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Category Symbol Source: HGNC EntrezGene Ensembl GeneCards RNA genes CroW21

GIFtS
-

DNAJC18 Gene

protein-coding GIFtS: 45
GCID: GC05M138776

DnaJ (Hsp40) homolog, subfamily C, member 18

Explore 3 diseases affiliated with
DNAJC18 via

MalaCards

our new
Human Malady Compendium

(According to ¹HGNC, ²Entrez Gene,
³UniProtKB/Swiss-Prot, ⁴UniProtKB/TrEMBL, ⁵OMIM, ⁶GeneLoc, ⁷Ensembl, ⁸DME, ⁹miRBase, and/or ¹⁰fRNAdb)
About This Section

Aliases
DnaJ (Hsp40) Homolog, Subfamily C, Member 18¹ ²
MGC29463¹
DnaJ Homolog Subfamily C Member 18²

External Ids:	HGNC: 28429¹	Entrez Gene: 202052²	Ensembl: ENSG00000170464⁷	UniProtKB: Q9H819³

Export aliases for DNAJC18 gene to outside databases

Previous GC identifer: GC05M133935

(According to Entrez Gene, Tocris Bioscience, Wikipedia's Gene Wiki, PharmGKB,
UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL)
About This Section

(According to GeneLoc and/or HGNC, and/or
Entrez Gene (NCBI build 37),
and/or miRBase,
Genomic Views according to UCSC (hg19) and Ensembl (release 69), Regulatory elements and Epigenetics data according to QIAGEN, SABiosciences, and/or SwitchGear Genomics)
About This Section

RefSeq DNA sequence:

NC_000005.9 NC_018916.1 NT_034772.6

Regulatory elements:

SABiosciences Regulatory transcription factor binding sites in the DNAJC18 gene promoter:
HFH-3 AhR Lmo2 Olf-1 HNF-4alpha1 Arnt FOXI1 c-Myb Msx-1
Other transcription factors

SwitchGear Promoter luciferase reporter plasmid: DNAJC18 promoter sequence

Search SABiosciences Chromatin IP Primers for DNAJC18

Epigenetics:

QIAGEN PyroMark CpG Assay predesigned Pyrosequencing DNA Methylation assays in human, mouse, rat DNAJC18

Genomic Location:
Genomic View: UCSC Golden Path with GeneCards custom track

Entrez Gene cytogenetic band: 5q31.2 Ensembl cytogenetic band: 5q31.2 HGNC cytogenetic band: 5q31.2

DNAJC18 Gene in genomic location: bands according to Ensembl, locations according to (and/or Entrez Gene and/or Ensembl if different)
DNAJC18 gene location

GeneLoc information about chromosome 5 GeneLoc Exon Structure

GeneLoc location for GC05M138776: view genomic region (about GC identifiers)

Start:	138,744,279 bp from pter	End:	138,780,180 bp from pter
Size:	35,902 bases	Orientation:	minus strand

(According to ¹UniProtKB, HORDE, neXtProt, Ensembl, and/or Reactome, Modification sites according to ²PhosphoSitePlus, Specific Peptides from DME, Protein expression images according to data from SPIRE MOPED and PaxDb, RefSeq according to NCBI, PDB rendering according to OCA and/or Proteopedia, Recombinant Proteins from EMD Millipore, R&D Systems, GenScript, Enzo Life Sciences, OriGene, Novus Biologicals, Sino Biological, ProSpec, and/or Uscn,
Biochemical Assays by EMD Millipore, R&D Systems, OriGene, GenScript, Cell Signaling Technology, Enzo Life Sciences, and/or Uscn, Ontologies according to Gene Ontology Consortium 01 Mar 2013 and Entrez Gene, Antibodies by EMD Millipore, R&D Systems, GenScript, Cell Signaling Technology, OriGene, Novus Biologicals, Thermo Fisher Scientific, Abcam, and/or Uscn)
About This Section

UniProtKB/Swiss-Prot: DJC18_HUMAN, Q9H819 (See protein sequence)

Recommended Name: DnaJ homolog subfamily C member 18

Size: 358 amino acids; 41551 Da

Subcellular location: Membrane; Single-pass membrane protein (Potential)

Explore the universe of human proteins at neXtProt for DNAJC18: NX_Q9H819

Post-translational modifications:

View modification sites using PhosphoSitePlus²

View neXtProt modification sites for NX_Q9H819

DNAJC18 Protein expression data from MOPED and PaxDb: About this image

REFSEQ proteins: NP_689899.1

ENSEMBL proteins:

ENSP00000302843 ENSP00000426338 ENSP00000425192 ENSP00000424438 ENSP00000427700

ENSP00000424572 ENSP00000425523 ENSP00000426819

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	Uscn Proteins for DNAJC18

Gene Ontology (GO): 1 cellular component term (GO ID links to tree view): About this table

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0016021	integral to membrane	IEA	--

DNAJC18 for ontologies About GeneDecksing

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(According to InterPro, ProtoNet, UniProtKB, and/or BLOCKS, Sets of similar genes according to GeneDecks)
About This Section

DNAJC18 for domains About GeneDecksing

3 InterPro domains/families:

IPR015399 DUF1977_DnaJ-like

IPR001623 DnaJ_domain

IPR018253 DnaJ_domain_CS

Graphical View of Domain Structure for InterPro Entry Q9H819

ProtoNet protein and cluster: Q9H819

1 Blocks protein family: IPB002052 N-6 Adenine-specific DNA methylase

UniProtKB/Swiss-Prot: DJC18_HUMAN, Q9H819

Similarity: Contains 1 J domain

(According to ¹UniProtKB, Genatlas, LifeMap Discovery™, IUBMB, and/or ²DME, Human phenotypes from GenomeRNAi, Animal models from MGI Mar 06 2013,
bound targets from SABiosciences, miRNA Gene Targets from miRTarBase shRNA from OriGene, RNAi from EMD Millipore, siRNAs from OriGene, QIAGEN, microRNA from QIAGEN, Gene Editing from DNA2.0, Clones from EMD Millipore, OriGene, SwitchGear Genomics, GenScript, Sino Biological, DNA2.0, and Vector BioLabs, Cell Lines from GenScript, LifeMap BioReagents, In Situ Hybridization Assays from Advanced Cell Diagnostics, Ontologies according to Gene Ontology Consortium 01 Mar 2013 via Entrez Gene.)
About This Section

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In Situ Assay
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Gene Ontology (GO): 2 molecular function terms (GO ID links to tree view): About this table

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0031072	heat shock protein binding	IEA	--
GO:0051082	unfolded protein binding	IEA	--

DNAJC18 for ontologies About GeneDecksing

(Pathways according to EMD Millipore, R&D Systems, Cell Signaling Technology, KEGG, PharmGKB, BioSystems, Reactome, Tocris Bioscience, GeneGo (Thomson Reuters), QIAGEN, and/or UniProtKB, Sets of similar genes according to GeneDecks, Interaction Networks according to SABiosciences, and/or STRING, Interactions according to ¹UniProtKB, ²MINT, ³I2D, and/or ⁴STRING, with links to IntAct and Ensembl, Ontologies according to Gene Ontology Consortium 01 Mar 2013 via Entrez Gene).
About This Section

Interactions:

Search SABiosciences Gene Network Central^TM Interacting Genes and Proteins Networks for DNAJC18

Gene Ontology (GO): 1 biological process term (GO ID links to tree view): About this table

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0006457	protein folding	IEA	--

DNAJC18 for ontologies About GeneDecksing

(Chemical Compounds according to UniProtKB, Enzo Life Sciences, EMD Millipore, Tocris Bioscience HMDB, BitterDB, and/or Novoseek, and Drugs according to DrugBank, Enzo Life Sciences, and/or PharmGKB, with drugs/clinical trials/news search links to CenterWatch)
About This Section

Browse Small Molecules at EMD Millipore

Browse drugs & compounds from Enzo Life Sciences

Browse Tocris compounds for DNAJC18
Search CenterWatch for drugs/clinical trials and news about DNAJC18 / DJC18

(Secondary structures according to fRNAdb,
GenBank/EMBL/DDBJ Accessions according to
Unigene (Build 235 Homo sapiens; Mar 10 2013) or GenBank,
RefSeq according to Entrez Gene,
DOTS (version 10), and/or AceView, transcript ids from Ensembl with links to UCSC,
exon structure from GeneLoc, alternative splicing isoforms according to ASD and/or ECgene,
RNAi Products from EMD Millipore,
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Tagged/untagged cDNA clones from OriGene, SwitchGear Genomics, GenScript, DNA2.0, Vector BioLabs, Primers from OriGene, SABiosciences, and/or QIAGEN )
About This Section

