA Gene Targets from miRTarBase shRNA from OriGene, RNAi from EMD Millipore, siRNAs from OriGene, QIAGEN, microRNA from QIAGEN, Gene Editing from DNA2.0, Clones from EMD Millipore, OriGene, SwitchGear Genomics, GenScript, Sino Biological, DNA2.0, and Vector BioLabs, Cell Lines from GenScript, LifeMap BioReagents, In Situ Hybridization Assays from Advanced Cell Diagnostics, Ontologies according to Gene Ontology Consortium 01 Mar 2013 via Entrez Gene.)
About This Section

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1 GenomeRNAi human phenotype for METRNL:

Synthetic lethal with Ras

(Pathways according to EMD Millipore, R&D Systems, Cell Signaling Technology, KEGG, PharmGKB, BioSystems, Reactome, Tocris Bioscience, GeneGo (Thomson Reuters), QIAGEN, and/or UniProtKB, Sets of similar genes according to GeneDecks, Interaction Networks according to SABiosciences, and/or STRING, Interactions according to ¹UniProtKB, ²MINT, ³I2D, and/or ⁴STRING, with links to IntAct and Ensembl, Ontologies according to Gene Ontology Consortium 01 Mar 2013 via Entrez Gene).
About This Section

Interactions:

Search SABiosciences Gene Network Central^TM Interacting Genes and Proteins Networks for METRNL

(Chemical Compounds according to UniProtKB, Enzo Life Sciences, EMD Millipore, Tocris Bioscience HMDB, BitterDB, and/or Novoseek, and Drugs according to DrugBank, Enzo Life Sciences, and/or PharmGKB, with drugs/clinical trials/news search links to CenterWatch)
About This Section

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(Secondary structures according to fRNAdb,
GenBank/EMBL/DDBJ Accessions according to
Unigene (Build 235 Homo sapiens; Mar 10 2013) or GenBank,
RefSeq according to Entrez Gene,
DOTS (version 10), and/or AceView, transcript ids from Ensembl with links to UCSC,
exon structure from GeneLoc, alternative splicing isoforms according to ASD and/or ECgene,
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About This Section

REFSEQ mRNAs for METRNL gene:

NM_001004431.1

Unigene Cluster for METRNL:

Meteorin, glial cell differentiation regulator-like

Hs.378821 [show with all ESTs]

Unigene Representative Sequence: AK093748

5 Ensembl transcripts including schematic representations, and UCSC links where relevant:

ENST00000320095(uc002kgh.3) ENST00000570778 ENST00000571814(uc002kgi.3)

ENST00000571940 ENST00000574053

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Additional cDNA sequence:

AF086149.1 AK093748.1 BC050568.1 BC082252.1 BC118558.1 BC118638.1

5 DOTS entries:

DT.40120570 DT.100691512 DT.120935027 DT.100060301 DT.100783269

24/111 AceView cDNA sequences (see all 111):

BM671791 BQ014457 CF264891 BU733639 BM710375 CF272400 BQ772108 NM_001004431

CD369635 AA454519 BU734204 CB850650 BM710224 BQ575901 AI798485 AI879700

CF264755 BU681631 BC082252 BM671640 BQ027062 AI813302 BM709698 AX748286

GeneLoc Exon Structure

(RNA expression data according to H-InvDB, NONCODE, miRBase, and RNAdb, Expression images according to data from BioGPS, Illumina Human BodyMap, and CGAP SAGE, Sets of similar genes according to GeneDecks, in vivo and in vitro expression data from LifeMap Discovery™, plus additional links to Genevestigator, and/or SOURCE, and/or BioGPS, and/or UniProtKB,
PCR Arrays from SABiosciences, Primers from OriGene, SABiosciences, and/or QIAGEN, In Situ Hybridization Assays from Advanced Cell Diagnostics)
About This Section

METRNL expression in normal human tissues (normalized intensities)
See probesets specificity/sensitivity at GeneAnnot
About this imageBioGPS
CGAP TAG: AGCTTCCAGC

About this image

METRNL expression in embryonic tissues and stem cells

Expression by the Database of Embryonic development, Stem cell research, and Regenerative medicine About this table

8 LifeMap In Vivo Development Anatomical Compartments/Cells

Tissue

Anatomical Compartment

Cell

Category (developmental path)

Spinal Cord

Spinal Floor Plate

Floor Plate Cells

Neural Ectoderm

Bone

Bone

Bone

Brain

Mesencephalic Ventricular Zone

Neural Tube

Metencephalon

Neural Tube

Spinal Cord

Spinal Floor Plate

Spinal Cord

Expression:

Positive

Negative

Selective marker

Experimental details:

Curated

Microarrays

In-situ hybridization

See METRNL Protein Expression from SPIRE MOPED and PaxDB
Genevestigator expression for METRNL

SOURCE GeneReport for Unigene cluster: Hs.378821

SABiosciences Custom PCR Arrays for METRNL

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In Situ
Assay Products:

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(Orthologs according to ^1,2HomoloGene (²older version, for species not in ¹newer version), ³euGenes, ⁴SGD , ⁵MGI Mar 06 2013, with possible further links to Flybase and/or WormBase, and/or ⁶Ensembl pan taxonomic compara , Gene Trees according to Ensembl and TreeFam)
About This Section

This gene was present in the common ancestor of animals.

