LILRA4 Gene
protein-coding GIFtS: 51
GCID: GC19M054844
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leukocyte immunoglobulin-like receptor, subfamily A (with...
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Aliases
for LILRA4 gene
(According to
1HGNC,
2Entrez Gene,
3UniProtKB/Swiss-Prot,
4UniProtKB/TrEMBL, 5OMIM, 6GeneLoc,
7Ensembl,
8DME,
9miRBase,
and/or 10fRNAdb)
About This Section
|
| Aliases |
|---|
| Leukocyte Immunoglobulin-Like Receptor, Subfamily A (With TM Domain), Member41 2 | | CD85 Antigen-Like Family Member G2 3 | | ILT71 2 3 | | Leukocyte Immunoglobulin-Like Receptor Subfamily A Member 42 | | CD85g1 2 | | Leukocyte Immunoglobulin-Like Receptor, Subfamily A (Without TM Domain), Member42 | | Immunoglobulin-Like Transcript 72 3 | | CD85g Antigen3 | | ILT-72 3 | | |
Export aliases for LILRA4 gene to outside databasesPrevious GC identifers: GC19M059540 GC19M051166 |
Summaries
for LILRA4 gene(According to Entrez Gene,
Tocris Bioscience,
Wikipedia's
Gene Wiki,
PharmGKB,
UniProtKB/Swiss-Prot,
and/or
UniProtKB/TrEMBL)
About This Section
|
Entrez Gene summary for LILRA4:
This gene encodes an immunoglobulin-like cell surface protein preferentially expressed in plasmacytoid dendritic cells(PDCs). This gene is highly expressed in PDCs, and is found to be rapidly down-regulated by interleukin 3 (IL3). Thisgene is one of the 19 highly related genes that form a leukocyte receptor gene cluster (LRC) at chromosomal region19q13.4. (provided by RefSeq, Jul 2008)
UniProtKB/Swiss-Prot: LIRA4_HUMAN, P59901Function: May act as receptor for class I MHC antigens. Ligand binding leads to the activation of eosinophils and tothe release of RNASE2, IL4 and leukotriene C4
Gene Wiki entry for LILRA4
|
Genomic Views
for LILRA4 gene
(According to
GeneLoc and/or
HGNC, and/or
Entrez Gene (NCBI build 37),
and/or miRBase,
Genomic Views according to
UCSC (hg19) and
Ensembl (release 69),
Regulatory elements and Epigenetics data according to
QIAGEN,
SABiosciences, and/or
SwitchGear Genomics)
About This Section
| RefSeq DNA sequence:- NC_000019.9 NC_018930.1 NT_011109.16
Regulatory elements:
 SABiosciences Regulatory transcription factor binding sites in the LILRA4 gene promoter:
ER-alpha AML1a SREBP-1c SREBP-1b AREB6 SREBP-1a POU2F1 POU2F1a
Other transcription factors
Search SABiosciences Chromatin IP Primers for LILRA4
Epigenetics:
| | QIAGEN PyroMark CpG Assay predesigned Pyrosequencing DNA Methylation assays in human, mouse, rat LILRA4 |
Genomic Location:
Genomic View: UCSC Golden Path with GeneCards custom track
Entrez Gene cytogenetic band: 19q13.4 Ensembl cytogenetic band: 19q13.42 HGNC cytogenetic band: 19q13.4LILRA4 Gene in genomic location: bands according to Ensembl, locations according to
 (and/or Entrez Gene and/or Ensembl if different)
GeneLoc information about chromosome 19 GeneLoc Exon Structure GeneLoc location for GC19M054844: view genomic region
(about GC identifiers)
Start:
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54,844,456 bp from pter |
End:
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54,850,421 bp from pter |
Size:
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5,966 bases |
Orientation:
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minus strand |
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Proteins
for LILRA4 gene
(According to
1UniProtKB,
HORDE,
neXtProt,
Ensembl,
and/or Reactome,
Modification sites according to 2PhosphoSitePlus,
Specific Peptides from DME,
Protein expression images according to data from SPIRE MOPED and PaxDb,
RefSeq according to NCBI,
PDB rendering according to OCA and/or Proteopedia,
Recombinant Proteins
from
EMD Millipore,
R&D Systems,
GenScript,
Enzo Life Sciences,
OriGene,
Novus Biologicals,
Sino Biological,
ProSpec, and/or
Uscn,
Biochemical Assays by
EMD Millipore,
R&D Systems,
OriGene,
GenScript,
Cell Signaling Technology,
Enzo Life Sciences, and/or
Uscn,
Ontologies according to Gene
Ontology Consortium 01 Mar 2013 and
