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Category   Symbol Source: HGNC EntrezGene Ensembl GeneCards RNA genes CroW21
GIFtS

HHAT Gene

protein-coding   GIFtS: 53
GCID: GC01P210501

hedgehog acyltransferase

 Explore 3 diseases affiliated with
HHAT via our new
 Human Malady Compendium 
Biological research products
for HHAT
    

(According to 1HGNC, 2Entrez Gene,
3UniProtKB/Swiss-Prot, 4UniProtKB/TrEMBL, 5OMIM, 6GeneLoc, 7Ensembl, 8DME, 9miRBase, and/or 10fRNAdb)
About This Section

Aliases
Hedgehog Acyltransferase1 2 3     GUP21
MART21 2 3 5     Rasp1
SKI12 3 5     Sit1
Skn1 2     Ski1
MART-21 3     Melanoma Antigen Recognized By T Cells 22
Melanoma Antigen Recognized By T-Cells 22 3     Protein-Cysteine N-Palmitoyltransferase HHAT2
Skinny Hedgehog Protein 12 3     EC 2.3.1.-3
FLJ107241     

External Ids:    HGNC: 182701   Entrez Gene: 557332   Ensembl: ENSG000000543927   OMIM: 6057435   UniProtKB: Q5VTY93   

Export aliases for HHAT gene to outside databases

Previous GC identifers: GC01P206890 GC01P208568 GC01P181175


(According to Entrez Gene, Tocris Bioscience, Wikipedia's Gene Wiki, PharmGKB,
UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL)
About This Section

Entrez Gene summary for HHAT:
'Skinny hedgehog' (SKI1) encodes an enzyme that acts within the secretory pathway to catalyze amino-terminal
palmitoylation of 'hedgehog' (see MIM 600725).(supplied by OMIM, Jul 2002)

UniProtKB/Swiss-Prot: HHAT_HUMAN, Q5VTY9
Function: Catalyzes N-terminal palmitoylation of SHH; which is required for SHH signaling (By similarity). May bind GTP




(According to GeneLoc and/or HGNC, and/or
Entrez Gene (NCBI build 37),
and/or miRBase,
Genomic Views according to UCSC (hg19) and Ensembl (release 69), Regulatory elements and Epigenetics data according to QIAGEN, SABiosciences, and/or SwitchGear Genomics)
About This Section
RefSeq DNA sequence:
NC_000001.10  NC_018912.1  NT_167186.1  
Regulatory elements:
   SABiosciences Regulatory transcription factor binding sites in the HHAT gene promoter:
         MEF-2A   HOXA3   ATF-2   POU6F1 (c2)   STAT5A   
         Other transcription factors

SwitchGear Promoter luciferase reporter plasmidHHAT promoter sequence
   Search SABiosciences Chromatin IP Primers for HHAT

Epigenetics:
QIAGEN PyroMark CpG Assay predesigned Pyrosequencing DNA Methylation assays in human, mouse, rat HHAT


Genomic Location:
Genomic View: UCSC Golden Path with GeneCards custom track

Entrez Gene cytogenetic band: 1q32   Ensembl cytogenetic band:  1q32.2   HGNC cytogenetic band: 1q32

HHAT Gene in genomic location: bands according to Ensembl, locations according to (and/or Entrez Gene and/or Ensembl if different)
HHAT gene location

GeneLoc information about chromosome 1         GeneLoc Exon Structure

GeneLoc location for GC01P210501:  view genomic region     (about GC identifiers)

Start:
210,501,596 bp from pter      End:
210,849,638 bp from pter
Size:
348,043 bases      Orientation:
plus strand

(According to 1UniProtKB, HORDE, neXtProt, Ensembl, and/or Reactome, Modification sites according to 2PhosphoSitePlus, Specific Peptides from DME, Protein expression images according to data from SPIRE MOPED and PaxDb, RefSeq according to NCBI, PDB rendering according to OCA and/or Proteopedia, Recombinant Proteins from EMD Millipore, R&D Systems, GenScript, Enzo Life Sciences, OriGene, Novus Biologicals, Sino Biological, ProSpec, and/or Uscn,
Biochemical Assays by EMD Millipore, R&D Systems, OriGene, GenScript, Cell Signaling Technology, Enzo Life Sciences, and/or Uscn, Ontologies according to Gene Ontology Consortium 01 Mar 2013 and Entrez Gene, Antibodies by EMD Millipore, R&D Systems, GenScript, Cell Signaling Technology, OriGene, Novus Biologicals, Thermo Fisher Scientific, Abcam, and/or Uscn)
About This Section