Orthologs for METRNL gene from 5/15 species (see all 15) About this table

Organism	Taxonomic classification	Gene	Description	Human Similarity	Orthology Type	Details
chicken (Gallus gallus)	Aves	METRNL¹	meteorin, glial cell differentiation regulator-like	72.49(n) 75(a)		417332 XM_415598.3 XP_415598.1
lizard (Anolis carolinensis)	Reptilia	METRNL⁶	--	69(a)	1 ↔ 1	2(107924834-107956652)
African clawed frog (Xenopus laevis)	Amphibia	48050172²	--	75.42(n)		48050172
zebrafish (Danio rerio)	Actinopterygii	Dr.17200²	Danio rerio mRNA similar to cDNA sequence BC019776 more	76.3(n)		BC046006.1
honey bee (Apis mellifera)	Insecta	--	--	24(a)	1 → many	Group14.1(27428-28452)

ENSEMBL Gene Tree for METRNL (if available)
TreeFam Gene Tree for METRNL (if available)

(Paralogs according to ¹HomoloGene,
²Ensembl, and ³SIMAP, Pseudogenes according to ⁴Pseudogene.org Build 68)
About This Section

Paralogs for METRNL gene

METRN²

1 SIMAP similar gene for METRNL using alignment to 4 protein entries: METRL_HUMAN (see all proteins):

METRN

METRNL for paralogs About GeneDecksing

(SNPs/Variants according to the ¹NCBI SNP Database, ²Ensembl, ³PupaSUITE, UniProtKB, and DNA2.0, Linkage Disequilibrium by HapMap, Structural Variations(CNVs/InDels/Inversions) from the Database of Genomic Variants, Mutations from the Human Gene Mutation Database (HGMD) and the Locus Specific Mutation Databases (LSDB), Blood group antigen gene mutations by BGMUT, Resequencing Primers from QIAGEN, Cancer Mutation PCR Arrays and Assays and Copy Number PCR Arrays from SABiosciences)
About This Section

SNP ID	Valid	Clinical significance	Chr 17 pos	Sequence	#	AA Chg	Type	More	#	Allele freq	Pop	Total sample	More
Genomic Data					Transcription Related Data				Allele Frequencies
				--	--	--		--	--
rs6416856^1,2	C,F,H,	--	81035574(+)	`AGAGGT/CGGCCT`	1	--	us2k¹		22		NS EA NA WA CSA	2344
rs71376548^1,2	C,F,	--	81035656(+)	`ATGTCC/TGCATC`	1	--	us2k¹		3		NA	124
rs138893326^1,2		--	81035703(+)	`CCGTGA/GTGTGA`	1	--	us2k¹		0	--	--	--	--
rs7501672^1,2	C,F,A,H,	--	81035823(+)	`AAGCCG/ACAGGT`	1	--	us2k¹		18		NS EA NA WA	2342
rs190031458^1,2		--	81035834(+)	`CCCTTC/TCCACA`	1	--	us2k¹		0	--	--	--	--
rs7501852^1,2	C,F,	--	81035930(+)	`GCAGAC/GGGGGG`	1	--	us2k¹		7		NA WA EA	366
rs60975725^1,2	C,F,	--	81035979(+)	`CCAGAC/TCTTGA`	1	--	us2k¹		5		WA NA EA	362
rs58633906^1,2	C,F,	--	81036061(+)	`GAACAG/CATCCT`	1	--	us2k¹		5		WA NA EA	362
rs182319846^1,2		--	81036096(+)	`CACTGC/TCATCG`	1	--	us2k¹		0	--	--	--	--
rs117325878^1,2	F,	--	81036175(+)	`TTTGCC/TGGTGA`	1	--	us2k¹		1		EA	120

HapMap Linkage Disequilibrium report for METRNL (81037567 - 81052864 bp)
Structural Variations (Copy Number Variations, Insertions/Deletions, Inversions)
Database of Genomic Variants (DGV): 4 variations for METRNL
4 CNVs: 67284 59566 73078 73080

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(in which this Gene is Involved, According to MalaCards, OMIM, UniProtKB, the University of Copenhagen DISEASES database, Genatlas, GeneTests, GAD, HuGE Navigator, and/or TGDB.)
About This Section

(in PubMed. Associations of this gene to articles via ¹Entrez Gene, ²UniProtKB/Swiss-Prot, ³HGNC, ⁴GAD, ⁵PharmGKB, ⁶HMDB, ⁷DrugBank, ⁸UniProtKB/TrEMBL, ⁹ Novoseek, and/or ¹⁰fRNAdb)
About This Section

