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KATNAL1 Gene

protein-coding GIFtS: 47

GC13M029680

katanin p60 subunit A-like 1
Symbol approved by the HUGO Gene Nomenclature Committee (HGNC) database

(According to ¹HGNC, ²Entrez Gene,
³UniProtKB/Swiss-Prot, ⁴UniProtKB/TrEMBL, ⁵OMIM, ⁶GeneLoc , and/or ⁷Ensembl, ⁸miRBase)
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Aliases
EC 3.6.4.3 ^³
MGC2599 ^¹, ²
OTTHUMP00000018194 ^²
OTTHUMP00000042415 ^²

Descriptions
katanin p60 subunit A-like 1 ^², ³
p60 katanin-like 1 ^³

External Ids
HGNC: 28361¹
Entrez Gene: 84056²
UniProtKB: Q9BW62³
Ensembl: ENSG00000102781⁷

Search outside databases for aliases for KATNAL1 gene

Previous GC identifer: GC13M028581

(According to Entrez Gene, Wikipedia's Gene Wiki,
UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL)
About This Section

UniProtKB/Swiss-Prot: KATL1_HUMAN, Q9BW62

Function: Severs microtubules in vitro in an ATP-dependent manner. This activity may promote rapid

reorganization of cellular microtubule arrays (By similarity)

(According to GeneLoc and/or HGNC, and/or
Entrez Gene (NCBI build 36),
and/or miRBase,
Genomic Views according to UCSC and Ensembl, Transcription factor binding sites according to SABiosciences)
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Genomic View:

UCSC Golden Path with GeneCards custom track

Transcription factor binding sites upstream to the KATNAL1 gene

Entrez Gene cytogenetic band: 13q12.3 Ensembl cytogenetic band: 13q12.3 HGNC cytogenetic band: 13q12.3

KATNAL1 Gene in genomic location: bands according to Ensembl, locations according to (and/or Entrez Gene and/or Ensembl if different)

GeneLoc gene densities for chromosome 13 GeneLoc Exon Structure

GeneLoc location for GC13M029680: (about GC identifiers)

Start:	29,674,767 bp from pter
End:	29,779,584 bp from pter
Size:	104,818 bases
Orientation:	minus strand

RefSeq DNA sequence:

NC_000013.9 NT_024524.13

(According to ¹UniProtKB, and/or Ensembl, Phosphorylation sites according to ²Phosphosite, RefSeq according to NCBI, PDB rendering according to OCA and/or Proteopedia, Recombinant Proteins from Invitrogen, Millipore, Sigma-Aldrich, R&D Systems, Enzo Life Sciences, Abnova, OriGene and/or, Abcam,
Biochemical Assays by Invitrogen, Millipore, R&D Systems, Cell Signaling Technology, and/or Enzo Life Sciences, Ontologies according to Gene Ontology Consortium 01 Apr 2009 and Entrez Gene, Antibodies by Invitrogen, Millipore, Sigma-Aldrich, R&D Systems, Cell Signaling Technology, Abcam, Abnova, and/or Novus Biologicals)
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UniProtKB/Swiss-Prot: KATL1_HUMAN, Q9BW62 (See protein sequence)

Recommended Name: Katanin p60 ATPase-containing subunit A-like 1

Size: 490 amino acids; 55392 Da

REFSEQ proteins (2 alternative transcripts):

NP_001014402.1 NP_115492.1

ENSEMBL proteins:

ENSP00000369989 ENSP00000369991

Human Recombinant Proteins

	Browse Drug Discovery Central at Invitrogen for human recombinant proteins
	Browse Purified and Recombinant Proteins at Millipore
	Browse Human Recombinant Proteins at Sigma-Aldrich
	Browse R&D Systems for human recombinant proteins
	Browse recombinant and purified proteins available from Enzo Life Sciences
	Recombinant Proteins from Abcam (KATNAL1)
	Browse Abnova for Human Recombinant Proteins

OriGene Purified Recombinant Human Protein: KATNAL1

1 Gene Ontology (GO) cellular component term (links to tree view):

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0005874	microtubule	IEA	--

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Antibodies for KATNAL1:

	Browse Antibodies Central at Invitrogen
	Browse Millipore's Extensive Line of Mono- and Polyclonal Antibodies
	Browse Antibodies at Sigma-Aldrich
	Browse R&D Systems for Antibodies
	Antibodies from Abcam (KATNAL1), each with their Abpromise^SM
	Browse Abnova for Monoclonal and Polyclonal Antibodies
	Search Novus for antibodies for KATNAL1

Assays for KATNAL1:

	Browse Invitrogen for biochemical assays
	Browse Kits and Assays available from Millipore
	Browse R&D Systems for biochemical assays
	Browse biochemical assays available from Enzo Life Sciences

(According to InterPro, ProtoNet, UniProtKB, and/or BLOCKS, Sets of similar genes according to GeneDecks)
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Graphical View of Domain Structure for InterPro Entry Q9BW62

ProtoNet protein and cluster: Q9BW62

1 Blocks protein family: IPB003960 AAA-protein subdomain

UniProtKB/Swiss-Prot: KATL1_HUMAN, Q9BW62

Similarity: Belongs to the AAA ATPase family

(According to MGI Jun 06 2009, UniProtKB, IUBMB,and/or Genatlas,
shRNA from OriGene, Sigma-Aldrich, RNAi from Sigma-Aldrich,
RNAi Products, Clones, and Q-PCR Products from Invitrogen, Millipore, OriGene, and/or Abnova, siRNAs from Applied Biosystems, SYBR primers from OriGene, Cell-based Assays from Millipore, Ontologies according to Gene Ontology Consortium 01 Apr 2009 via Entrez Gene.)
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Inhib. RNA:		Invitrogen RNAi Products for gene knock-down (KATNAL1)
		Browse for Gene Knock-down Tools from Millipore
		Browse Abnova for Chimera RNAi Products

OriGene 29mer shRNA kit in GFP-retroviral vectors (see all 2): NM_032116

Applied Biosystems Silencer® siRNAs for KATNAL1

Sigma-Aldrich siRNA and siRNA Panels for KATNAL1

Sigma-Aldrich shRNA for KATNAL1

Explore Sigma-Aldrich super-pooled esiRNAs

Clones:		Invitrogen Clones for KATNAL1
		Browse Clones for the Expression of Recombinant Proteins Available from Millipore

OriGene GFP tagged cDNA clones in CMV expression vector (see all 2): NM_001014380
Myc/DDK tagged cDNA clones in CMV expression vector (see all 2): NM_001014380
untagged cDNA clones in CMV expression vector (see all 3): NM_001014380

Primers:

Browse Quantitative PCR Central at Invitrogen for Q-PCR LUX™ Primers

OriGene genome-wide validated SYBR primer pairs: NM_032116

UniProtKB/Swiss-Prot: KATL1_HUMAN, Q9BW62

Function: Severs microtubules in vitro in an ATP-dependent manner. This activity may promote rapid

reorganization of cellular microtubule arrays (By similarity)

Catalytic activity: ATP + H(2)O = ADP + phosphate

Enzyme Number (IUBMB): EC 3.6.4.3

5 Gene Ontology (GO) molecular function terms (links to tree view):

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0000166	nucleotide binding	IEA	--
GO:0005524	ATP binding	IEA	--
GO:0008568	microtubule-severing ATPase activity	IEA	--
GO:0016787	hydrolase activity	IEA	--
GO:0017111	nucleoside-triphosphatase activity	IEA	--

About this table

(Pathways according to Invitrogen (maps by GeneGo), Millipore, Cell Signaling Technology, Sigma-Aldrich, KEGG and/or UniProtKB,
Sets of similar genes according to GeneDecks, Proteins Network according to SABiosciences, Interactions according to ¹UniProtKB, ²MINT, and/or ³STRING, with links to IntAct and Ensembl, Ontologies according to Gene Ontology Consortium 01 Apr 2009 via Entrez Gene.)
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2 Interacting proteins for KATNAL1 (ENSP00000369989³ Q9BW62²) via UniProtKB, MINT, and/or STRING

