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EVI5L Gene

protein-coding   GIFtS: 47
GCID: GC19P007895

Ecotropic Viral Integration Site 5-Like

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(According to 1HGNC, 2Entrez Gene,
3UniProtKB/Swiss-Prot, 4UniProtKB/TrEMBL, 5OMIM, 6GeneLoc, 7Ensembl, 8DME, 9miRBase, 10fRNAdb, 12H-InvDB, 13NCBI, 14NONCODE, and/or 15RNAdb)
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Aliases
Ecotropic Viral Integration Site 5-Like1 2
Ecotropic Viral Integration Site 5-Like Protein2 3
EVI5-Like Protein2

External Ids:    HGNC: 304641   Entrez Gene: 1157042   Ensembl: ENSG000001424597   UniProtKB: Q96CN43   

Export aliases for EVI5L gene to outside databases

Previous GC identifers: GC19P007817 GC19P007802 GC19P007569


(According to Entrez Gene, GeneCards, Tocris Bioscience, Wikipedia's Gene Wiki, PharmGKB,
UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL)
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GeneCards Summary for EVI5L Gene:
EVI5L (ecotropic viral integration site 5-like) is a protein-coding gene. GO annotations related to this gene include Rab GTPase activator activity. An important paralog of this gene is TBC1D12.

UniProtKB/Swiss-Prot: EVI5L_HUMAN, Q96CN4
Function: Functions as a GTPase-activating protein (GAP) with a broad specificity




(According to GeneLoc and/or HGNC, and/or
Entrez Gene (NCBI build 37),
and/or miRBase,
Genomic Views according to UCSC (hg19) and Ensembl (release 75), Regulatory elements and Epigenetics data according to QIAGEN, and/or SwitchGear Genomics)
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RefSeq DNA sequence at NCBI GenBank:
NC_000019.9  NT_011295.12  NC_018930.2  
Regulatory elements:
   Regulatory transcription factor binding sites in the EVI5L gene promoter:
         POU2F2 (Oct-2.1)   oct-B3   oct-B2   POU2F2   Gfi-1   POU2F2C   AREB6   POU2F1   POU2F1a   POU2F2B   
         Other transcription factors

Browse SwitchGear Promoter luciferase reporter plasmids
   Search Chromatin IP Primers for EVI5L

Epigenetics:
DNA Methylation CpG Assay Predesigned for Pyrosequencing in human, mouse, rat EVI5L


Genomic Location:
Genomic View: UCSC Golden Path with GeneCards custom track

Entrez Gene cytogenetic band: 19p13.2   Ensembl cytogenetic band:  19p13.2   HGNC cytogenetic band: 19p13

EVI5L Gene in genomic location: bands according to Ensembl, locations according to (and/or Entrez Gene and/or Ensembl if different)
EVI5L gene location

GeneLoc information about chromosome 19         GeneLoc Exon Structure

GeneLoc location for GC19P007895:  view genomic region     (about GC identifiers)

Start:
7,895,119 bp from pter      End:
7,929,862 bp from pter
Size:
34,744 bases      Orientation:
plus strand

(According to 1UniProtKB, HORDE, 2neXtProt, Ensembl, and/or Reactome, Modification sites according to PhosphoSitePlus, Specific Peptides from DME, RefSeq according to NCBI, PDB rendering according to OCA and/or Proteopedia, Recombinant Proteins from EMD Millipore, R&D Systems, GenScript, Enzo Life Sciences, OriGene, Novus Biologicals, Sino Biological, ProSpec, Cloud-Clone Corp., eBioscience, and/or antibodies-online,
Biochemical Assays by EMD Millipore, R&D Systems, OriGene, GenScript, Cell Signaling Technology, Enzo Life Sciences, Cloud-Clone Corp., eBioscience, and/or antibodies-online, Antibodies by EMD Millipore, R&D Systems, Cell Signaling Technology, OriGene, Novus Biologicals, Thermo Fisher Scientific, Abcam, Cloud-Clone Corp, antibodies-online, and/or others.)
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UniProtKB/Swiss-Prot: EVI5L_HUMAN, Q96CN4 (See protein sequence)
Recommended Name: EVI5-like protein  
Size: 794 amino acids; 91376 Da
Subunit: May interact with RAB10
Secondary accessions: B9A6I9
Alternative splicing: 2 isoforms:  Q96CN4-1   Q96CN4-2   (No experimental confirmation available)