UniProtKB/Swiss-Prot: HHAT_HUMAN, Q5VTY9 (See protein sequence)
Recommended Name: Protein-cysteine N-palmitoyltransferase HHAT  
Size: 493 amino acids; 57313 Da
Subcellular location: Endoplasmic reticulum membrane; Multi-pass membrane protein (By similarity)
Secondary accessions: B7Z4D5 B7Z5I1 B7Z868 B7ZA75 D3DT91 F5H444 Q17RZ7 Q4G0K3 Q5CZ95 Q5TGI2 Q9NVH9
Q9Y3N8
Alternative splicing: 7 isoforms:  Q5VTY9-1   Q5VTY9-2   Q5VTY9-3   Q5VTY9-4   Q5VTY9-5   Q5VTY9-6   Q5VTY9-7   (No experimental confirmation available)

Explore the universe of human proteins at neXtProt for HHAT: NX_Q5VTY9

Post-translational modifications:

  • View modification sites using PhosphoSitePlus2
  • View neXtProt modification sites for NX_Q5VTY9

  • HHAT Protein expression data from MOPED and PaxDb:    About this image 
    Estimated protein expression log10 (pmol).

    REFSEQ proteins (6 alternative transcripts): 
    NP_001116306.1  NP_001164035.1  NP_001164051.1  NP_001164058.1  NP_001164059.1  NP_060664.2  

    ENSEMBL proteins: 
     ENSP00000261458   ENSP00000355977   ENSP00000413399   ENSP00000355976   ENSP00000438468  
     ENSP00000416845   ENSP00000444995   ENSP00000442625   ENSP00000375773   ENSP00000308628  
     ENSP00000439229  

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    Novus Biologicals HHAT Lysate
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    Browse ProSpec Recombinant Proteins
    Uscn Proteins for HHAT

    Gene Ontology (GO): 2 cellular component terms (GO ID links to tree view):    About this table

    GO IDQualified GO termEvidencePubMed IDs
    GO:0005789endoplasmic reticulum membrane IEA--
    GO:0016021integral to membrane IEA--


    HHAT for ontologies           About GeneDecksing



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    (According to InterPro, ProtoNet, UniProtKB, and/or BLOCKS, Sets of similar genes according to GeneDecks)
    About This Section

    HHAT for domains           About GeneDecksing

    1 InterPro domain/family:
     IPR004299 MBOAT_fam

    Graphical View of Domain Structure for InterPro Entry Q5VTY9

    ProtoNet protein and cluster: Q5VTY9

    UniProtKB/Swiss-Prot: HHAT_HUMAN, Q5VTY9
    Similarity: Belongs to the membrane-bound acyltransferase family. HHAT subfamily


    (According to 1UniProtKB, Genatlas, LifeMap Discovery™, IUBMB, and/or 2DME, Human phenotypes from GenomeRNAi, Animal models from MGI Mar 06 2013,
    bound targets from SABiosciences, miRNA Gene Targets from miRTarBase shRNA from OriGene, RNAi from EMD Millipore, siRNAs from OriGene, QIAGEN, microRNA from QIAGEN, Gene Editing from DNA2.0, Clones from EMD Millipore, OriGene, SwitchGear Genomics, GenScript, Sino Biological, DNA2.0, and Vector BioLabs, Cell Lines from GenScript, LifeMap BioReagents, In Situ Hybridization Assays from Advanced Cell Diagnostics, Ontologies according to Gene Ontology Consortium 01 Mar 2013 via Entrez Gene.)
    About This Section

    Function Summary:

         UniProtKB/Swiss-Prot: HHAT_HUMAN, Q5VTY9
    Function: Catalyzes N-terminal palmitoylation of SHH; which is required for SHH signaling (By similarity). May bind GTP

    Enzyme Number (IUBMB): EC 2.3.1.-1

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    hsa-miR-4324 hsa-miR-3671 hsa-miR-34c-3p hsa-miR-607 hsa-miR-1283 hsa-miR-522 hsa-miR-432 hsa-miR-224*
    SwitchGear 3'UTR luciferase reporter plasmidHHAT 3' UTR sequence
    Inhib. RNA
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    Gene Ontology (GO): 4 molecular function terms (GO ID links to tree view):    About this table