Interactant		Interaction Details
GeneCard	External ID(s)	Interaction Details
COX17	Q14061²	MINT-66362
USP54	ENSP00000326547³	STRING (score=.712)

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(Chemical Compounds according to UniProtKB, Enzo Life Sciences, Sigma-Aldrich, Tocris Bioscience, and/or Novoseek and Drugs according to Enzo Life Sciences and/or PharmGKB)
About This Section

Browse drugs & compounds from Enzo Life Sciences

Browse Small Molecules at Sigma-Aldrich

Browse Tocris compounds for KATNAL1

(GenBank/EMBL/DDBJ Accessions according to
Unigene (Build 219 Homo sapiens; Jun 2 2009) or GenBank,
RefSeq according to Entrez Gene,
DOTS (version 10), and/or AceView,
non coding RNAs according to RNAdb,
ESTs according to GeneTide,
exon structure from GeneLoc, alternative splicing isoforms according to ASD and/or ECgene,
RNAi Products from Invitrogen, Millipore, and/or Abnova,
siRNAs from Applied Biosystems, Sigma-Aldrich,
shRNA from Sigma-Aldrich, OriGene,
Tagged/untagged cDNA clones from OriGene,
Expression Assays from Applied Biosystems)
About This Section

Inhib. RNA:		Invitrogen RNAi Products for gene knock-down (KATNAL1)
		Browse for Gene Knock-down Tools from Millipore
		Browse Abnova for Chimera RNAi Products

OriGene 29mer shRNA kit in GFP-retroviral vectors (see all 2): NM_032116

Sigma-Aldrich siRNA and siRNA Panels for KATNAL1

Sigma-Aldrich shRNA for KATNAL1

Explore Sigma-Aldrich super-pooled esiRNAs

Applied Biosystems Silencer® siRNAs:

NM_001014380 NM_032116

REFSEQ mRNAs for KATNAL1 gene (2 alternative transcripts):

NM_001014380.1 NM_032116.3

Applied Biosystems TaqMan ® Gene Expression Assays:

NM_001014380 NM_032116

               OriGene GFP tagged cDNA clones in CMV expression vector (see all 2): NM_001014380
                                 Myc/DDK tagged cDNA clones in CMV expression vector (see all 2): NM_001014380
                                 untagged cDNA clones in CMV expression vector (see all 3): NM_001014380

Additional cDNA sequence:

AK097423.1 AK291439.1 AK315724.1 BC000612.2 BX647147.1

4 DOTS entries:

DT.40129064 DT.97798511 DT.70105251 DT.95157638

22 AceView cDNA sequences:

AK097423 BU634530 NM_032116 BC000612 AI379588 AI039250 BE311939 AL704125

AI809648 BU568394 AL702104 BI462044 BX954382 BG913072 BQ772434 BM152637

BU568862 CD694916 AA910028 AA284402 CA419634 AA292530

highest scoring ESTs for KATNAL1:

AI809648 BC000612 BE311939 BE390393 BI462044 AI039250 AI080268 AI379588 AI732299 AI732303

Unigene Cluster for KATNAL1:

Katanin p60 subunit A-like 1

Hs.243596 [show with all ESTs]

Unigene Representative Sequence: NM_032116

GeneLoc Exon Structure

2 Ensembl transcripts including schematic representations:

ENST00000380615 ENST00000380617

(Experimental results according to ¹GeneNote and GNF BioGPS,
probe sets-to-genes annotations according to ²GeneAnnot , ³GeneTide , Sets of similar genes according to GeneDecks, Electronic Northern calculations according to data from UniGene (Build 219 Homo sapiens), SAGE tags according to CGAP, plus additional links to SOURCE, and/or GNF BioGPS, and/or EXPOLDB, and/or UniProtKB,
Expression Assays from Applied Biosystems )
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KATNAL1 expression in normal and diseased human tissues