REFSEQ mRNAs for DNAJC18 gene:

NM_152686.3

Unigene Cluster for DNAJC18:

DnaJ (Hsp40) homolog, subfamily C, member 18

Hs.483537 [show with all ESTs]

Unigene Representative Sequence: NM_152686

12 Ensembl transcripts including schematic representations, and UCSC links where relevant:

ENST00000515559 ENST00000302060(uc003len.3 uc010jff.3) ENST00000510770

ENST00000508445 ENST00000514052 ENST00000507405 ENST00000505568 ENST00000506870

ENST00000515581 ENST00000515277 ENST00000512473(uc011czd.2) ENST00000505268

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		hsa-miR-576-3p hsa-miR-3164 hsa-miR-328 hsa-miR-429 hsa-miR-1197 hsa-miR-134 hsa-miR-3916 hsa-miR-649

		SwitchGear 3'UTR luciferase reporter plasmid: DNAJC18 3' UTR sequence

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Additional cDNA sequence:

AK001329.1 AK304726.1 AK308138.1 BC030162.2

10 DOTS entries:

DT.110337 DT.120805043 DT.423799 DT.100813482 DT.95188934 DT.100813481 DT.120805033 DT.95374681

DT.95374680 DT.40229279

24/117 AceView cDNA sequences (see all 117):

BM993756 BE382440 AA621729 AI301373 AI125822 F03940 BQ425372 BF032590

AI239431 BI826438 NM_152686 AA807384 CB241698 BM696532 AI566804 F04462

F10478 BE935916 AW162126 AI193417 BC030162 AI884305 BM554404 BM824961

GeneLoc Exon Structure

4 Alternative Splicing Database (ASD) splice patterns (SP) for DNAJC18 About this scheme

ExUns:	1	^	2a	·	2b	·	2c	^	3	^	4a	·	4b	^	5	^	6a	·	6b	^	7	^	8	^	9
SP1:
SP2:
SP3:									-
SP4:											-		-

ECgene alternative splicing isoforms for DNAJC18

(RNA expression data according to H-InvDB, NONCODE, miRBase, and RNAdb, Expression images according to data from BioGPS, Illumina Human BodyMap, and CGAP SAGE, Sets of similar genes according to GeneDecks, in vivo and in vitro expression data from LifeMap Discovery™, plus additional links to Genevestigator, and/or SOURCE, and/or BioGPS, and/or UniProtKB,
PCR Arrays from SABiosciences, Primers from OriGene, SABiosciences, and/or QIAGEN, In Situ Hybridization Assays from Advanced Cell Diagnostics)
About This Section

DNAJC18 expression in normal human tissues (normalized intensities)
See probesets specificity/sensitivity at GeneAnnot
About this imageBioGPS
CGAP TAG: GCAGCGATGA

About this image

DNAJC18 expression in embryonic tissues and stem cells

Expression by the Database of Embryonic development, Stem cell research, and Regenerative medicine About this table

5 LifeMap In Vivo Development Anatomical Compartments/Cells

Tissue

Anatomical Compartment

Cell

Category (developmental path)

Bone

Endochondral Facial Bones

Bone

Bone

Bone

Bone

Cartilage

Expression:

Positive

Negative

Selective marker

Experimental details:

Curated

Microarrays

In-situ hybridization

See DNAJC18 Protein Expression from SPIRE MOPED and PaxDB
Genevestigator expression for DNAJC18

SOURCE GeneReport for Unigene cluster: Hs.483537

SABiosciences Expression via Pathway-Focused PCR Array including DNAJC18:

Heat Shock Proteins & Chaperones in human mouse rat

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In Situ
Assay Products:

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(Orthologs according to ^1,2HomoloGene (²older version, for species not in ¹newer version), ³euGenes, ⁴SGD , ⁵MGI Mar 06 2013, with possible further links to Flybase and/or WormBase, and/or ⁶Ensembl pan taxonomic compara , Gene Trees according to Ensembl and TreeFam)
About This Section

This gene was present in the common ancestor of eukaryotes.