Entrez Gene,
Antibodies by
EMD Millipore,
R&D Systems,
GenScript,
Cell Signaling Technology,
OriGene,
Novus Biologicals,
Thermo Fisher Scientific,
Abcam, and/or
Uscn)
About This Section
|
UniProtKB/Swiss-Prot: LIRA4_HUMAN, P59901 (See
protein sequence)Recommended Name: Leukocyte immunoglobulin-like receptor subfamily A member 4 precursor Size: 499 amino acids; 55165 Da
Subunit: Interacts with BST2
Subcellular location: Membrane; Single-pass type I membrane protein
Secondary accessions: Q32MC4Alternative splicing: 2 isoforms: P59901-1 P59901-2 Explore the universe of human proteins at neXtProt for LILRA4: NX_P59901
Post-translational modifications:
View modification sites using PhosphoSitePlus2View neXtProt modification sites for NX_P59901
LILRA4 Protein expression data from MOPED and PaxDb: About this image 
REFSEQ proteins: NP_036408.3
ENSEMBL proteins: ENSP00000291759
Human Recombinant Protein Products:
Gene Ontology (GO): 1 cellular component term (GO ID links to tree view): About this table | GO ID | Qualified GO term | Evidence | PubMed IDs |
|---|
| GO:0016021 | integral to membrane |
IEA | -- |
LILRA4 for ontologies About GeneDecksing
LILRA4 Antibody Products:
Assay Products for LILRA4: |
Protein
Domains /
Families
for LILRA4 gene(According to InterPro, ProtoNet,
UniProtKB, and/or BLOCKS,
Sets of similar genes according to GeneDecks)
About This Section
|
LILRA4 for domains About GeneDecksing
4 InterPro domains/families:Graphical View of Domain Structure for InterPro Entry P59901ProtoNet protein and cluster: P59901 1 Blocks protein family: IPB003599 Immunoglobulin subtype
UniProtKB/Swiss-Prot: LIRA4_HUMAN, P59901Similarity: Contains 4 Ig-like C2-type (immunoglobulin-like) domains |
Function
for LILRA4 gene
(According to 1UniProtKB,
Genatlas,
LifeMap Discovery™,
IUBMB, and/or
2DME,
Human phenotypes from GenomeRNAi,
Animal models from MGI Mar 06 2013,
bound targets from SABiosciences,
miRNA Gene Targets from miRTarBase
shRNA from
OriGene,
RNAi from
EMD Millipore,
siRNAs from
OriGene,
QIAGEN,
microRNA from QIAGEN,
Gene Editing from DNA2.0,
Clones from EMD Millipore,
OriGene,
SwitchGear Genomics,
GenScript,
Sino Biological,
DNA2.0,
and Vector BioLabs,
Cell Lines from GenScript,
LifeMap BioReagents,
In Situ Hybridization Assays from Advanced Cell Diagnostics,
Ontologies according to Gene Ontology Consortium 01 Mar 2013 via
Entrez Gene.)
About This Section
|
Function Summary:
UniProtKB/Swiss-Prot: LIRA4_HUMAN, P59901Function: May act as receptor for class I MHC antigens. Ligand binding leads to the activation of eosinophils and tothe release of RNASE2, IL4 and leukotriene C4
Clone
Products: |  | Browse Clones for the Expression of Recombinant Proteins Available from EMD Millipore | |  | OriGene Myc/DDK tagged cDNA clones in CMV expression vector in human, mouse, rat for LILRA4
OriGene untagged cDNA clones in CMV expression vector in human, mouse, rat for LILRA4
OriGene custom cloning services – gene synthesis, subcloning, mutagenesis,
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Synthesis Service for LILRA4 | |  | Vector BioLabs ready-to-use adenovirus/AAV for human, mouse, rat LILRA4 |
In Situ Assay
Products: |
 | Advanced Cell Diagnostics RNAscope RNA in situ hybridization assays for LILRA4 |
Gene Ontology (GO): 2 molecular function terms (GO ID links to tree view): About this table
LILRA4 for ontologies About GeneDecksing
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Pathways & Interactions
for LILRA4 gene
(Pathways according to
EMD Millipore,
R&D Systems,
Cell Signaling Technology,
KEGG,
PharmGKB,
BioSystems,
Reactome,
Tocris Bioscience,
GeneGo (Thomson Reuters),
QIAGEN,
and/or UniProtKB,
Sets of similar genes according to GeneDecks,
Interaction Networks according to
SABiosciences,
and/or STRING,
Interactions according to 1UniProtKB,
2MINT,
3I2D, and/or
4STRING,
with links to IntAct and
Ensembl,
Ontologies according to Gene Ontology Consortium 01 Mar 2013 via
Entrez Gene).