    GO IDQualified GO termEvidencePubMed IDs
    GO:0005525GTP binding IEA--
    GO:0016409palmitoyltransferase activity ----
    GO:0016740transferase activity ----
    GO:0016746transferase activity, transferring acyl groups IEA--


    HHAT for ontologies           About GeneDecksing


    Animal Models:
         Mouse knock-outs for HHAT: Hhattm2Ptch Hhattm1Ptch
         6 MGI mutant phenotypes (inferred from 2 alleles(MGI details for Hhat):
     embryogenesis  growth/size  limbs/digits/tail  mortality/aging  nervous system 
     skeleton 

    HHAT for phenotypes           About GeneDecksing


    (Pathways according to EMD Millipore, R&D Systems, Cell Signaling Technology, KEGG, PharmGKB, BioSystems, Reactome, Tocris Bioscience, GeneGo (Thomson Reuters), QIAGEN, and/or UniProtKB, Sets of similar genes according to GeneDecks, Interaction Networks according to SABiosciences, and/or STRING, Interactions according to 1UniProtKB, 2MINT, 3I2D, and/or 4STRING, with links to IntAct and Ensembl, Ontologies according to Gene Ontology Consortium 01 Mar 2013 via Entrez Gene).
    About This Section

    Unified GeneCards pathways  About this table 
    See pathways by source

    Super-pathwaycontained gene-specific pathways
    1Signaling events mediated by the Hedgehog family
    Signaling events mediated by the Hedgehog family1.00


    1 BioSystems Pathway for HHAT 
        Signaling events mediated by the Hedgehog family


    Interactions:

        Search SABiosciences Gene Network CentralTM Interacting Genes and Proteins Networks for HHAT

    STRING Interaction Network Preview (showing 2 interactants - click image to see more details)

    5 Interacting proteins for HHAT (Q5VTY93 ENSP000002614584) via UniProtKB, MINT, STRING, and/or I2D
    InteractantInteraction Details
    GeneCardExternal ID(s)
    ACTR5Q9H9F93I2D: score=5 
    ARP102753I2D: score=1 
    DHHO433233I2D: score=1 
    SHHENSP000002972614STRING: ENSP00000297261
    UBCENSP000003448184STRING: ENSP00000344818
    About this table

    Gene Ontology (GO): 3 biological process terms (GO ID links to tree view):    About this table

    GO IDQualified GO termEvidencePubMed IDs
    GO:0007224smoothened signaling pathway ----
    GO:0007275multicellular organismal development IEA--
    GO:0018345protein palmitoylation ----


    HHAT for ontologies           About GeneDecksing



    (Chemical Compounds according to UniProtKB, Enzo Life Sciences, EMD Millipore, Tocris Bioscience HMDB, BitterDB, and/or Novoseek, and Drugs according to DrugBank, Enzo Life Sciences, and/or PharmGKB, with drugs/clinical trials/news search links to CenterWatch)
    About This Section
    Browse Small Molecules at EMD Millipore
    Browse drugs & compounds from Enzo Life Sciences

    Browse Tocris compounds for HHAT

    1 HMDB Compound for HHAT    About this table
    CompoundSynonyms CAS #PubMed Ids
    Guanosine triphosphate5'-GTP (see all 10)86-01-1--
    Search CenterWatch for drugs/clinical trials and news about HHAT 

    (Secondary structures according to fRNAdb,
    GenBank/EMBL/DDBJ Accessions according to
    Unigene (Build 235 Homo sapiens; Mar 10 2013) or GenBank,
    RefSeq according to Entrez Gene,
    DOTS (version 10), and/or AceView, transcript ids from Ensembl with links to UCSC,
    exon structure from GeneLoc, alternative splicing isoforms according to ASD and/or ECgene,
    RNAi Products from EMD Millipore,
    siRNAs from OriGene, QIAGEN, shRNA from OriGene, microRNA from QIAGEN,
    Tagged/untagged cDNA clones from OriGene, SwitchGear Genomics, GenScript, DNA2.0, Vector BioLabs, Primers from OriGene, SABiosciences, and/or QIAGEN )
    About This Section

    REFSEQ mRNAs for HHAT gene (6 alternative transcripts): 
    NM_001122834.2  NM_001170564.1  NM_001170580.1  NM_001170587.1  NM_001170588.1  NM_018194.4  

    Unigene Cluster for HHAT:

    Hedgehog acyltransferase
    Hs.58650  [show with all ESTs]
    Unigene Representative Sequence: BC051191
    11 Ensembl transcripts including schematic representations, and UCSC links where relevant:
    ENST00000261458(uc001hhz.4 uc010pss.2) ENST00000367010(uc021pip.1)
    ENST00000426968 ENST00000367009(uc001hia.4) ENST00000545154(uc010psr.2 uc010pst.2)
    ENST00000413764(uc009xcx.3) ENST00000541565(uc010psq.2) ENST00000537898(uc009xcy.3 uc010psu.2)
    ENST00000391905 ENST00000308852 ENST00000545781

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    hsa-miR-4324 hsa-miR-3671 hsa-miR-34c-3p hsa-miR-607 hsa-miR-1283 hsa-miR-522 hsa-miR-432 hsa-miR-224*
    SwitchGear 3'UTR luciferase reporter plasmidHHAT 3' UTR sequence
    Inhib. RNA
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    Additional cDNA sequence: 

    AK001586.1 AK092186.1 AK295130.1 AK297193.1 AK298991.1 AK299200.1 AK302955.1 AK316190.1 
    AK316524.1 AL049848.1 BC039071.1 BC051191.1 BC117130.1 CR936628.1 

    6 DOTS entries:

    DT.109649  DT.100017974  DT.121393550  DT.95239376  DT.75131978  DT.121393519 

    24/44 AceView cDNA sequences (see all 44):

    CN478989 AA659886 BX117517 BI820245 AL049848 AK092186 BI491131 CK903268 
    BE856979 BI560701 BG702982 AL710925 F37336 BI551726 NM_018194 W92273 
    AK001586 W81463 AU130633 BC039071 BQ722190 BI711404 AA010728 W81493 

    GeneLoc Exon Structure

    4 Alternative Splicing Database (ASD) splice patterns (SP) for HHAT    About this scheme

    ExUns: 1 ^ 2a · 2b · 2c ^ 3 ^ 4 ^ 5 ^ 6 ^ 7a · 7b ^ 8 ^ 9 ^ 10 ^ 11 ^ 12 ^ 13 ^ 14 ^ 15a · 15b
    SP1:        -     -     -     -                                               -                                       
    SP2:                    -     -                                               -                                       
    SP3:                                                                                                                  
    SP4:                                -     -                                                                           


    ECgene alternative splicing isoforms for HHAT

    (RNA expression data according to H-InvDB, NONCODE, miRBase, and RNAdb, Expression images according to data from BioGPS, Illumina Human BodyMap, and CGAP SAGE, Sets of similar genes according to GeneDecks, in vivo and in vitro expression data from LifeMap Discovery™, plus additional links to Genevestigator, and/or SOURCE, and/or BioGPS, and/or UniProtKB,
    PCR Arrays from SABiosciences, Primers from OriGene, SABiosciences, and/or QIAGEN, In Situ Hybridization Assays from Advanced Cell Diagnostics)
    About This Section

    HHAT expression in normal human tissues (normalized intensities)
    See probesets specificity/sensitivity at GeneAnnot
    About this imageBioGPS
    CGAP TAG: GACCAACAGT

    Microarray
    RNAseq (Illumina Body Map)
    (100×FPKM)½
    SAGE (Serial Analysis of Gene Expression)

    About this image
    See HHAT Protein Expression from SPIRE MOPED and PaxDB
    Genevestigator expression for HHAT

    SOURCE GeneReport for Unigene cluster: Hs.58650

    UniProtKB/Swiss-Prot: HHAT_HUMAN, Q5VTY9
    Tissue specificity: Ubiquitously expressed in normal tissues and cancer cell lines

        SABiosciences Expression via Pathway-Focused PCR Array including HHAT: 
              Hedgehog Signaling Pathway in human mouse rat

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    In Situ
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    (Orthologs according to 1,2HomoloGene (2older version, for species not in 1newer version), 3euGenes, 4SGD , 5MGI Mar 06 2013, with possible further links to Flybase and/or WormBase, and/or 6Ensembl pan taxonomic compara , Gene Trees according to Ensembl and TreeFam)
    About This Section

    This gene was present in the common ancestor of eukaryotes.

