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Category Symbol Source: HGNC EntrezGene Ensembl GeneCards RNA genes CroW21

GIFtS
-

KRT19 Gene

protein-coding GIFtS: 67
GCID: GC17M039679

keratin 19

Explore 141 diseases affiliated with
KRT19 via

MalaCards

our new
Human Malady Compendium

(According to ¹HGNC, ²Entrez Gene,
³UniProtKB/Swiss-Prot, ⁴UniProtKB/TrEMBL, ⁵OMIM, ⁶GeneLoc, ⁷Ensembl, ⁸DME, ⁹miRBase, and/or ¹⁰fRNAdb)
About This Section

Aliases
Keratin 19¹ ²		Cytokeratin 19²
K19¹ ² ³		Cytokeratin-19³
CK19¹ ²		Keratin, Type I Cytoskeletal 19²
K1CS¹ ²		Keratin, Type I, 40-Kd²
CK-19² ³		Keratin-19³
MGC15366¹		Cytokeratin-19³
40-KDa Keratin Intermediate Filament²		Keratin-19³

External Ids:	HGNC: 6436¹	Entrez Gene: 3880²	Ensembl: ENSG00000171345⁷	OMIM: 148020⁵	UniProtKB: P08727³

Export aliases for KRT19 gene to outside databases

Previous GC identifers: GC17M039264 GC17P041708 GC17M039588 GC17M040053 GC17M036933 GC17M035443

(According to Entrez Gene, Tocris Bioscience, Wikipedia's Gene Wiki, PharmGKB,
UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL)
About This Section

Entrez Gene summary for KRT19:

The protein encoded by this gene is a member of the keratin family. The keratins are intermediate filament proteins

responsible for the structural integrity of epithelial cells and are subdivided into cytokeratins and hair keratins.

The type I cytokeratins consist of acidic proteins which are arranged in pairs of heterotypic keratin chains. Unlike

its related family members, this smallest known acidic cytokeratin is not paired with a basic cytokeratin in

epithelial cells. It is specifically expressed in the periderm, the transiently superficial layer that envelopes the

developing epidermis. The type I cytokeratins are clustered in a region of chromosome 17q12-q21. (provided by RefSeq,

Jul 2008)

UniProtKB/Swiss-Prot: K1C19_HUMAN, P08727

Function: Involved in the organization of myofibers. Together with KRT8, helps to link the contractile apparatus to

dystrophin at the costameres of striated muscle

Gene Wiki entry for KRT19 (Keratin 19)

(According to GeneLoc and/or HGNC, and/or
Entrez Gene (NCBI build 37),
and/or miRBase,
Genomic Views according to UCSC (hg19) and Ensembl (release 69), Regulatory elements and Epigenetics data according to QIAGEN, SABiosciences, and/or SwitchGear Genomics)
About This Section

RefSeq DNA sequence:

NC_000017.10 NC_018928.1 NT_010783.15

Regulatory elements:

SABiosciences Regulatory transcription factor binding sites in the KRT19 gene promoter:
PPAR-gamma1 AP-2alpha isoform 2 PPAR-gamma2 HNF-4alpha1
Other transcription factors

SwitchGear Promoter luciferase reporter plasmid: KRT19 promoter sequence

Search SABiosciences Chromatin IP Primers for KRT19

Epigenetics:

QIAGEN PyroMark CpG Assay predesigned Pyrosequencing DNA Methylation assays in human, mouse, rat KRT19

Genomic Location:
Genomic View: UCSC Golden Path with GeneCards custom track

Entrez Gene cytogenetic band: 17q21.2 Ensembl cytogenetic band: 17q21.2 HGNC cytogenetic band: 17q21-q23

KRT19 Gene in genomic location: bands according to Ensembl, locations according to (and/or Entrez Gene and/or Ensembl if different)
KRT19 gene location

GeneLoc information about chromosome 17 GeneLoc Exon Structure

GeneLoc location for GC17M039679: view genomic region (about GC identifiers)

Start:	39,679,869 bp from pter	End:	39,684,641 bp from pter
Size:	4,773 bases	Orientation:	minus strand