About This Section
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Unified GeneCards pathways About this table
See pathways by source
| Super-pathway | contained gene-specific pathways |
|---|
| 1 | Osteoclast differentiation | |
Pathway sources
See GeneCards unified pathways
Show all pathways
1 
Kegg Pathway (Kegg details for LILRA4):
LILRA4 for pathways About GeneDecksing
Interactions:
Search SABiosciences Gene Network CentralTM Interacting Genes and Proteins Networks for LILRA4
STRING Interaction
Network Preview (showing 5 interactants - click image to see 6)
5/7 Interacting proteins for LILRA4 (P599013 ENSP000002917594) via UniProtKB, MINT, STRING, and/or I2D (see all 7)About this table
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Drugs & Compounds
for LILRA4 gene(Chemical Compounds according to UniProtKB, Enzo Life Sciences,
EMD Millipore, Tocris Bioscience
HMDB,
BitterDB, and/or
Novoseek, and Drugs according to
DrugBank,
Enzo Life Sciences, and/or
PharmGKB, with drugs/clinical trials/news
search links to CenterWatch)
About This Section
|
LILRA4 for compounds About GeneDecksing
 Browse Tocris compounds for LILRA4
1 Novoseek chemical compound relationship for LILRA4 gene About this table
| Compound |
-log (P-Val) |
Hits |
PubMed IDs for Articles with Shared Sentences (# sentences) |
| tyrosine |
16.3 |
2 |
12384430 (1), 16735691 (1) |
Search CenterWatch for drugs/clinical trials and news about LILRA4 / LIRA4
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Transcripts
for LILRA4 gene(Secondary structures according to
fRNAdb,
GenBank/EMBL/DDBJ Accessions according to
Unigene
(Build 235 Homo sapiens; Mar 10 2013) or GenBank,
RefSeq according to Entrez Gene,
DOTS (version 10), and/or
AceView,
transcript ids from Ensembl
with links to UCSC,
exon structure from GeneLoc,
alternative splicing isoforms according to ASD and/or
ECgene,
RNAi Products from
EMD Millipore,
siRNAs from
OriGene,
QIAGEN,
shRNA from
OriGene,
microRNA from QIAGEN,
Tagged/untagged cDNA clones from
OriGene,
SwitchGear Genomics,
GenScript,
DNA2.0,
Vector BioLabs,
Primers from
OriGene,
SABiosciences, and/or
QIAGEN
)
About This Section
|
REFSEQ mRNAs for LILRA4 gene: NM_012276.3 Unigene Cluster for LILRA4: Leukocyte immunoglobulin-like receptor, subfamily A (with TM domain), member 4 Hs.406708 [show with all ESTs]Unigene Representative Sequence: BM2853912 Ensembl transcripts including schematic representations, and UCSC links where relevant: ENST00000291759(uc002qfi.3 uc002qfj.3) ENST00000421056
Clone
Products: | | OriGene Myc/DDK tagged cDNA clones in CMV expression vector in human, mouse, rat for LILRA4
OriGene untagged cDNA clones in CMV expression vector in human, mouse, rat for LILRA4
OriGene custom cloning services – gene synthesis, subcloning, mutagenesis,
variant library, vector shuttling
| | | GenScript: all cDNA clones in your preferred vector: LILRA4 (NM_012276) | | | DNA2.0 Custom Codon Optimized Gene
Synthesis Service for LILRA4 | | | Vector BioLabs ready-to-use adenovirus/AAV for human, mouse, rat LILRA4 |
Additional cDNA sequence: AF041261.1 AF283989.1 BC038829.1 BC109198.1 BC109199.1 5 DOTS entries: DT.95263522 DT.95263521 DT.99934982 DT.121487405 DT.95091468 3 AceView cDNA sequences: NM_012276 BG059554 AF283989
GeneLoc Exon Structure
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Expression
for LILRA4 gene
(RNA expression data according to
H-InvDB,
NONCODE,
miRBase, and
RNAdb,
Expression images according to data from
BioGPS,
Illumina Human BodyMap, and
CGAP
SAGE,
Sets of similar genes according to GeneDecks,
in vivo and in vitro expression data from LifeMap Discovery™,
plus additional links to
Genevestigator, and/or
SOURCE, and/or
BioGPS, and/or
UniProtKB,
PCR Arrays from
SABiosciences,
Primers from