Explore the universe of human proteins at neXtProt for EVI5L: NX_Q96CN4

Explore proteomics data for EVI5L at MOPED

Post-translational modifications: 

  • Modification sites at neXtProt
  • Modification sites at PhosphoSitePlus

  • See EVI5L Protein Expression from SPIRE MOPED, PaxDB, and MaxQB

    REFSEQ proteins (2 alternative transcripts): 
    NP_001153416.1  NP_660288.1  

    ENSEMBL proteins: 
     ENSP00000270530   ENSP00000445905   ENSP00000472181   ENSP00000470145  

    EVI5L Human Recombinant Protein Products:

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    Cloud-Clone Corp. Proteins for EVI5L

     
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    antibodies-online proteins for EVI5L (3 products) 

     
    antibodies-online peptides for EVI5L

    EVI5L Antibody Products:

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    antibodies-online antibodies for EVI5L (12 products) 

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    (According to HGNC, IUPHAR, InterPro, ProtoNet, UniProtKB, and/or BLOCKS, Sets of similar genes according to GeneDecks)
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    1 InterPro protein domain:
     IPR000195 Rab-GTPase-TBC_dom

    Graphical View of Domain Structure for InterPro Entry Q96CN4

    ProtoNet protein and cluster: Q96CN4

    2 Blocks protein domains:
    IPB000195 RabGAP/TBC domain
    IPB000515 Binding-protein-dependent transport systems inner membrane component


    UniProtKB/Swiss-Prot: EVI5L_HUMAN, Q96CN4
    Similarity: Contains 1 Rab-GAP TBC domain


    Find genes that share domains with EVI5L           About GenesLikeMe


    (According to 1UniProtKB, Genatlas, LifeMap Discovery™, IUBMB, and/or 2DME, Human phenotypes from GenomeRNAi, Animal models from MGI Mar 06 2013, genOway,
    transcription factor targeting from QIAGEN and/or HOMER, miRNA Gene Targets from miRTarBase, shRNA from OriGene, siRNAs from OriGene, QIAGEN, microRNA from QIAGEN, SwitchGear Genomics, Gene Editing from DNA2.0, Clones from OriGene, GenScript, Sino Biological, DNA2.0, Vector BioLabs and/or Addgene, Cell Lines from GenScript, ESI BIO, In Situ Hybridization Assays from Advanced Cell Diagnostics, Flow cytometry from eBioscience, Ontologies according to Gene Ontology Consortium 01 Apr 2014 via Entrez Gene.)
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    Molecular Function:

         UniProtKB/Swiss-Prot Summary: EVI5L_HUMAN, Q96CN4
    Function: Functions as a GTPase-activating protein (GAP) with a broad specificity

         Gene Ontology (GO): 2 molecular function terms:    About this table

    GO IDQualified GO termEvidencePubMed IDs
    GO:0005097Rab GTPase activator activity IEA--
    GO:0005515protein binding IPI16189514
         
    Find genes that share ontologies with EVI5L           About GenesLikeMe


    Animal Models:
       genOway: Develop your customized and physiologically relevant rodent model for EVI5L

    miRNA
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    Selected qRT-PCR Assays for microRNAs that regulate EVI5L (see all 28):
    hsa-miR-30c hsa-miR-34b* hsa-miR-522 hsa-miR-1271 hsa-miR-200a hsa-miR-30d hsa-miR-449a hsa-miR-1205
    SwitchGear 3'UTR luciferase reporter plasmidEVI5L 3' UTR sequence
    Inhib. RNA
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    (According to UniProtKB, COMPARTMENTS Subcellular localization database, Ontologies according to Gene Ontology Consortium 01 Apr 2014 via Entrez Gene.)
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    Subcellular locations from COMPARTMENTS: 

    CompartmentConfidence
    cytosol2
    nucleus2

    (SuperPaths according to PathCards, Pathways according to R&D Systems, Cell Signaling Technology, KEGG, PharmGKB, BioSystems, Sino Biological, Reactome, Tocris Bioscience, GeneGo (Thomson Reuters), QIAGEN, and/or UniProtKB, Sets of similar genes according to GeneDecks, Interaction Networks according to QIAGEN, and/or STRING, Interactions according to 1UniProtKB, 2MINT, 3I2D, and/or 4STRING, with links to IntAct and Ensembl, Ontologies according to Gene Ontology Consortium 01 Apr 2014 via Entrez Gene).
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        Custom Pathway & Disease-focused RT2 Profiler PCR Arrays for EVI5L
    Interactions:

        Search GeneGlobe Interaction Network for EVI5L

    STRING Interaction Network Preview (showing 5 interactants - click image to see more details)

    Selected Interacting proteins for EVI5L (Q96CN41, 3 ENSP000002705304) via UniProtKB, MINT, STRING, and/or I2D (see all 26)
    InteractantInteraction Details
    GeneCardExternal ID(s)
    NUDT18Q6ZVK81, 3, ENSP000003078524EBI-749523,EBI-740486 I2D: score=3 STRING: ENSP00000307852
    ENSG00000258947Q135093I2D: score=1 
    TUBB3Q135093I2D: score=1 
    SH3YL1Q96HL83I2D: score=6 
    RAB8BQ929303I2D: score=4 
    About this table

    Gene Ontology (GO): 1 biological process term:    About this table

    GO IDQualified GO termEvidencePubMed IDs
    GO:0032313regulation of Rab GTPase activity ----

    Find genes that share ontologies with EVI5L           About GenesLikeMe



    (Chemical Compounds according to UniProtKB, Enzo Life Sciences, EMD Millipore, Tocris Bioscience, ApexBio, HMDB, BitterDB, and/or Novoseek, Ligands according to IUPHAR, and Drugs according to DrugBank, Enzo Life Sciences, and/or PharmGKB)
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    Browse Small Molecules at EMD Millipore
       Browse drugs & compounds from Enzo Life Sciences
      Browse compounds at ApexBio 

    Browse Tocris compounds for EVI5L



    (Secondary structures according to fRNAdb,
    GenBank/EMBL/DDBJ Accessions according to
    Unigene (Build 236 Homo sapiens; Apr 25 2013) or GenBank,
    RefSeq according to Entrez Gene,
    DOTS (version 10), and/or AceView, transcript ids from Ensembl with links to UCSC,
    exon structure from GeneLoc, alternative splicing isoforms according to ASD and/or ECgene,
    siRNAs from OriGene, QIAGEN, shRNA from OriGene, microRNA from QIAGEN, SwitchGear Genomics,
    Tagged/untagged cDNA clones from OriGene, GenScript, DNA2.0, Vector BioLabs, and/or Addgene, Primers from OriGene, and/or QIAGEN, Flow cytometry from eBioscience )
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    REFSEQ mRNAs for EVI5L gene (2 alternative transcripts): 
    NM_001159944.2  NM_145245.4  

    Unigene Cluster for EVI5L:

    Ecotropic viral integration site 5-like
    Hs.26870  [show with all ESTs]
    Unigene Representative Sequence: NM_145245
    7 Ensembl transcripts including schematic representations, and UCSC links where relevant:
    ENST00000270530(uc002min.3) ENST00000538904(uc010xjz.2 uc002mio.1)
    ENST00000597440 ENST00000599036 ENST00000601984 ENST00000600802 ENST00000601766

    miRNA
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    Selected qRT-PCR Assays for microRNAs that regulate EVI5L (see all 28):
    hsa-miR-30c hsa-miR-34b* hsa-miR-522 hsa-miR-1271 hsa-miR-200a hsa-miR-30d hsa-miR-449a hsa-miR-1205
    SwitchGear 3'UTR luciferase reporter plasmidEVI5L 3' UTR sequence
    Inhib. RNA
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    Pre-validated RT2 qPCR Primer Assay in human, mouse, rat EVI5L
      QuantiTect SYBR Green Assays in human, mouse, rat EVI5L
      QuantiFast Probe-based Assays in human, mouse, rat EVI5L

    Additional mRNA sequence: 

    AB449874.1 AK091609.1 BC014111.2 HM370400.1 

    8 DOTS entries:

    DT.445809  DT.95269450  DT.100036532  DT.95088313  DT.100670115  DT.100750195  DT.91745017  DT.97831238 

    Selected AceView cDNA sequences (see all 135):