(According to ¹UniProtKB, HORDE, neXtProt, Ensembl, and/or Reactome, Modification sites according to ²PhosphoSitePlus, Specific Peptides from DME, Protein expression images according to data from SPIRE MOPED and PaxDb, RefSeq according to NCBI, PDB rendering according to OCA and/or Proteopedia, Recombinant Proteins from EMD Millipore, R&D Systems, GenScript, Enzo Life Sciences, OriGene, Novus Biologicals, Sino Biological, ProSpec, and/or Uscn,
Biochemical Assays by EMD Millipore, R&D Systems, OriGene, GenScript, Cell Signaling Technology, Enzo Life Sciences, and/or Uscn, Ontologies according to Gene Ontology Consortium 01 Mar 2013 and Entrez Gene, Antibodies by EMD Millipore, R&D Systems, GenScript, Cell Signaling Technology, OriGene, Novus Biologicals, Thermo Fisher Scientific, Abcam, and/or Uscn)
About This Section

UniProtKB/Swiss-Prot: K1C19_HUMAN, P08727 (See protein sequence)

Recommended Name: Keratin, type I cytoskeletal 19

Size: 400 amino acids; 44106 Da

Subunit: Heterotetramer of two type I and two type II keratins. Interacts with PNN and the actin-binding domain of DMD.

Interacts with HCV core protein

Developmental stage: Present in hair follicles at all stages of development

Miscellaneous: There are two types of cytoskeletal and microfibrillar keratin: I (acidic; 40-55 kDa) and II (neutral to

basic; 56-70 kDa)

Secondary accessions: B2R874 Q5XG83 Q6NW33 Q7L5M9 Q96A53 Q96FV1 Q9BYF9 Q9P1Y4

Explore the universe of human proteins at neXtProt for KRT19: NX_P08727

Post-translational modifications:

View modification sites using PhosphoSitePlus²

View neXtProt modification sites for NX_P08727

KRT19 Protein expression data from MOPED and PaxDb: About this image

REFSEQ proteins: NP_002267.2

ENSEMBL proteins:

ENSP00000355124 ENSP00000466625 ENSP00000408759

Human Recombinant Protein Products:

	Browse Purified and Recombinant Proteins at EMD Millipore
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	Novus Biologicals KRT19 Protein Novus Biologicals KRT19 Lysates
	Browse Sino Biological Recombinant Proteins
	ProSpec Recombinant Protein for KRT19
	Uscn Proteins for KRT19

Gene Ontology (GO): 5/6 cellular component terms (GO ID links to tree view) (see all 6): About this table

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0005882	intermediate filament	IEA	--
GO:0016010	dystrophin-associated glycoprotein complex	IEA	--
GO:0030018	Z disc	IEA	--
GO:0042383	sarcolemma	IEA	--
GO:0043034	costamere	IDA	16000376

KRT19 for ontologies About GeneDecksing

KRT19 Antibody Products:

	EMD Millipore Mono- and Polyclonal Antibodies for the study of KRT19
	R&D Systems Antibodies for KRT19 (Cytokeratin 19)
	Cell Signaling Technology (CST) Antibodies for KRT19 (K19)
	OriGene Antibodies (see all 10): KRT19 OriGene Custom Antibody Services for KRT19
	GenScript Custom Superior Antibodies Services for KRT19
	Novus Biologicals KRT19 Antibodies
	Abcam antibodies for KRT19
	Uscn Antibodies for KRT19
	ThermoFisher Antibody for KRT19

Assay Products for KRT19:

	Browse Kits and Assays available from EMD Millipore
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	GenScript Custom Assay Services for KRT19
	Browse Enzo Life Sciences for kits & assays
	Uscn ELISAs and CLIAs for KRT19

(According to InterPro, ProtoNet, UniProtKB, and/or BLOCKS, Sets of similar genes according to GeneDecks)
About This Section

KRT19 for domains About GeneDecksing

5 InterPro domains/families:

IPR018039 Intermediate_filament_CS

IPR001664 IF

IPR002957 Keratin_I

IPR009053 Prefoldin

IPR016044 F

Graphical View of Domain Structure for InterPro Entry P08727

ProtoNet protein and cluster: P08727

2 Blocks protein families:

IPB001664 Intermediate filament protein

IPB002957 Type I keratin signature

UniProtKB/Swiss-Prot: K1C19_HUMAN, P08727

Domain: This keratin differs from all other IF proteins in lacking the C-terminal tail domain

Similarity: Belongs to the intermediate filament family

(According to ¹UniProtKB, Genatlas, LifeMap Discovery™, IUBMB, and/or ²DME, Human phenotypes from GenomeRNAi, Animal models from MGI Mar 06 2013,
bound targets from SABiosciences, miRNA Gene Targets from miRTarBase shRNA from OriGene, RNAi from EMD Millipore, siRNAs from OriGene, QIAGEN, microRNA from QIAGEN, Gene Editing from DNA2.0, Clones from EMD Millipore, OriGene, SwitchGear Genomics, GenScript, Sino Biological, DNA2.0, and Vector BioLabs, Cell Lines from GenScript, LifeMap BioReagents, In Situ Hybridization Assays from Advanced Cell Diagnostics, Ontologies according to Gene Ontology Consortium 01 Mar 2013 via Entrez Gene.)
About This Section