    Orthologs for HHAT gene from 7/21 species (see all 21)    About this table
    Organism Taxonomic
    classification
    Gene Description Human
    Similarity
    Orthology
    Type
    Details
    chicken
    (Gallus gallus)
    Aves HHAT1 hedgehog acyltransferase 65.99(n)
    65.1(a)
      771269  XM_001234560.1  XP_001234561.1 
    lizard
    (Anolis carolinensis)
    Reptilia --
    --
    --
    65(a)
    62(a)
    1 ↔ many
    1 ↔ many
    1(237390935-237407112)
    1(237261311-237317591)
    zebrafish
    (Danio rerio)
    Actinopterygii hhat1 hedgehog acyltransferase 57.72(n)
    49.27(a)
      560334  NM_001114890.1  NP_001108362.1 
    fruit fly
    (Drosophila melanogaster)
    Insecta rasp1 CG11495-PA 44.02(n)
    28.98(a)
      44098  NM_079174.2  NP_523898.1 
    worm
    (Caenorhabditis elegans)
    Secernentea hhat-11 Protein HHAT-1 40.9(n)
    27.42(a)
      259478  NM_001027318.4  NP_001022489.1 
    thale cress
    (Arabidopsis thaliana)
    eudicotyledons AT1G576006
    MBOAT (membrane bound O-acyl transferase)-like pro...
    17(a)
    1 → many
    1(21330512-21334727)
    rice
    (Oryza sativa)
    Liliopsida --
    O-acyltransferase, putative, expressed
    17(a)
    1 → many
    5(2528880-2536229)


    ENSEMBL Gene Tree for HHAT (if available)
    TreeFam Gene Tree for HHAT (if available) 

    (Paralogs according to 1HomoloGene,
    2Ensembl, and 3SIMAP, Pseudogenes according to 4Pseudogene.org Build 68)
    About This Section
    Paralogs for HHAT gene
    HHATL2  

    HHAT for paralogs           About GeneDecksing



    (SNPs/Variants according to the 1NCBI SNP Database, 2Ensembl, 3PupaSUITE, UniProtKB, and DNA2.0, Linkage Disequilibrium by HapMap, Structural Variations(CNVs/InDels/Inversions) from the Database of Genomic Variants, Mutations from the Human Gene Mutation Database (HGMD) and the Locus Specific Mutation Databases (LSDB), Blood group antigen gene mutations by BGMUT, Resequencing Primers from QIAGEN, Cancer Mutation PCR Arrays and Assays and Copy Number PCR Arrays from SABiosciences)
    About This Section

    10/6974 NCBI SNPs in HHAT are shown (see all 6974    About this table
    Genomic DataTranscription Related DataAllele Frequencies
    SNP IDValidClinical
    significance
    Chr 1 posSequence#AA
    Chg
    TypeMore#Allele
    freq
    PopTotal
    sample
    More
    ----------
    rs348286631,2
    --210499631(+) ATGCCC/TGGCTA 2 -- us2k10--------
    rs1396148891,2
    --210499695(+) GGTCTC/TGATCT 2 -- us2k10--------
    rs1920572681,2
    --210499699(+) TTGATC/TTCCTG 2 -- us2k10--------
    rs1808357521,2
    --210499709(+) GACTTC/TGTGAT 2 -- us2k10--------
    rs75371331,2
    C,--210499721(+) cacccA/Gccttg 2 -- us2k13Minor allele frequency- G:0.17NA WA 6
    rs1464807281,2
    --210499799(+) GCTTTC/TTGCAA 2 -- us2k10--------
    rs75372501,2
    C,F,--210499859(+) TGTCTA/GTAAAT 2 -- us2k15Minor allele frequency- G:0.15NA WA 126
    rs1858193111,2
    --210499868(+) ATGTGC/TCTTGC 2 -- us2k10--------
    rs1133035091,2
    C--210499937(+) TTAAAA/TTTACA 2 -- us2k10--------
    rs114246581,2
    C,--210499939(+) AAATTA/-ACACT 2 -- us2k11Minor allele frequency- -:0.00CSA 2

    HapMap Linkage Disequilibrium report for HHAT (210501596 - 210751596 bp, first 250kb of HHAT)
    Structural Variations (Copy Number Variations, Insertions/Deletions, Inversions)
    Database of Genomic Variants (DGV): 29 variations for HHAT
         15/24 CNVs (see all 24): 34770 58361 97625 84842 74895 84843 38166 34771 65800 9913 30454 74893 84841 23014 2346
         5 Indels: 44503 24053 41776 24052 44504

    SABiosciences Cancer Mutation PCR Assays
    QIAGEN SeqTarget long-range PCR primers in human, mouse, rat for resequencing HHAT
    DNA2.0 Custom Variant and Variant Library Synthesis for HHAT