OriGene,
SABiosciences, and/or
QIAGEN,
In Situ Hybridization Assays from Advanced Cell Diagnostics)
About This Section
| LILRA4 expression in normal human tissues (normalized intensities)
See probesets specificity/sensitivity at GeneAnnot
About this imageBioGPS
CGAP TAG: --
About this image
See LILRA4 Protein Expression from SPIRE MOPED and PaxDB
Genevestigator expression for LILRA4
SOURCE GeneReport for Unigene cluster: Hs.406708
UniProtKB/Swiss-Prot: LIRA4_HUMAN, P59901Tissue specificity: Detected in eosinophils, neutrophils and monocytes
SABiosciences Custom PCR Arrays for LILRA4
Primer
Products: | | OriGene genome-wide validated SYBR primer pairs in human, mouse, rat for LILRA4
Browse OriGene validated miRNA SYBR primer pairs
| | | SABiosciences RT2 qPCR Primer Assay in human, mouse / rat LILRA4 | | | QIAGEN QuantiTect SYBR Green Assays in human, mouse, rat LILRA4 | | | QIAGEN QuantiFast Probe-based Assays in human, mouse, rat LILRA4 | In Situ
Assay Products: |
| Advanced Cell Diagnostics RNAscope RNA in situ hybridization assays for LILRA4 |
Orthologs
for LILRA4 gene
(Orthologs according to
1,2HomoloGene (2older version, for species not in 1newer version),
3euGenes,
4SGD
,
5MGI Mar 06 2013,
with possible further links to
Flybase
and/or
WormBase,
and/or
6Ensembl pan taxonomic compara ,
Gene Trees according to Ensembl and
TreeFam)
About This Section
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This gene was present in the common ancestor of chordates.
Orthologs for LILRA4 gene from 2/6 species (see all 6) About this table
ENSEMBL Gene Tree for LILRA4 (if available)
TreeFam Gene Tree for LILRA4 (if available) |
Paralogs
for LILRA4 gene(Paralogs according to
1HomoloGene,
2Ensembl, and 3SIMAP, Pseudogenes according to 4Pseudogene.org Build 68)
About This Section
| Paralogs for LILRA4 gene
- KIR2DL42 LILRB42 LILRB12 LILRA62 IGSF12 LILRA52 KIR3DL12 OSCAR2
- NCR12 ENSG000002177922 TARM12 KIR3DX12 LILRA32 LILRB52 KIR3DL32 LILRA12
- KIR2DL12 LILRB22 A1BG2 KIR2DL22 LILRA22 KIR3DL22 KIR2DL32 FCAR2
- LILRB32 GP62
18/49 SIMAP similar genes for LILRA4 using alignment to 1 protein entry: LIRA4_HUMAN(see all similar genes):LILRA6 LILRB3 LILRA1 LILRA2 LILRA5 LILRA3
LILRB1 LILRB5 LILRB2 MIR cl-10 LILRB4 KIR-K36
KIR412 KIR-K39 KIR-K64 KIR-K9 KIR-K3 KIR-K65
LILRA4 for paralogs About GeneDecksing
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Genomic Variants
for LILRA4 gene(SNPs/Variants according to the
1NCBI SNP Database,
2Ensembl,
3PupaSUITE,
UniProtKB, and
DNA2.0,
Linkage Disequilibrium by HapMap,
Structural Variations(CNVs/InDels/Inversions) from the Database of Genomic Variants, Mutations from the Human Gene
Mutation Database (HGMD) and the Locus Specific Mutation
Databases (LSDB), Blood group antigen gene mutations by BGMUT,
Resequencing Primers from QIAGEN,
Cancer Mutation PCR Arrays and Assays and Copy Number PCR Arrays from SABiosciences)
About This Section
|
| Genomic Data | Transcription Related Data | Allele Frequencies | | SNP ID | Valid | Clinical
significance | Chr 19 pos | Sequence | # | AA
Chg | Type | More | # | Allele
freq | Pop | Total
sample | More |
|---|
HapMap Linkage Disequilibrium report for LILRA4 (54844456 - 54850421 bp)
Structural Variations (Copy Number Variations, Insertions/Deletions, Inversions)
Database of Genomic Variants (DGV): 2 variations for LILRA4
2 CNVs: 4088 5116 | SABiosciences Cancer Mutation PCR Assays |
| | QIAGEN SeqTarget long-range PCR primers in human, mouse, rat for resequencing LILRA4 |
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Disorders
/ Diseases
for LILRA4 gene
(in which this Gene is Involved, According to MalaCards,
OMIM, UniProtKB,
the University of Copenhagen DISEASES
database,
Genatlas, GeneTests,
GAD,
HuGE Navigator,
and/or TGDB.)