Function Summary:

UniProtKB/Swiss-Prot: K1C19_HUMAN, P08727

Function: Involved in the organization of myofibers. Together with KRT8, helps to link the contractile apparatus to

dystrophin at the costameres of striated muscle

Genatlas biochemistry entry for KRT19:

keratin 19,type I (maybe included in KRTI@1),simple epithelia,hair follicle,nail bed

miRNA Products:		OriGene 3'-UTR Clone: KRT19 Browse MicroRNA Expression Plasmids
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Inhib. RNA Products:		Browse for Gene Knock-down Tools from EMD Millipore
		OriGene 29mer shRNA kits in GFP-retroviral vector in human, mouse, rat for KRT19 (see all 7) OriGene shRNA RFP: KRT19 OriGene siRNA: KRT19
		QIAGEN FlexiTube/FlexiPlate siRNA for gene silencing in human, mouse, rat KRT19

Gene Editing
Products:

DNA2.0 Custom Protein Engineering Service for KRT19

Clone Products:		Browse Clones for the Expression of Recombinant Proteins Available from EMD Millipore
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		GenScript: all cDNA clones in your preferred vector: KRT19 (NM_002276)
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Cell Line Products:		GenScript Custom overexpressing Cell Line Services for KRT19
		LifeMap BioReagents: cell line associated with KRT19: PureStem Progenitor E68

In Situ Assay
Products:

Advanced Cell Diagnostics RNAscope RNA in situ hybridization assays for KRT19

Gene Ontology (GO): 5 molecular function terms (GO ID links to tree view): About this table

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0005198	structural molecule activity	--	--
GO:0005200	structural constituent of cytoskeleton	TAS	2448790
GO:0005515	protein binding	IPI	16000376
GO:0008307	structural constituent of muscle	IDA	16000376
GO:0032403	protein complex binding	IEA	--

KRT19 for ontologies About GeneDecksing

Animal Models:
14 MGI mutant phenotypes (inferred from 4 alleles) (MGI details for Krt19):

cardiovascular system	cellular	digestive/alimentary	embryogenesis	endocrine/exocrine gland
growth/size	integument	mortality/aging	nervous system	no phenotypic analysis
normal	reproductive system	skeleton	tumorigenesis

KRT19 for phenotypes About GeneDecksing

(Pathways according to EMD Millipore, R&D Systems, Cell Signaling Technology, KEGG, PharmGKB, BioSystems, Reactome, Tocris Bioscience, GeneGo (Thomson Reuters), QIAGEN, and/or UniProtKB, Sets of similar genes according to GeneDecks, Interaction Networks according to SABiosciences, and/or STRING, Interactions according to ¹UniProtKB, ²MINT, ³I2D, and/or ⁴STRING, with links to IntAct and Ensembl, Ontologies according to Gene Ontology Consortium 01 Mar 2013 via Entrez Gene).
About This Section

Unified GeneCards pathways About this table

See pathways by source

Super-pathway

contained gene-specific pathways

Cytoskeleton remodeling Keratin filaments

Cytoskeleton remodeling Keratin filaments

1.00

Cytoskeleton remodeling_Keratin filaments

0.91

Cytoskeletal Signaling

Cytoskeletal Signaling

1.00

Signaling mediated by p38-alpha and p38-beta

Signaling mediated by p38-alpha and p38-beta

1.00

Pathway sources
See GeneCards unified pathways
Show all pathways

1 EMD Millipore Pathway for KRT19

Cytoskeleton remodeling Keratin filaments

1 Cell Signaling Technology (CST) Pathway for KRT19

Cytoskeletal Signaling

1 GeneGo (Thomson Reuters) Pathway for KRT19

Cytoskeleton remodeling Keratin filaments

1 BioSystems Pathway for KRT19

Signaling mediated by p38-alpha and p38-beta

KRT19 for pathways About GeneDecksing

Interactions:

SABiosciences Gene Network Central^TM Interacting Genes and Proteins Network for KRT19

STRING Interaction Network Preview (showing 5 interactants - click image to see 25)

5/74 Interacting proteins for KRT19 (P08727^{1, 2, 3} ENSP00000355124⁴) via UniProtKB, MINT, STRING, and/or I2D (see all 74)