    (in which this Gene is Involved, According to MalaCards, OMIM, UniProtKB, the University of Copenhagen DISEASES database, Genatlas, GeneTests, GAD, HuGE Navigator, and/or TGDB.)
    About This Section

    HHAT for disorders           About GeneDecksing

    OMIM gene information: 605743    OMIM disorders: --

    3 diseases for HHAT:    About MalaCards
    thoracic outlet syndrome    melanoma    aspergillosis

    2 diseases from the University of Copenhagen DISEASES database for HHAT:
    Aspergillosis     Ancylostomiasis
    Human Genome Epidemiology (HuGE) Navigator: HHAT (2 documents)

    Export disorders for HHAT gene to outside databases

    (in PubMed. Associations of this gene to articles via 1Entrez Gene, 2UniProtKB/Swiss-Prot, 3HGNC, 4GAD, 5PharmGKB, 6HMDB, 7DrugBank, 8UniProtKB/TrEMBL, 9 Novoseek, and/or 10fRNAdb)
    About This Section

    PubMed articles for HHAT gene, integrated from 9 sources (see all 12):
    (articles sorted by number of sources associating them with HHAT)
        Utopia: connect your pdf to the dynamic
    world of online information

    1. Isolation of a new melanoma antigen, MART-2, containing a mutated epitope recognized by autologous tumor-infiltrating T lymphocytes. (PubMed id 11160356)1, 2, 3 Kawakami Y.... Rosenberg S. (2001)
    2. The DNA sequence and biological annotation of human chromosome 1. (PubMed id 16710414)1, 2 Gregory S.G.... Bentley D.R. (2006)
    3. The status, quality, and expansion of the NIH full-length cDNA project: the Mammalian Gene Collection (MGC). (PubMed id 15489334)1, 2 Gerhard D.S....Malek J. (2004)
    4. Complete sequencing and characterization of 21,243 full-length human cDNAs. (PubMed id 14702039)1, 2 Ota T.... Sugano S. (2004)
    5. Skinny hedgehog, an acyltransferase required for palmitoylation and activity of the hedgehog signal. (PubMed id 11486055)1, 2 Chamoun Z.... Basler K. (2001)
    6. SKI-1/S1P inhibition: a promising surrogate to statins to block hepatitis C virus replication. (PubMed id 22626636)1 Blanchet M....Labonte P. (2012)
    7. Identification of N-terminal residues of Sonic Hedgeho g important for palmitoylation by Hedgehog acyltransferase. (PubMed id 23112049)1 Hardy R.Y. and Resh M.D. (2012)
    8. Systematic and quantitative assessment of the ubiquiti n-modified proteome. (PubMed id 21906983)1 Kim W....Gygi S.P. (2011)
    9. Personalized smoking cessation: interactions between nicotine dose, dependence and quit-success genotype score. (PubMed id 20379614)1 Rose J.E....Uhl G.R. (2010)
    10. The full-ORF clone resource of the German cDNA consortium. (PubMed id 17974005)2 Bechtel S.... Schupp I. (2007)

    (in PubMed, OMIM, and NCBI Bookshelf)
    About This Section
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    (According to Entrez Gene, HGNC, AceView, euGenes, Ensembl, miRBase, ECgene, Kegg, and/or H-InvDB)
    About This Section
    Entrez Gene: 55733 HGNC: 18270 AceView: HHAT Ensembl:ENSG00000054392 euGenes: HUgn55733
    ECgene: HHAT H-InvDB: HHAT

    (According to HUGE)
    About This Section
      --

    (According to PharmGKB, ATLAS, HORDE, IMGT, LEIDEN, UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL,
    Wikipedia and/or GeneReviews via UniProtKB/Swiss-Prot)
    About This Section
    NameDescription
    PharmGKB entry for HHAT Pharmacogenomics, SNPs, Pathways
    ATLAS Chromosomes in Cancer entry for HHAT Genetics and Cytogenetics in Oncology and Haematology

    (Patent information from GeneIP,
    Licensable technologies from WIS Yeda, Salk, Tufts,
    IP news from LifeMap Sciences, Inc.)
    About This Section
    Patent Information for HHAT gene:
    Search GeneIP for patents involving HHAT

    GeneCards and IP:
    Japan Patent Office Licenses GeneCards     European Patent Office Licenses GeneCards     Improving the IP Search



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    GeneCards Homepage - Last full update: 19 Mar 2013 - Incrementals: 21 Mar 2013 , 15 Apr 2013 , 26 Apr 2013

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