Orthologs for DNAJC18 gene from 8/27 species (see all 27) About this table

Organism	Taxonomic classification	Gene	Description	Human Similarity	Orthology Type	Details
mouse (Mus musculus)	Mammalia	Dnajc18^{1 , 5}	DnaJ (Hsp40) homolog, subfamily C, member 18^{1, 5}	87.49(n)¹ 90.2(a)¹		18 (19.21 cM)⁵ 76594¹ NM_029669.4¹ NP_083945.1¹ 35671103⁵
chicken (Gallus gallus)	Aves	DNAJC18¹	DnaJ (Hsp40) homolog, subfamily C, member 18	72.34(n) 76.53(a)		416189 NM_001025609.1 NP_001020780.1
lizard (Anolis carolinensis)	Reptilia	DNAJC18⁶	--	68(a)	1 ↔ 1	1(132159121-132169439)
zebrafish (Danio rerio)	Actinopterygii	dnajc18¹	DnaJ (Hsp40) homolog, subfamily C, member 18	61.97(n) 59.53(a)		559928 NM_001113588.1 NP_001107060.1
fruit fly (Drosophila melanogaster)	Insecta	CG3061⁶ CG30156⁶	--	30(a) 13(a)	many ↔ many many ↔ many	3R(9858089-9860764) 2R(2868951-2870491)
worm (Caenorhabditis elegans)	Secernentea	dnj-1⁶	DNaJ domain (prokaryotic heat shock protein) famil...	29(a)	1 → many	IV(11298986-11304765)
thale cress (Arabidopsis thaliana)	eudicotyledons	AT3G57340⁶ AT5G05750⁶ (see all 3)	DnaJ/Hsp40 domain-containing protein DNAJ heat shock N-terminal domain-containing prote... (see all 3)	23(a) 21(a) (see all 3)	many ↔ many many ↔ many (see all 3)	3(21218973-21220485) 5(1727429-1728933)
rice (Oryza sativa)	Liliopsida	-- --	chaperone protein dnaJ 49, putative, expressed	24(a) 22(a)	many ↔ many many ↔ many	1(20986715-20991489) 5(17381202-17385570)

ENSEMBL Gene Tree for DNAJC18 (if available)
TreeFam Gene Tree for DNAJC18 (if available)

(Paralogs according to ¹HomoloGene,
²Ensembl, and ³SIMAP, Pseudogenes according to ⁴Pseudogene.org Build 68)
About This Section

Paralogs for DNAJC18 gene

DNAJB12² DNAJC16² DNAJB14²

14 SIMAP similar genes for DNAJC18 using alignment to 9 protein entries: DJC18_HUMAN (see all proteins):

LOC51277 DNAJC5B DNAJC1 DNAJB14 DNAJB5 DNAJB4

DNAJB6 DNAJA4 DNAJB12 DNAJC17 ZCSL3 DNAJC24

DNAJB2 DNAJB3

DNAJC18 for paralogs About GeneDecksing

(SNPs/Variants according to the ¹NCBI SNP Database, ²Ensembl, ³PupaSUITE, UniProtKB, and DNA2.0, Linkage Disequilibrium by HapMap, Structural Variations(CNVs/InDels/Inversions) from the Database of Genomic Variants, Mutations from the Human Gene Mutation Database (HGMD) and the Locus Specific Mutation Databases (LSDB), Blood group antigen gene mutations by BGMUT, Resequencing Primers from QIAGEN, Cancer Mutation PCR Arrays and Assays and Copy Number PCR Arrays from SABiosciences)
About This Section