Interactant		Interaction Details
GeneCard	External ID(s)	Interaction Details
FAM107A	O95990²^,³, ENSP00000354270⁴	MINT-68366 I2D: score=5 STRING: ENSP00000354270
FANCG	O15287²^,³, ENSP00000367910⁴	MINT-68101 I2D: score=5 STRING: ENSP00000367910
KRT15	P19012²^,³, ENSP00000254043⁴	MINT-67018 I2D: score=5 STRING: ENSP00000254043
ZNF638	Q14966²^,³, ENSP00000264447⁴	MINT-67324 I2D: score=5 STRING: ENSP00000264447
EXOC8	Q8IYI6²^,³, ENSP00000353564⁴	MINT-67176 I2D: score=4 STRING: ENSP00000353564

About this table

Gene Ontology (GO): 5 biological process terms (GO ID links to tree view): About this table

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0019048	virus-host interaction	IEA	--
GO:0043627	response to estrogen stimulus	IEP	10037815
GO:0044419	interspecies interaction between organisms	--	--
GO:0045214	sarcomere organization	IDA	16000376
GO:0060706	cell differentiation involved in embryonic placenta development	IEA	--

KRT19 for ontologies About GeneDecksing

(Chemical Compounds according to UniProtKB, Enzo Life Sciences, EMD Millipore, Tocris Bioscience HMDB, BitterDB, and/or Novoseek, and Drugs according to DrugBank, Enzo Life Sciences, and/or PharmGKB, with drugs/clinical trials/news search links to CenterWatch)
About This Section

KRT19 for compounds About GeneDecksing

Browse Small Molecules at EMD Millipore

Browse drugs & compounds from Enzo Life Sciences

Browse Tocris compounds for KRT19
10/53 Novoseek chemical compound relationships for KRT19 gene (see all 53) About this table

Compound	-log (P-Val)	Hits	PubMed IDs for Articles with Shared Sentences (# sentences)
smcc	54	2	14652811 (1)
hematoxylin	48.2	9	11519700 (1), 16006800 (1), 15659042 (1), 17851895 (1) (see all 7)
arotinoid	45.8	1	1698888 (1)
paraffin	39.8	18	7567941 (3), 16911464 (1), 9372313 (1), 15659042 (1) (see all 13)
acitretin	35	1	17637822 (1)
toluene diisocyanate	33.5	14	15478392 (6), 17191304 (2), 19187332 (1)
glycogen	31.6	2	19781766 (1), 15144794 (1)
matrigel	30.5	3	19513553 (1), 12021244 (1)
estrogen	27.9	16	20044607 (2), 11026574 (2), 12481020 (1), 16086826 (1) (see all 12)
b 723	26.8	3	12514785 (1), 12481020 (1), 12647218 (1)

Search CenterWatch for drugs/clinical trials and news about KRT19 / K1C19

(Secondary structures according to fRNAdb,
GenBank/EMBL/DDBJ Accessions according to
Unigene (Build 235 Homo sapiens; Mar 10 2013) or GenBank,
RefSeq according to Entrez Gene,
DOTS (version 10), and/or AceView, transcript ids from Ensembl with links to UCSC,
exon structure from GeneLoc, alternative splicing isoforms according to ASD and/or ECgene,
RNAi Products from EMD Millipore,
siRNAs from OriGene, QIAGEN, shRNA from OriGene, microRNA from QIAGEN,
Tagged/untagged cDNA clones from OriGene, SwitchGear Genomics, GenScript, DNA2.0, Vector BioLabs, Primers from OriGene, SABiosciences, and/or QIAGEN )
About This Section

REFSEQ mRNAs for KRT19 gene:

NM_002276.4

Unigene Cluster for KRT19:

Keratin 19

Hs.654568 [show with all ESTs]

Unigene Representative Sequence: BQ073256

7 Ensembl transcripts including schematic representations, and UCSC links where relevant:

ENST00000361566(uc002hxd.4) ENST00000468880 ENST00000593096 ENST00000471565

ENST00000479031 ENST00000455635 ENST00000462611

miRNA Products:		OriGene 3'-UTR Clone: KRT19 Browse OriGene MicroRNA Expression Plasmids
		QIAGEN Custom miScript Target Protector blocks miRNA-binding site of human, mouse, rat KRT19
		Search QIAGEN for miScript miRNA Assays for microRNAs that regulate KRT19
		Browse SwitchGear 3'UTR luciferase reporter plasmids