Applied Biosystems TaqMan ® Gene Expression Assays for KATNAL1

¹ / ² / ³

7 probe-sets matching KATNAL1 gene

Affymetrix probe-set	Array	# genes	Sensitivity	Specificity	Correlation	Length	Gb_Accession	Consensus	Uniqueness	Score	Rank
Affymetrix probe-set	Array	GeneAnnot data			GeneNote data		GeneTide data
78104_at²^{, 3}	U95-D	1	1.00	1.00	0.86	0.99	AI809648	0.80	1.00	0.91	1
67180_at²^{, 3}	U95-D	1	1.00	1.00	0.78	0.84	AI080268	0.60	1.00	0.82	1
78461_at²^{, 3}	U95-E	1	0.75	1.00	0.96	1.17	AI807482	0.60	1.00	0.82	1
227713_at²^{, 3}	U133-B	1	1.00	1.00	--	--	AI807482	0.60	1.00	0.82	1
223790_at²^{, 3}	U133-B	1	1.00	1.00	--	--	BC000612	0.80	1.00	0.91	1
227713_at²	U133Plus2	1	1.00	1.00	--	--	--	--	--	--	--
223790_at²	U133Plus2	1	1.00	1.00	--	--	--	--	--	--	--

GeneDecks KATNAL1 for binary patterns associated with the probe-sets selected above

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Data from

(Publications) and GNF BioGPS
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About these images

CGAP SAGE TAG: TTATGATGGC

SOURCE GeneReport for Unigene cluster: Hs.243596

(Orthologs according to ^1,2HomoloGene (²older version, for species not in ¹newer version), ³euGenes, ⁴SGD and/or ⁵MGI Jun 06 2009, with possible further links to Flybase and/or WormBase, Gene Trees according to Ensembl)
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Orthologs for KATNAL1 gene from 5/13 species (see all 13 )

Organism	Gene	Locus	Description	Human Similarity	NCBI accessions
dog (Canis familiaris)	KATNAL1¹	--	katanin p60 subunit A-like 1	94.76(n) 97.96(a)	486020 XM_543146.2 XP_543146.2
chimpanzee (Pan troglodytes)	KATNAL1¹	--	katanin p60 subunit A-like 1	99.86(n) 99.8(a)	452517 XM_509610.2 XP_509610.2
cow (Bos taurus)	KATNAL1¹	--	katanin p60 subunit A-like 1	90.54(n) 97.14(a)	537739 XM_617920.3 XP_617920.2
rat (Rattus norvegicus)	Katnal1¹	--	katanin p60 subunit A-like 1	84.56(n) 95.49(a)	288449 NM_001006956.1 NP_001006957.1
mouse (Mus musculus)	Katnal1^{1, 5}	5⁵	katanin p60 subunit A-like 1^{1, 5}	85.66(n)¹ 94.67(a)¹	231912¹ NM_153572.1¹ NP_705800.1¹ AK049620⁵ AK180354⁵ (see all 6)

About this table Species with no ortholog for KATNAL1

ENSEMBL Gene Tree for KATNAL1

(Paralogs according to ¹HomoloGene
and ²Ensembl, Pseudogenes according to ³Pseudogene.org)
About This Section

Paralogs for KATNAL1 gene

KATNAL2² KATNA1²

(According to the ¹NCBI SNP Database, ²Ensembl, ³PupaSUITE, and UniProtKB, Linkage Disequilibrium by HapMap, Genotyping Reagents from Applied Biosystems)
About This Section

10/652 NCBI SNPs in KATNAL1 are shown (see all 652 )
(Click for Applied Biosystems TaqMan ® Genotyping Assay) (see all 193)