    BM722689 AW572975 BM677555 BQ012107 CR613564 AI139560 BQ778293 T32165 
    BX412718 BQ268859 BM703366 Z38319 AI359135 AJ708251 BQ777177 BF195724 
    BG150278 BQ189190 BX404784 AI382542 AI283121 CD251248 BQ012122 BQ184638 

    GeneLoc Exon Structure

    3 Alternative Splicing Database (ASD) splice patterns (SP) for EVI5L    About this scheme

    ExUns: 1 ^ 2 ^ 3 ^ 4 ^ 5 ^ 6a · 6b ^ 7a · 7b ^ 8 ^ 9 ^ 10 ^ 11a · 11b · 11c ^ 12 ^ 13 ^ 14a · 14b ^ 15 ^ 16 ^ 17 ^ 18 ^ 19
    SP1:                                                                          -     -     -                                                         
    SP2:                                                                          -           -                                                         
    SP3:                                                                                                                                                


    ECgene alternative splicing isoforms for EVI5L

    (RNA expression data according to H-InvDB, NONCODE, miRBase, and RNAdb, Expression images according to data from BioGPS, Illumina Human BodyMap, and CGAP SAGE, Sets of similar genes according to GeneDecks, in vivo and in vitro expression data from LifeMap Discovery™, Protein expression images according to data from SPIRE 1MOPED, 2PaxDb, and 3MaxQB, plus additional links to SOURCE, and/or BioGPS, and/or UniProtKB,
    PCR Arrays from QIAGEN, Primers from OriGene, and/or QIAGEN, In Situ Hybridization Assays from Advanced Cell Diagnostics)
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    EVI5L expression in normal human tissues (normalized intensities)
    See probesets specificity/sensitivity at GeneAnnot
    About this imageBioGPS <intensity>2/3
    CGAP TAG: TCGCCCAGGC
    EVI5L Expression
    About this image

    EVI5L Protein expression data from MOPED1, PaxDb2 and MaxQB3    About this image

    EVI5L Protein Expression

    SOURCE GeneReport for Unigene cluster: Hs.26870
        Custom PCR Arrays for EVI5L
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    In Situ
    Assay Products:
     

     
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    (Orthologs according to 1,2HomoloGene (2older version, for species not in 1newer version), 3euGenes, 4SGD , 5MGI Mar 06 2013, with possible further links to Flybase and/or WormBase, and/or 6Ensembl pan taxonomic compara , Gene Trees according to Ensembl and TreeFam)
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    This gene was present in the common ancestor of animals and fungi.

    Orthologs for EVI5L gene from Selected species (see all 14)    About this table
    Organism Taxonomic
    classification
    Gene Description Human
    Similarity
    Orthology
    Type
    Details
    mouse
    (Mus musculus)
    Mammalia Evi5l5 ecotropic viral integration site 5 like   --   8 (1.95 cM) 4166567 
    chicken
    (Gallus gallus)
    Aves EVI5L6
    ecotropic viral integration site 5-like
    82(a)
    1 ↔ 1
    AADN03013353.1(1987-11350)
    lizard
    (Anolis carolinensis)
    Reptilia EVI5L6
    ecotropic viral integration site 5-like
    91(a)
    1 ↔ 1
    GL343286.1(305-26785)
    tropical clawed frog
    (Xenopus tropicalis)
    Amphibia evi5l1 ecotropic viral integration site 5-like 69.81(n)
    74.44(a)
      100145757  NM_001127128.1  NP_001120600.1 
    zebrafish
    (Danio rerio)
    Actinopterygii si:ch211-239f4.11 si:ch211-239f4.1 70.24(n)
    76(a)
      566318  XM_689590.6  XP_694682.6 
    fruit fly
    (Drosophila melanogaster)
    Insecta CG117276
    --
    42(a)
    1 → many
    X(11324556-11348371)
    worm
    (Caenorhabditis elegans)
    Secernentea eat-176
    Protein F01G12.6 (F01G12.6) mRNA, complete cds
    35(a)
    1 → many
    X(16399210-16416840) WBGene00020770
    baker's yeast
    (Saccharomyces cerevisiae)
    Saccharomycetes GYP56
    GYL16
    GTPase-activating protein (GAP) for yeast Rab fami...
    Putative GTPase activating protein (GAP) with a ro...
    17(a)
    16(a)
    many ↔ many
    many ↔ many
    XVI(76669-79353) YPL249C
    XIII(647118-649280) YMR192W