Inhib. RNA Products:		Browse for Gene Knock-down Tools from EMD Millipore
		OriGene 29mer shRNA kits in GFP-retroviral vector in human, mouse, rat for KRT19 (see all 7) OriGene shRNA RFP: KRT19 OriGene siRNA: KRT19
		QIAGEN FlexiTube/FlexiPlate siRNA for gene silencing in human, mouse, rat KRT19

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		GenScript: all cDNA clones in your preferred vector: KRT19 (NM_002276)
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		QIAGEN QuantiFast Probe-based Assays in human, mouse, rat KRT19

Additional cDNA sequence:

AB041267.1 AB041268.1 AK313261.1 BC002539.2 BC007628.1 BC010409.2 BC067744.1 BC084574.1

Y00503.1

24/55 DOTS entries (see all 55):

DT.100641390 DT.97863192 DT.102839922 DT.120986422 DT.91777282 DT.102839919 DT.91956476 DT.95308977

DT.91726920 DT.120986466 DT.120986024 DT.120986457 DT.120986319 DT.91911344 DT.97848055 DT.97847997

DT.120986146 DT.91827630 DT.120985849 DT.100670686 DT.121334571 DT.95148013 DT.120986130 DT.91982684

24/112 AceView cDNA sequences (see all 112):

BG752693 BG821421 BQ926703 H26997 BG250957 BG831983 BQ932068 BM809792

BM017426 AI738888 BP368720 BG830765 BQ949556 BQ948762 BE394826 BM019344

BG749554 AA632038 BM562993 H30189 BP351939 BQ278796 BQ962606 BQ685491

GeneLoc Exon Structure

5/7 Alternative Splicing Database (ASD) splice patterns (SP) for KRT19 (see all 7) About this scheme

ExUns:	1a	·	1b	·	1c	^	2	^	3	^	4a	·	4b	^	5a	·	5b	^	6a	·	6b	^	7a	·	7b	·	7c	^	8a	·	8b	·	8c	·	8d	·	8e
SP1:									-		-
SP2:											-
SP3:									-		-										-		-
SP4:
SP5:					-		-		-		-		-		-		-		-		-		-		-

ECgene alternative splicing isoforms for KRT19

(RNA expression data according to H-InvDB, NONCODE, miRBase, and RNAdb, Expression images according to data from BioGPS, Illumina Human BodyMap, and CGAP SAGE, Sets of similar genes according to GeneDecks, in vivo and in vitro expression data from LifeMap Discovery™, plus additional links to Genevestigator, and/or SOURCE, and/or BioGPS, and/or UniProtKB,
PCR Arrays from SABiosciences, Primers from OriGene, SABiosciences, and/or QIAGEN, In Situ Hybridization Assays from Advanced Cell Diagnostics)
About This Section

KRT19 expression in normal human tissues (normalized intensities)
See probesets specificity/sensitivity at GeneAnnot
About this imageBioGPS
CGAP TAG: GACATCAAGT

About this image

KRT19 expression in embryonic tissues and stem cells

Expression by the Database of Embryonic development, Stem cell research, and Regenerative medicine About this table

10/17 LifeMap In Vivo Development Anatomical Compartments/Cells (see all 17)

Tissue

Anatomical Compartment

Cell

Category (developmental path)

Cartilage

Cervical Intervertebral Disc

Intervertebral Disc Nucleus Pulposus Cells

Selective marker (cell-identifying gene)

Cartilage

Lumbar Intervertebral Disc

Intervertebral Disc Nucleus Pulposus Cells

Cartilage

Sacral Intervertebral Disc

Intervertebral Disc Nucleus Pulposus Cells

Cartilage

Thoracic Intervertebral Disc

Intervertebral Disc Nucleus Pulposus Cells

Cartilage

Liver

Intrahepatic Biliary Tree

Biliary Epithelial Cells

Liver

Intrahepatic Biliary Tree

Immature Biliary Epithelial Cells

Liver

Liver Lobule

Hepatocytes

Liver

Intrahepatic Biliary Tree

Oval Cells

Liver

Adipose

Visceral White Adipose

Visceral White Adipocytes

Adipose

Gut Tube

Midgut

Midgut Endoderm Cells

Endoderm

Expression:

Positive

Negative

Selective marker

Experimental details:

Curated

Microarrays

In-situ hybridization

Stem Cell Differentiation: 6 LifeMap Cells

Name

Category

PureStem™ epithelial progenitor E164 (Embryonic Progenitor Cell)

Intermediate Mesoderm, Kidney

PureStem™ progenitor F15 (Embryonic Progenitor Cell)

PureStem™ progenitor RASMO12 (Embryonic Progenitor Cell)

PureStem™ progenitor W10 (Embryonic Progenitor Cell)

Hepatic progenitor cells (Functional hepatocyt...)