About This Section
|
LILRA4 for disorders About GeneDecksing
OMIM gene information: 607517
OMIM disorders: --
4  diseases for LILRA4: About MalaCardspituitary adenoma hepatitis c adenoma hepatitis
Human Genome Epidemiology (HuGE) Navigator: LILRA4 (4 documents)
Export disorders for LILRA4 gene to outside databases
|
Publications
for LILRA4 gene (in
PubMed.
Associations of this gene to articles via
1Entrez Gene,
2UniProtKB/Swiss-Prot,
3HGNC,
4GAD,
5PharmGKB,
6HMDB,
7DrugBank,
8UniProtKB/TrEMBL,
9 Novoseek, and/or
10fRNAdb)
About This Section
|
PubMed articles for LILRA4 gene, integrated from 9 sources (see all 17):
(articles sorted by number of sources associating them with LILRA4) | |  | Utopia: connect your pdf to the dynamic
world of online information |
- Regulation of TLR7/9 responses in plasmacytoid dendri tic cells by BST2 and ILT7 receptor interaction. (PubMed id 19564354)1, 2, 9 Cao W....Liu Y.J. (2009)
- Nitroproteins from a human pituitary adenoma tissue discovered with a nitrotyrosine affinity column and tandem mass spectrometry. (PubMed id 16777052)1, 2, 9 Zhan X. and Desiderio D.M. (2006)
- Activation of human eosinophils through leukocyte immunoglobulin-like receptor 7. (PubMed id 12529506)1, 2 Tedla N.... Arm J.P. (2003)
- Extensive gene duplications and a large inversion characterize the human leukocyte receptor cluster. (PubMed id 10941842)1, 3 Wende H.... Ziegler A. (2000)
- Plasmacytoid dendritic cell-specific receptor ILT7-Fc epsilonRI gamma inhibits Toll-like receptor-induced interferon production. (PubMed id 16735691)1, 9 Cao W....Liu Y.J. (2006)
- Immunoglobulin-like transcripts ILT2, ILT3 and ILT7 are expressed by human dendritic cells and down-regulated following activation. (PubMed id 15094202)1, 9 Ju X.S....Zenke M. (2004)
- Subtractive hybridization reveals the expression of immunoglobulin like transcript 7, Eph-B1, granzyme B, and 3 novel transcripts in human plasmacytoid dendritic cells. (PubMed id 12384430)1, 9 Rissoan M.-C.... Bates E.E.M. (2002)
- Large-scale candidate gene analysis of spontaneous cl earance of hepatitis C virus. (PubMed id 20331378)1 Mosbruger T.L....Thio C.L. (2010)
- SAGE library screening reveals ILT7 as a specific plasmacytoid dendritic cell marker that regulates type I IFN production. (PubMed id 18048391)1 Cho M....Kamogawa-Schifter Y. (2008)
- Scan of 977 nonsynonymous SNPs in CLL4 trial patients for the identification of genetic variants influencing prognosis. (PubMed id 18006695)1 Sellick G.S....Houlston R.S. (2008)
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External Searches for LILRA4 gene
(in PubMed,
OMIM, and NCBI Bookshelf)
About This Section
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Genome Databases showing LILRA4 gene
(According to
Entrez Gene,
HGNC,
AceView,
euGenes,
Ensembl,
miRBase,
ECgene,
Kegg,
and/or
H-InvDB)
About This Section
|
|
Other Databases showing LILRA4 gene
(According to HUGE)
About This Section
| -- |
Specialized Databases showing LILRA4 gene(According to PharmGKB,
ATLAS, HORDE, IMGT, LEIDEN, UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL,
Wikipedia and/or GeneReviews via UniProtKB/Swiss-Prot)
About This Section
|