PubMed articles for METRNL gene integrated from 9 sources:
(articles sorted by number of sources associating them with METRNL)

Utopia: connect your pdf to the dynamic
world of online information

The status, quality, and expansion of the NIH full-length cDNA project: the Mammalian Gene Collection (MGC). (PubMed id 15489334)² Gerhard D.S....Malek J. (2004)
Complete sequencing and characterization of 21,243 full-length human cDNAs. (PubMed id 14702039)¹ Ota T.... Sugano S. (2004)
Generation and initial analysis of more than 15,000 full-length human and mouse cDNA sequences. (PubMed id 12477932)¹ Strausberg R.L....Marra M.A. (2002)

(in PubMed, OMIM, and NCBI Bookshelf)
About This Section

		AND	OR
Aliases
Free Text

Query String		PubMed OMIM NCBI Bookshelf
	(Note: In FireFox, select the above section and copy using Ctrl-C)

(According to Entrez Gene, HGNC, AceView, euGenes, Ensembl, miRBase, ECgene, Kegg, and/or H-InvDB)
About This Section

Entrez Gene: 284207	HGNC: 27584	AceView: LOC284207	Ensembl:ENSG00000176845	euGenes: HUgn284207
ECgene: METRNL	H-InvDB: METRNL

(According to HUGE)
About This Section

(According to PharmGKB, ATLAS, HORDE, IMGT, LEIDEN, UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL,
Wikipedia and/or GeneReviews via UniProtKB/Swiss-Prot)
About This Section

Name	Description
PharmGKB entry for METRNL	Pharmacogenomics, SNPs, Pathways

Intellectual Property for METRNL gene
(Patent information from GeneIP, Licensable technologies from WIS Yeda, Salk, Tufts, IP news from LifeMap Sciences, Inc.)

About This Section

Patent Information for METRNL gene:
Search GeneIP for patents involving METRNL

GeneCards and IP:
Japan Patent Office Licenses GeneCards European Patent Office Licenses GeneCards Improving the IP Search

(Antibodies, recombinant proteins, and assays from EMD Millipore, R&D Systems, OriGene, QIAGEN, GenScript, Cell Signaling Technology, SABiosciences, Novus Biologicals, Sino Biological, Enzo Life Sciences, Abcam, ProSpec, Uscn, Thermo Fisher Scientific, Gene Editing from DNA2.0, Clones from EMD Millipore, OriGene, GenScript, Sino Biological, DNA2.0, SwitchGear Genomics, Vector BioLabs, Cell lines from GenScript and LifeMap BioReagents, PCR Arrays from SABiosciences, Drugs and/or compounds from EMD Millipore, Tocris Bioscience, and/or Enzo Life Sciences),
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About This Section


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Search ThermoFisher Antibodies for METRNL

Vector BioLabs ready-to-use adenovirus/AAV for human, mouse, rat METRNL

GeneCards Homepage - Last full update: 19 Mar 2013 - Incrementals: 21 Mar 2013 , 15 Apr 2013 , 26 Apr 2013

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Category			VWF (GIFTS: 73) von Willebrand factor
GIFtS Group

The GeneCards human gene database gene index: 1 3 5 6 A B C D E F G H I J K L M N O P Q R S T U V W X Y Z

The GeneCards Human Gene Database

Developed at the Crown Human Genome Center, Department of Molecular Genetics, the Weizmann Institute of Science

Hot genes Disease genes METRNL gene at Home site.

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METRNL Gene

meteorin, glial cell differentiation regulator-like

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Export list of displayed genes

METRNL Gene

meteorin, glial cell differentiation regulator-like

Aliasesfor METRNL gene

Summariesfor METRNL gene

Genomic Viewsfor METRNL gene

Proteinsfor METRNL gene

Protein Domains /Familiesfor METRNL gene

Functionfor METRNL gene

Pathways & Interactionsfor METRNL gene

Drugs & Compoundsfor METRNL gene

Transcriptsfor METRNL gene

Expression for METRNL gene

Orthologsfor METRNL gene

Paralogsfor METRNL gene

Genomic Variantsfor METRNL gene

Disorders / Diseasesfor METRNL gene

Publicationsfor METRNL gene

External Searches for METRNL gene

Genome Databases showing METRNL gene

Other Databases showing METRNL gene

Specialized Databases showing METRNL gene

Intellectual Propertyfor METRNL gene

Productsfor METRNL gene

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Aliases
for METRNL gene

Summaries
for METRNL gene

Genomic Views
for METRNL gene

Proteins
for METRNL gene

Protein Domains /
Families
for METRNL gene

Function
for METRNL gene

Pathways & Interactions
for METRNL gene

Drugs & Compounds
for METRNL gene

Transcripts
for METRNL gene

Expression
for METRNL gene

Orthologs
for METRNL gene

Paralogs
for METRNL gene

Genomic Variants
for METRNL gene

Disorders / Diseases
for METRNL gene

Publications
for METRNL gene

Intellectual Property
for METRNL gene

Products
for METRNL gene