Intervertebral disc (IVD) annulus fibrosus (AF) cells (Differentiation of s...)

Cartilage

See KRT19 Protein Expression from SPIRE MOPED and PaxDB
Genevestigator expression for KRT19

SOURCE GeneReport for Unigene cluster: Hs.654568

UniProtKB/Swiss-Prot: K1C19_HUMAN, P08727

Tissue specificity: Expressed in a defined zone of basal keratinocytes in the deep outer root sheath of hair follicles.

Also observed in sweat gland and mammary gland ductal and secretory cells, bile ducts, gastrointestinal tract, bladder

urothelium, oral epithelia, esophagus, ectocervical epithelium (at protein level). Expressed in epidermal basal cells,

in nipple epidermis and a defined region of the hair follicle. Also seen in a subset of vascular wall cells in both

the veins and artery of human umbilical cord, and in umbilical cord vascular smooth muscle. Observed in muscle fibers

accumulating in the costameres of myoplasm at the sarcolemma in structures that contain dystrophin and spectrin

SABiosciences Expression via Pathway-Focused PCR Arrays including KRT19:

Cell Lineage Identification in human mouse rat

Terminal Differentiation Markers in human mouse rat

Breast Cancer in human mouse rat

Epithelial to Mesenchymal Transition (EMT) in human mouse rat

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In Situ
Assay Products:

Advanced Cell Diagnostics RNAscope RNA in situ hybridization assays for KRT19

(Orthologs according to ^1,2HomoloGene (²older version, for species not in ¹newer version), ³euGenes, ⁴SGD , ⁵MGI Mar 06 2013, with possible further links to Flybase and/or WormBase, and/or ⁶Ensembl pan taxonomic compara , Gene Trees according to Ensembl and TreeFam)
About This Section

This gene was present in the common ancestor of chordates.

Orthologs for KRT19 gene from 4/10 species (see all 10) About this table

Organism	Taxonomic classification	Gene	Description	Human Similarity	Orthology Type	Details
chicken (Gallus gallus)	Aves	KRT19¹	keratin 19	73.16(n) 77.72(a)		395861 NM_205009.1 NP_990340.1
lizard (Anolis carolinensis)	Reptilia	KRT19⁶	--	69(a)	1 ↔ 1	6(70650436-70661158)
African clawed frog (Xenopus laevis)	Amphibia	48040981²	--	73.77(n)		48040981
zebrafish (Danio rerio)	Actinopterygii	krt17⁶ si:rp71-1n18.1⁶ (see all 15)	si:rp71-1n18.1 (see all 15)	50(a) 50(a) (see all 15)	many ↔ many many ↔ many (see all 15)	19(5991533-5996775) 22(11885419-11889710)

ENSEMBL Gene Tree for KRT19 (if available)
TreeFam Gene Tree for KRT19 (if available)

(Paralogs according to ¹HomoloGene,
²Ensembl, and ³SIMAP, Pseudogenes according to ⁴Pseudogene.org Build 68)
About This Section

Paralogs for KRT19 gene

KRT17² KRT13² KRT16² KRT26² KRT10² KRT12² KRT18² KRT25²

KRT27² KRT15² KRT20² KRT24² KRT9² KRT28² KRT14² KRT23²

18/59 SIMAP similar genes for KRT19 using alignment to 3 protein entries: K1C19_HUMAN (see all proteins) (see all similar genes):

keratin KRT15 GUCA1B K14 KRT33B KRT31

KRT34 KRT33A KRT12 KRT17 JUP KRT16

KRT35 KRT20 KRT36 KRT24 KRT32 KRT10

KRT19 for paralogs About GeneDecksing

5 Pseudogenes.org Pseudogenes for KRT19
PGOHUM00000238825 PGOHUM00000239898 PGOHUM00000245622 PGOHUM00000245653 PGOHUM00000243669