    ENSEMBL Gene Tree for EVI5L (if available)
    TreeFam Gene Tree for EVI5L (if available)

    (Paralogs according to 1HomoloGene,
    2Ensembl, and 3SIMAP, Pseudogenes according to 4Pseudogene.org Build 68)
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    Paralogs for EVI5L gene
    TBC1D122  TBC1D42  TBC1D142  RABGAP1L2  TBC1D12  EVI52  RABGAP12  
    1 SIMAP similar gene for EVI5L using alignment to 3 protein entries:     EVI5L_HUMAN (see all proteins):
    EVI5

    Find genes that share paralogs with EVI5L           About GenesLikeMe


    1 Pseudogenes.org Pseudogene for EVI5L
    PGOHUM00000263027


    (SNPs/Variants according to the 1NCBI SNP Database, 2Ensembl, 3PupaSUITE, 4UniProtKB, and DNA2.0, Linkage Disequilibrium by HapMap, Structural Variations(CNVs/InDels/Inversions) from the Database of Genomic Variants, Mutations from the Human Gene Mutation Database (HGMD), the Human Cytochrome P450 Allele Nomenclature Database, and the Locus Specific Mutation Databases (LSDB), Blood group antigen gene mutations by BGMUT, Resequencing Primers, Cancer Mutation PCR Arrays and Assays, and Copy Number PCR Arrays from QIAGEN)
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    Selected SNPs for EVI5L (see all 810)    About this table                                 

    Genomic DataTranscription Related DataAllele Frequencies
    SNP IDValidClinical
    significance
    Chr 19 posSequence#AA
    Chg
    TypeMore#Allele
    freq
    PopTotal
    sample
    More
    ----------
    rs1824734561,2
    --7893183(+) AGGCAC/TCCGCC 1 -- us2k10--------
    rs1879742141,2
    --7893185(+) GCACCC/TGCCTC 1 -- us2k10--------
    rs1145915611,2
    F--7893497(+) TTGAGC/ATCTCA 1 -- us2k11Minor allele frequency- A:0.01WA 118
    rs765303631,2
    C--7893528(+) TGCCCC/GTCCCT 1 -- us2k10--------
    rs1923123701,2
    --7893581(+) GCTGGA/GCGAGG 1 -- us2k10--------
    rs1842169791,2
    --7893599(+) TGCCTG/TTAATC 1 -- us2k10--------
    rs1434453601,2
    --7893697(+) CAAAAA/TAATTA 1 -- us2k10--------
    rs1467652951,2
    --7893768(+) TCATTC/GGAGTC 1 -- us2k10--------
    rs1404964591,2
    --7893778(+) CCAAGA/GGTTCG 1 -- us2k10--------
    rs1888152341,2
    --7893888(+) GTCACC/TCTCTC 1 -- us2k10--------

    HapMap Linkage Disequilibrium report for EVI5L (7895119 - 7929862 bp)

    Structural Variations
         Database of Genomic Variants (DGV) 8 variations for EVI5L:    About this table    
    Variant IDTypeSubtypePubMed ID
    nsv910986CNV Loss21882294
    nsv910984CNV Loss21882294
    nsv525499CNV Loss19592680
    dgv3715n71CNV Loss21882294
    nsv910985CNV Loss21882294
    nsv910980CNV Loss21882294
    nsv833733CNV Gain17160897
    nsv833732CNV Gain17160897

    Site Specific Mutation Identification with PCR Assays
    SeqTarget long-range PCR primers for resequencing EVI5L
    DNA2.0 Custom Variant and Variant Library Synthesis for EVI5L

    (in which this Gene is Involved, According to MalaCards, OMIM, UniProtKB, the University of Copenhagen DISEASES database, Genatlas, GeneTests, GAD, HuGE Navigator, and/or TGDB.)
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    Find genes that share disorders with EVI5L           About GenesLikeMe