SNP ID	Valid	Clinical significance	Chr 5 pos	Sequence	#	AA Chg	Type	More	#	Allele freq	Pop	Total sample	More
Genomic Data					Transcription Related Data				Allele Frequencies
				--	--	--		--	--
rs184259532^1,2		--	138745416(+)	`CACCTA/GTTCCC`	1	--	int¹		0	--	--	--	--
rs62385298^1,2	C,F,	--	138745616(+)	`AGGAAC/GTGGGA`	1	--	int¹		6		NA CSA WA EA	364
rs190419940^1,2		--	138745625(+)	`GATAAC/TACCTG`	1	--	int¹		0	--	--	--	--
rs144891021^1,2		--	138745659(+)	`TTGATC/TTGTAT`	1	--	int¹		0	--	--	--	--
rs4315920^1,2	C,F,A,H,	--	138745685(+)	`CTGCAG/ACTGCT`	1	--	int¹		36		NS EA NA MN WA CSA	3256
rs182006961^1,2		--	138745805(+)	`CTTTAG/TTCGGA`	1	--	int¹		0	--	--	--	--
rs186238720^1,2		--	138745812(+)	`CGGAAA/GTGGCT`	1	--	int¹		0	--	--	--	--
rs146745281^1,2		--	138746115(+)	`TTTCTC/TGGTGG`	1	--	ut3¹		0	--	--	--	--
rs140356741^1,2		--	138746211(+)	`CTAAGC/TTTATA`	1	--	ut3¹		0	--	--	--	--
rs139940065^1,2	C,	--	138746248(+)	`ATGAA-/GTGTGCA`	1	--	ut3¹		0	--	--	--	--

HapMap Linkage Disequilibrium report for DNAJC18 (138744279 - 138780180 bp)
Structural Variations (Copy Number Variations, Insertions/Deletions, Inversions)
Database of Genomic Variants (DGV) variations for DNAJC18: --

SABiosciences Cancer Mutation PCR Assays

QIAGEN SeqTarget long-range PCR primers in human, mouse, rat for resequencing DNAJC18

DNA2.0 Custom Variant and Variant Library Synthesis for DNAJC18

(in which this Gene is Involved, According to MalaCards, OMIM, UniProtKB, the University of Copenhagen DISEASES database, Genatlas, GeneTests, GAD, HuGE Navigator, and/or TGDB.)
About This Section

DNAJC18 for disorders About GeneDecksing

3 diseases for DNAJC18: About MalaCards

myelodysplastic syndrome pulmonary disease malaria

Human Genome Epidemiology (HuGE) Navigator: DNAJC18 (5 documents)

Export disorders for DNAJC18 gene to outside databases

(in PubMed. Associations of this gene to articles via ¹Entrez Gene, ²UniProtKB/Swiss-Prot, ³HGNC, ⁴GAD, ⁵PharmGKB, ⁶HMDB, ⁷DrugBank, ⁸UniProtKB/TrEMBL, ⁹ Novoseek, and/or ¹⁰fRNAdb)
About This Section

PubMed articles for DNAJC18 gene, integrated from 9 sources (see all 11):
(articles sorted by number of sources associating them with DNAJC18)

Utopia: connect your pdf to the dynamic
world of online information

The status, quality, and expansion of the NIH full-length cDNA project: the Mammalian Gene Collection (MGC). (PubMed id 15489334)^{1, 2} Gerhard D.S....Malek J. (2004)
Complete sequencing and characterization of 21,243 full-length human cDNAs. (PubMed id 14702039)^{1, 2} Ota T.... Sugano S. (2004)
Genetic susceptibility to distinct bladder cancer sub phenotypes. (PubMed id 19692168)¹ Guey L.T....Malats N. (2010)
Integrated genomic analysis implicates haploinsufficiency of multiple chromosome 5q31.2 genes in de novo myelodysplastic syndromes pathogenesis. (PubMed id 19240791)¹ Graubert T.A....Walter M.J. (2009)
Polymorphisms in innate immunity genes and lung cancer risk in Xuanwei, China. (PubMed id 19170196)¹ Shen M....Lan Q. (2009)
PTEN identified as important risk factor of chronic o bstructive pulmonary disease. (PubMed id 19625176)¹ Hosgood H.D....Lan Q. (2009)
Pathway-based evaluation of 380 candidate genes and lung cancer susceptibility suggests the importance of the cell cycle pathway. (PubMed id 18676680)¹ Hosgood H.D....Lan Q. (2008)
hORFeome v3.1: a resource of human open reading frames representing over 10,000 human genes. (PubMed id 17207965)¹ Lamesch P.... Vidal M. (2007)
Diversification of transcriptional modulation: large-scale identification and characterization of putative alternative promoters of human genes. (PubMed id 16344560)¹ Kimura K.... Sugano S. (2006)
Generation and initial analysis of more than 15,000 full-length human and mouse cDNA sequences. (PubMed id 12477932)¹ Strausberg R.L....Marra M.A. (2002)