(SNPs/Variants according to the ¹NCBI SNP Database, ²Ensembl, ³PupaSUITE, UniProtKB, and DNA2.0, Linkage Disequilibrium by HapMap, Structural Variations(CNVs/InDels/Inversions) from the Database of Genomic Variants, Mutations from the Human Gene Mutation Database (HGMD) and the Locus Specific Mutation Databases (LSDB), Blood group antigen gene mutations by BGMUT, Resequencing Primers from QIAGEN, Cancer Mutation PCR Arrays and Assays and Copy Number PCR Arrays from SABiosciences)
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SNP ID	Valid	Clinical significance	Chr 17 pos	Sequence	#	AA Chg	Type	More	#	Pop	Total sample
Genomic Data					Transcription Related Data				Allele Frequencies
				--	--	--		--	--
rs200777034^1,2	C	untested	35657647(+)	`CAGGCC/TTCCGA`	2	S G	mis¹		1	EU	299
rs11550883^1,2	C,F,	untested	35657688(-)	`GTCGCT/CTTTCG`	2	/A	syn¹		8	NA WA CSA EA	367
rs4602^1,2	C,F,A,H,	untested	35657777(-)	`GGGGGG/CCTACG`	2	/A /G	mis¹ ese³		17	MN NA EA NS CSA WA EU	5278
rs4601^1,2	C,F,H,	untested	35660626(-)	`GACAAC/TGCCCG`	2	N	syn¹		20	MN NA WA CSA EA EU	6764
rs56051972^1,2	C,F,	untested	39684598(+)	`GCGGCC/GGGTGG`	1	--	ut5¹		5	NA CSA WA	244
rs113190172^1,2	C	--	35444867(+)	`CAAACC/G/TTTTCA`	1	--	spa¹		1	CSA	1
rs113521369^1,2	C	--	35444869(+)	`AACCTA/G/TTCAAA`	1	--	int¹		1	CSA	1
rs61320338^1,2	F,	--	35445288(+)	`GGCCCG/ACTCTC`	1	--	int¹		1	WA	118
rs113256846^1,2	C,	--	35445497(+)	`CACTAT/CGCGAA`	1	--	int¹		2	NA WA	120
rs113107551^1,2	C,	--	35445880(+)	`GTCACG/CCAGGC`	1	--	int¹		1	NA	2

HapMap Linkage Disequilibrium report for KRT19 (39679869 - 39684641 bp)
Structural Variations (Copy Number Variations, Insertions/Deletions, Inversions)
Database of Genomic Variants (DGV): 5 variations for KRT19
5 CNVs: 3145 37699 30831 8845 4036

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(in which this Gene is Involved, According to MalaCards, OMIM, UniProtKB, the University of Copenhagen DISEASES database, Novoseek, Genatlas, GeneTests, GAD, HuGE Navigator, and/or TGDB.)
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KRT19 for disorders           About GeneDecksing

OMIM gene information: 148020    OMIM disorders: --

20/141 diseases for KRT19 (see all 141):    About MalaCards

von hippel-lindau disease persistent hyperinsulinemic hypoglycemia of infancy pulmonary alveolar proteinosis mixed ductal-endocrine carcinoma

multinodular goiter anal canal adenocarcinoma hyperinsulinemic hypoglycemia graft versus host disease

squamous cell carcinoma non-small cell lung carcinoma pulmonary large cell neuroendocrine carcinoma lichen planus

bile duct adenoma collecting duct carcinoma laryngeal squamous cell carcinoma carcinoma

hypoglycemia of infancy oral lichen planus chromophobe renal cell carcinoma osteofibrous dysplasia

8 diseases from the University of Copenhagen DISEASES database for KRT19:

Carcinoma Lung cancer Breast cancer Papillary thyroid carcinoma

Cholangiocarcinoma Follicular adenoma Hepatocellular carcinoma Adenoma

10/97 Novoseek disease relationships for KRT19 gene (see all 97) About this table

Disease	-log (P-Val)	Hits	PubMed IDs for Articles with Shared Sentences (# sentences)
carcinoma squamous cell	71.4	73	9574953 (4), 8582085 (3), 17129430 (3), 10382940 (2) (see all 54)
adenoma follicular	70.4	20	17308358 (2), 12446919 (2), 19213370 (2), 7520169 (2) (see all 12)
tumors	69.3	425	19956064 (6), 20167095 (5), 15225427 (4), 19341176 (4) (see all 99)
carcinoma papillary	69.1	53	12708901 (6), 12446919 (4), 11038428 (3), 10843292 (3) (see all 26)
carcinoma	68.3	106	18095993 (4), 9833698 (3), 7509852 (2), 15272279 (2) (see all 63)
cholangiocarcinoma	67.6	20	16168519 (3), 19094709 (2), 11743639 (2), 15362741 (2) (see all 12)
thyroid papillary carcinoma	66.6	44	16998461 (4), 11038428 (3), 12746563 (3), 15716612 (3) (see all 20)
nsclc	64.4	73	16809939 (4), 18646695 (4), 18080980 (4), 19662363 (3) (see all 35)
metastasis	64.4	73	14593079 (3), 16788845 (3), 14556657 (2), 10741009 (2) (see all 47)
breast cancer	63	193	15946491 (6), 19087676 (5), 10908152 (5), 9148421 (4) (see all 85)