AB	SNP ID	Valid	Chr 13 pos	Sequence	Recs	AA Chg	Type	More	Recs	Pop	Total sample	More
AB	Genomic Data				Transcription Data				Allele Frequencies
				--	--	--		--	--			--
	rs4769824^1,2	A,C,F,H	29781339(+)	`CATTCC/TTAAAC`	1	--	ng3¹		4	EU EA WA	410
	rs9506275^1,2	A,C,F,H	29679111(+)	`TAGGTC/AAGACT`	2	--	ut3¹ ese³		16	EA NA EU WA	1474
	rs10478^1,2	A,C,F,H	29675202(-)	`ATACAA/GACTAA`	2	--	ut3¹ ese³		9	MN EU EA WA NA	928
	rs3825533^1,2	C,F,H	29679642(-)	`TACTTT/CTACAT`	2	--	ut3¹		11	NA EU EA WA	934
	rs10161651^1,2	C,F,H	29678458(+)	`aGTTAA/CATAGA`	2	--	ut3¹		4	EU EA WA	406
	rs7328946^1,2	C,F,H	29680099(+)	`ATTATT/GAAGTA`	2	--	ut3¹ ese³		9	NA EU EA WA	740
--	rs9551867^1,2	C,F,H	29676032(+)	`AAAAAA/TTTTAA`	2	--	ut3¹		8	EU EA WA	818
	rs4572235^1,2	C,F,H	29675579(+)	`AGGGCG/AATATT`	2	--	ut3¹ ese³		4	EU EA WA	420
	rs12428633^1,2	C,F,H	29676558(+)	`ATTTTA/TTCTCA`	2	--	ut3¹		3	EA WA	296
	rs17689385^1,2	C,F,H	29676781(+)	`ATTTGG/AGTTGT`	2	--	ut3¹		11	NA EU EA WA	978

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HapMap Linkage Disequilibrium images for KATNAL1 (up to first 250kb)

(in which this Gene is Involved, According to OMIM, UniProtKB, PharmGKB, Genatlas, GeneTests, Blood group antigen gene mutations by BGMUT, HGMD, GAD, HuGE Navigator, BCGD, and/or TGDB.)
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(Possibly Related Articles in Doctor's Guide)
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(in PubMed. Associations of this gene to articles via ¹Novoseek, ²HGNC, ³Entrez Gene, ⁴UniProtKB/Swiss-Prot, ⁵UniProtKB/TrEMBL, ⁶GAD, and/or ⁷PharmGKB)
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8 PubMed articles for KATNAL1 gene:

The DNA sequence and analysis of human chromosome 13. (PubMed id 15057823)^{3, 4} Dunham A.... Ross M.T. (2004)
The status, quality, and expansion of the NIH full-length cDNA project: the Mammalian Gene Collection (MGC). (PubMed id 15489334)^{3, 4} Gerhard D.S....Malek J. (2004)
Generation and initial analysis of more than 15,000 full-length human and mouse cDNA sequences. (PubMed id 12477932)^{2, 3} Strausberg R.L....Marra M.A. (2002)
Towards a proteome-scale map of the human protein-protein interaction network. (PubMed id 16189514)³ Rual J.F....Vidal M. (2005)
Complete sequencing and characterization of 21,243 full-length human cDNAs. (PubMed id 14702039)³ Ota T....Sugano S. (2004)
Yeast two-hybrid screens implicate DISC1 in brain development and function. (PubMed id 14623284)³ Millar J.K....Porteous D.J. (2003)
Isolation of novel and known genes from a human fetal cochlear cDNA library using subtractive hybridization and differential screening. (PubMed id 7829101)³ Robertson N.G....Morton C.C. (1994)
Drosophila IKK-related kinase Ik2 and Katanin p60-like 1 regulate dendrite pruning of sensory neuron during metamorphosis. (PubMed id 19329489)¹ Lee H.H....Jan Y.N. (2009)

(in PubMed, OMIM, and NCBI Bookshelf)
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		AND	OR
Aliases
Free Text

Query String		PubMed OMIM NCBI Bookshelf
	(Note: In FireFox, select the above section and copy using Ctrl-C)

(According to Entrez Gene, HGNC, AceView, euGenes, Ensembl, miRBase, ECgene, and/or H-InvDB)
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Entrez Gene: 84056	HGNC: 28361	AceView: KATNAL1	Ensembl:ENSG00000102781	euGenes: HUgn84056
ECgene: KATNAL1	H-InvDB: KATNAL1

(According to HUGE)
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(According to ATLAS, HORDE, IMGT, MTDB, LEIDEN, UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL,
Wikipedia and/or GeneReviews via UniProtKB/Swiss-Prot)
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Licensable Technologies for KATNAL1
(Available from WIS Yeda, Salk, Tufts)