| Name | Description |
| PharmGKB entry for LILRA4 | Pharmacogenomics, SNPs, Pathways |
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| | |
About This Section
| Patent Information for LILRA4 gene:
Search GeneIP for patents involving LILRA4
GeneCards and IP:
Japan Patent Office Licenses GeneCards European Patent Office Licenses GeneCards Improving the IP Search
|
Products
for LILRA4 gene(Antibodies, recombinant proteins, and assays from EMD Millipore, R&D Systems, OriGene, QIAGEN, GenScript, Cell Signaling Technology, SABiosciences, Novus Biologicals, Sino Biological, Enzo Life Sciences, Abcam, ProSpec, Uscn, Thermo Fisher Scientific, Gene Editing from DNA2.0, Clones from EMD Millipore, OriGene, GenScript, Sino Biological, DNA2.0, SwitchGear Genomics, Vector BioLabs, Cell lines from GenScript and LifeMap BioReagents, PCR Arrays from SABiosciences, Drugs and/or compounds from EMD Millipore, Tocris Bioscience, and/or
Enzo Life Sciences),
In Situ Hybridization Assays from
Advanced Cell Diagnostics
About This Section
|
 | |
 | |
 |
| | |  | Browse OriGene Antibodies | | OriGene shRNA RFP for LILRA4 | | | OriGene 29mer shRNA kits in GFP-retroviral vector in human, mouse, rat for LILRA4 | | OriGene genome-wide validated SYBR primer pairs in human, mouse, rat for LILRA4 | | | OriGene Protein Over-expression Lysate for LILRA4 | | Browse OriGene Fluorogenic Cell Assay Kits | | | OriGene siRNA for LILRA4 | | OriGene 3'-UTR Clone for LILRA4 | | | OriGene untagged cDNA clones in CMV expression vector in human, mouse, rat for LILRA4 | | OriGene Myc/DDK tagged cDNA clones in CMV expression vector in human, mouse, rat for LILRA4 | | | Browse OriGene GFP tagged cDNA clones in CMV expression vector | | Browse OriGene MicroRNA Expression Plasmids | | | Browse OriGene basic RS shRNAs | | Browse OriGene validated miRNA SYBR primer pairs | | | Browse OriGene full length recombinant human proteins expressed in human HEK293 cells | | OriGene custom cloning services - gene synthesis, subcloning, mutagenesis,
variant library, vector shuttling | | | OriGene Custom Antibody Services for LILRA4 | | OriGene Custom Protein Services for LILRA4 | | | OriGene Custom Immunoassay Development | | |
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| | | QIAGEN Custom miScript Target Protector blocks miRNA-binding site of in human, mouse, rat LILRA4 | | | QIAGEN SeqTarget long-range PCR primers in human, mouse, rat for resequencing LILRA4 | | | QIAGEN PyroMark CpG Assay predesigned Pyrosequencing DNA Methylation assays in human, mouse, rat LILRA4 | | | QIAGEN FlexiTube/FlexiPlate siRNA for gene silencing in human, mouse, rat LILRA4 | | | QIAGEN QuantiFast Probe-based Assays in human, mouse, rat LILRA4 | | | QIAGEN QuantiTect SYBR Green Assays in human, mouse, rat LILRA4 |
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 | | | LILRA4 Proteins, Antibodies, CLIAs, and ELISAs |
| | | | | Advanced Cell Diagnostics RNAscope RNA in situ hybridization assays for LILRA4 |
|  |  |  | | | | ThermoFisher Antibodies for LILRA4 |
| | | Vector BioLabs ready-to-use adenovirus/AAV for human, mouse, rat LILRA4 |
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