    Genetic Association Database (GAD): EVI5L

    Export disorders for EVI5L gene to outside databases

    (in PubMed. Associations of this gene to articles via 1Entrez Gene, 2UniProtKB/Swiss-Prot, 3HGNC, 4GAD, 5PharmGKB, 6HMDB, 7DrugBank, 8UniProtKB/TrEMBL, 9 Novoseek, and/or 10fRNAdb)
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    PubMed articles for EVI5L gene, integrated from 10 sources (see all 12):
    (articles sorted by number of sources associating them with EVI5L)
        Utopia: connect your pdf to the dynamic
    world of online information

    1. The Evi5 family in cellular physiology and pathology. (PubMed id 23669355)1, 3 Lim Y.S. and Tang B.L. (FEBS Lett. 2013)
    2. Host determinants of HIV-1 control in African Americans. (PubMed id 20205591)1, 4 Pelak K.... . (J. Infect. Dis. 2010)
    3. Identification and characterization of a novel Tre-2/Bub2/Cdc16 (TBC) protein that possesses Rab3A-GAP activity. (PubMed id 19077034)1, 2 Ishibashi K.... Fukuda M. (Genes Cells 2009)
    4. Screening for target Rabs of TBC (Tre-2/Bub2/Cdc16) domain-containing proteins based on their Rab-binding activity. (PubMed id 16923123)1, 2 Itoh T.... Fukuda M. (Genes Cells 2006)
    5. The status, quality, and expansion of the NIH full-length cDNA project: the Mammalian Gene Collection (MGC). (PubMed id 15489334)1, 2 Gerhard D.S.... Malek J. (Genome Res. 2004)
    6. The protein interaction landscape of the human CMGC kinase group. (PubMed id 23602568)1 Varjosalo M....Gstaiger M. (Cell Rep 2013)
    7. Interactome mapping suggests new mechanistic details underlying Alzheimer's disease. (PubMed id 21163940)1 Soler-LA^pez M....Aloy P. (Genome Res. 2011)
    8. An empirical framework for binary interactome mapping. (PubMed id 19060904)1 Venkatesan K....Vidal M. (Nat. Methods 2009)
    9. Towards a proteome-scale map of the human protein-protein interaction network. (PubMed id 16189514)1 Rual J.F....Vidal M. (Nature 2005)
    10. Complete sequencing and characterization of 21,243 full-length human cDNAs. (PubMed id 14702039)1 Ota T.... Sugano S. (Nat. Genet. 2004)

    (in PubMed, OMIM, and NCBI Bookshelf)
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      Query String
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    (According to Entrez Gene, HGNC, AceView, euGenes, Ensembl, miRBase, ECgene, Kegg, and/or H-InvDB)
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    Entrez Gene: 115704 HGNC: 30464 AceView: EVI5L Ensembl:ENSG00000142459 euGenes: HUgn115704
    ECgene: EVI5L H-InvDB: EVI5L

    (According to HUGE)
    About This Section

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    (According to PharmGKB, ATLAS, HORDE, IMGT, LEIDEN, UniProtKB/Swiss-Prot, UniProtKB/TrEMBL, and/or others, e.g. Wikipedia and GeneReviews, via UniProtKB/Swiss-Prot)
    About This Section

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    NameDescription
    PharmGKB entry for EVI5L Pharmacogenomics, SNPs, Pathways

    (Patent information from GeneIP,
    Licensable technologies from WIS Yeda, Salk, Tufts,
    IP news from LifeMap Sciences, Inc.)
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    Patent Information for EVI5L gene:
    Search GeneIP for patents involving EVI5L

    GeneCards and IP:
    Japan Patent Office Licenses GeneCards     European Patent Office Licenses GeneCards     Improving the IP Search



    (Antibodies, recombinant proteins, and assays from EMD Millipore, R&D Systems, OriGene, QIAGEN, GenScript, Cell Signaling Technology, Novus Biologicals, Sino Biological, Enzo Life Sciences, Abcam, ProSpec, Cloud-Clone Corp., Thermo Fisher Scientific, eBioscience, antibodies-online, and/or others, Gene Editing from DNA2.0. Clones from OriGene, GenScript, Sino Biological, DNA2.0, SwitchGear Genomics, Vector BioLabs, Addgene, Cell lines from GenScript, and ESI BIO, Flow cytometery from eBioscience, PCR Arrays from QIAGEN, Drugs and/or compounds from EMD Millipore, Tocris Bioscience, Enzo Life Sciences, and/or ApexBio, In Situ Hybridization Assays from
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