About This Section

(Reagents available from Applied Biosystems, Antibodies and assays by Cell Signaling Technology, Abcam, Novus Biologicals,
Sigma-Aldrich, R&D Systems, Millipore, Abnova, and/or Invitrogen, Clones available from OriGene,and/or Invitrogen, Drugs and/or compounds by Sigma-Aldrich,
Enzo Life Sciences, and/or Tocris Bioscience)
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Products for KATNAL1:

TaqMan ® Gene Expression Assays

TaqMan ® Genotyping Assays

Free SNP selection tool

Invitrogen iPath Pathways	Invitrogen BLOCK-iT™ RNAi
Invitrogen Antibodies	Invitrogen Assays
Invitrogen Clones	Invitrogen Q-PCR Products
Invitrogen Human Recombinant Kinases	Invitrogen Custom Antibody and Peptide Service
Invitrogen Proteins / Assays / Screening Services	Search Invitrogen catalog for KATNAL1-related products

	Millipore Custom Antibody & Bulk Services
	Millipore Preclinical / Clinical Development Services
	Millipore Immunoassay Services
	Millipore Target Screening & Profiling Services

	Predesigned and custom siRNAs for KATNAL1		Browse antibodies at Sigma-Aldrich
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	Lentivirus-delivered shRNAs for KATNAL1		Browse small molecules at Sigma-Aldrich
	"Your Favorite Gene" Pathways		feedback

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	Multiplex/Array Assay Kits & Reagents		ELISpot Kits & Development Modules
	Recombinant & Natural Proteins

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Search Tocris compounds for KATNAL1

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KATNAL1 Gene

protein-coding GIFtS: 47

Aliases &
Descriptions
for KATNAL1

Summaries
for KATNAL1

Genomic Location
for KATNAL1

Proteins
for KATNAL1

Protein Domains/
Families
for KATNAL1

Gene Function
for KATNAL1

Pathways & Interactions
for KATNAL1

Drugs & Compounds
for KATNAL1

Transcripts
for KATNAL1

Expression
for KATNAL1

Orthologs
for KATNAL1

Paralogs
for KATNAL1

SNPs/Variants
for KATNAL1

Disorders & Mutations
for KATNAL1

Medical News
for KATNAL1

Publications
for KATNAL1

Search for KATNAL1

Genome Databases showing KATNAL1

Other Databases showing KATNAL1

Specialized Databases showing KATNAL1

Licensable Technologies
for KATNAL1

Services
for KATNAL1

Gene Index: 3 5 A B C D E F G H I J K L M N O P Q R S T U V W X Y Z

KATNAL1 Gene

protein-coding GIFtS: 47

Aliases & Descriptionsfor KATNAL1

Summariesfor KATNAL1

Genomic Locationfor KATNAL1

Proteinsfor KATNAL1

Protein Domains/Familiesfor KATNAL1

Gene Functionfor KATNAL1

Pathways & Interactionsfor KATNAL1

Drugs & Compoundsfor KATNAL1

Transcriptsfor KATNAL1

Expression for KATNAL1

Orthologsfor KATNAL1

Paralogsfor KATNAL1

SNPs/Variantsfor KATNAL1

Disorders & Mutationsfor KATNAL1

Medical Newsfor KATNAL1

Publicationsfor KATNAL1

Search for KATNAL1

Genome Databases showing KATNAL1

Other Databases showing KATNAL1

Specialized Databases showing KATNAL1

Licensable Technologiesfor KATNAL1

Servicesfor KATNAL1

Gene Index: 3 5 A B C D E F G H I J K L M N O P Q R S T U V W X Y Z

Aliases &
Descriptions
for KATNAL1

Summaries
for KATNAL1

Genomic Location
for KATNAL1

Proteins
for KATNAL1

Protein Domains/
Families
for KATNAL1

Gene Function
for KATNAL1

Pathways & Interactions
for KATNAL1

Drugs & Compounds
for KATNAL1

Transcripts
for KATNAL1

Expression
for KATNAL1

Orthologs
for KATNAL1

Paralogs
for KATNAL1

SNPs/Variants
for KATNAL1

Disorders & Mutations
for KATNAL1

Medical News
for KATNAL1

Publications
for KATNAL1

Licensable Technologies
for KATNAL1

Services
for KATNAL1