(in PubMed, OMIM, and NCBI Bookshelf)
About This Section

		AND	OR
Aliases
Free Text

Query String		PubMed OMIM NCBI Bookshelf
	(Note: In FireFox, select the above section and copy using Ctrl-C)

(According to Entrez Gene, HGNC, AceView, euGenes, Ensembl, miRBase, ECgene, Kegg, and/or H-InvDB)
About This Section

Entrez Gene: 202052	HGNC: 28429	AceView: MGC29463	Ensembl:ENSG00000170464	euGenes: HUgn202052
ECgene: DNAJC18	H-InvDB: DNAJC18

(According to HUGE)
About This Section

(According to PharmGKB, ATLAS, HORDE, IMGT, LEIDEN, UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL,
Wikipedia and/or GeneReviews via UniProtKB/Swiss-Prot)
About This Section

Name	Description
PharmGKB entry for DNAJC18	Pharmacogenomics, SNPs, Pathways

Intellectual Property for DNAJC18 gene
(Patent information from GeneIP, Licensable technologies from WIS Yeda, Salk, Tufts, IP news from LifeMap Sciences, Inc.)

About This Section

Patent Information for DNAJC18 gene:
Search GeneIP for patents involving DNAJC18

GeneCards and IP:
Japan Patent Office Licenses GeneCards European Patent Office Licenses GeneCards Improving the IP Search

(Antibodies, recombinant proteins, and assays from EMD Millipore, R&D Systems, OriGene, QIAGEN, GenScript, Cell Signaling Technology, SABiosciences, Novus Biologicals, Sino Biological, Enzo Life Sciences, Abcam, ProSpec, Uscn, Thermo Fisher Scientific, Gene Editing from DNA2.0, Clones from EMD Millipore, OriGene, GenScript, Sino Biological, DNA2.0, SwitchGear Genomics, Vector BioLabs, Cell lines from GenScript and LifeMap BioReagents, PCR Arrays from SABiosciences, Drugs and/or compounds from EMD Millipore, Tocris Bioscience, and/or Enzo Life Sciences),
In Situ Hybridization Assays from
Advanced Cell Diagnostics
About This Section


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GeneCards Homepage - Last full update: 18 Mar 2013 - Incrementals: 21 Mar 2013 , 15 Apr 2013 , 26 Apr 2013

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Category			VWF (GIFTS: 73) von Willebrand factor
GIFtS Group

The GeneCards human gene database gene index: 1 3 5 6 A B C D E F G H I J K L M N O P Q R S T U V W X Y Z

The GeneCards Human Gene Database

Developed at the Crown Human Genome Center, Department of Molecular Genetics, the Weizmann Institute of Science

Hot genes Disease genes DNAJC18 gene at Home site.

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DNAJC18 Gene

DnaJ (Hsp40) homolog, subfamily C, member 18

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for DNAJC18 gene

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DNAJC18 Gene

DnaJ (Hsp40) homolog, subfamily C, member 18

Aliasesfor DNAJC18 gene

Summariesfor DNAJC18 gene

Genomic Viewsfor DNAJC18 gene

Proteinsfor DNAJC18 gene

Protein Domains /Familiesfor DNAJC18 gene

Functionfor DNAJC18 gene

Pathways & Interactionsfor DNAJC18 gene

Drugs & Compoundsfor DNAJC18 gene

Transcriptsfor DNAJC18 gene

Expression for DNAJC18 gene

Orthologsfor DNAJC18 gene

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Genomic Variantsfor DNAJC18 gene

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Genome Databases showing DNAJC18 gene

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Specialized Databases showing DNAJC18 gene

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for DNAJC18 gene

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for DNAJC18 gene

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for DNAJC18 gene

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Families
for DNAJC18 gene

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for DNAJC18 gene

Pathways & Interactions
for DNAJC18 gene

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for DNAJC18 gene

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for DNAJC18 gene

Expression
for DNAJC18 gene

Orthologs
for DNAJC18 gene

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for DNAJC18 gene

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for DNAJC18 gene

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for DNAJC18 gene

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for DNAJC18 gene

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for DNAJC18 gene

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for DNAJC18 gene