Genetic Association Database (GAD): KRT19
Human Genome Epidemiology (HuGE) Navigator: KRT19 (3 documents)

Export disorders for KRT19 gene to outside databases

(in PubMed. Associations of this gene to articles via ¹Entrez Gene, ²UniProtKB/Swiss-Prot, ³HGNC, ⁴GAD, ⁵PharmGKB, ⁶HMDB, ⁷DrugBank, ⁸UniProtKB/TrEMBL, ⁹ Novoseek, and/or ¹⁰fRNAdb)
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PubMed articles for KRT19 gene, integrated from 9 sources (see all 1117):
(articles sorted by number of sources associating them with KRT19)

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Characterization of the major physiologic phosphorylation site of human keratin 19 and its role in filament organization. (PubMed id 10212274)^{1, 2, 9} Zhou X.... Omary M.B. (1999)
Specific interaction of the actin-binding domain of dystrophin with intermediate filaments containing keratin 19. (PubMed id 16000376)^{1, 2, 9} Stone M.R.... Bloch R.J. (2005)
Cloning and characterization of the 5'-flanking region of human cytokeratin 19 gene in human cholangiocarcinoma cell line. (PubMed id 11682035)^{1, 2, 9} Kagaya M.... Kobayashi K. (2001)
Genomic organization and amplification of the human keratin 15 and 19 genes. (PubMed id 10623642)^{1, 2, 9} Whittock N.V.... McGrath J.A. (2000)
New consensus nomenclature for mammalian keratins. (PubMed id 16831889)^{1, 3} Schweizer J....Wright M.W. (2006)
The status, quality, and expansion of the NIH full-length cDNA project: the Mammalian Gene Collection (MGC). (PubMed id 15489334)^{1, 2} Gerhard D.S....Malek J. (2004)
Novel single nucleotide polymorphisms of the cytokeratin 19 pseudogene are associated with primary biliary cirrhosis. (PubMed id 12697249)^{1, 4} Daimon Y....Okanoue T. (2003)
Dissection of protein linkage between keratins and pinin, a protein with dual location at desmosome-intermediate filament complex and in the nucleus. (PubMed id 10809736)^{1, 2} Shi J. and Sugrue S.P. (2000)
Microsequences of 145 proteins recorded in the two-dimensional gel protein database of normal human epidermal keratinocytes. (PubMed id 1286667)^{1, 2} Rasmussen H.H....Vandekerckhove J. (1992)
Keratin 19: predicted amino acid sequence and broad tissue distribution suggest it evolved from keratinocyte keratins. (PubMed id 2469734)^{1, 2} Stasiak P.C.... Lane E.B. (1989)

(in PubMed, OMIM, and NCBI Bookshelf)
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		AND	OR
Aliases
Disorders
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(According to Entrez Gene, HGNC, AceView, euGenes, Ensembl, miRBase, ECgene, Kegg, and/or H-InvDB)
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Entrez Gene: 3880	HGNC: 6436	AceView: KRT19	Ensembl:ENSG00000171345	euGenes: HUgn3880
ECgene: KRT19	H-InvDB: KRT19

(According to HUGE)
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(According to PharmGKB, ATLAS, HORDE, IMGT, LEIDEN, UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL,
Wikipedia and/or GeneReviews via UniProtKB/Swiss-Prot)
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Name	Description
PharmGKB entry for KRT19	Pharmacogenomics, SNPs, Pathways
ATLAS Chromosomes in Cancer entry for KRT19	Genetics and Cytogenetics in Oncology and Haematology

Intellectual Property for KRT19 gene
(Patent information from GeneIP, Licensable technologies from WIS Yeda, Salk, Tufts, IP news from LifeMap Sciences, Inc.)

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Patent Information for KRT19 gene:
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GeneCards and IP:
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Cell Line associated with KRT19: PureStem Progenitor E68

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KRT19 Gene

keratin 19

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LUNG TUMOR
HEPATIC NECROSIS MASSIVE
Carcinoma Basal Cell
Oncocytoma
THYROID FOLLICULAR CARCINOMA
dysplasia
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KRT19 Gene

keratin 19

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