Aliases for DNAH14 Gene
Aliases for DNAH14 Gene
External Ids for DNAH14 Gene
- HGNC: 2945
- Entrez Gene: 127602
- Ensembl: ENSG00000185842
- OMIM: 603341
- UniProtKB: Q0VDD8
Previous HGNC Symbols for DNAH14 Gene
- C1orf67
Previous GeneCards Identifiers for DNAH14 Gene
- GC01U990036
- GC01P222303
- GC01P222302
- GC03M057407
- GC01P221576
- GC01P223398
- GC01P223285
- GC01P223184
- GC01P225083
- GC01P195635
Summaries for DNAH14 Gene
-
Dyneins are microtubule-associated motor protein complexes composed of several heavy, light, and intermediate chains. Two major classes of dyneins, axonemal and cytoplasmic, have been identified. DNAH14 is an axonemal dynein heavy chain (DHC) (Vaughan et al., 1996 [PubMed 8812413]).[supplied by OMIM, Mar 2008]
GeneCards Summary for DNAH14 Gene
DNAH14 (Dynein Axonemal Heavy Chain 14) is a Protein Coding gene. Diseases associated with DNAH14 include Communicating Hydrocephalus and Hydrocephalus, Normal-Pressure. Among its related pathways are Respiratory electron transport, ATP synthesis by chemiosmotic coupling, and heat production by uncoupling proteins.. Gene Ontology (GO) annotations related to this gene include microtubule motor activity. An important paralog of this gene is DNAH6.
UniProtKB/Swiss-Prot for DNAH14 Gene
-
Force generating protein of respiratory cilia. Produces force towards the minus ends of microtubules. Dynein has ATPase activity; the force-producing power stroke is thought to occur on release of ADP. Involved in sperm motility; implicated in sperm flagellar assembly (By similarity).
Additional gene information for DNAH14 Gene
No data available for CIViC summary , Tocris Summary , Gene Wiki entry , PharmGKB "VIP" Summary , fRNAdb sequence ontologies and piRNA Summary for DNAH14 Gene
Genomics for DNAH14 Gene
GeneHancer (GH) Regulatory Elements for DNAH14 Gene
- Top Transcription factor binding sites by QIAGEN in the DNAH14 gene promoter:
Regulatory Element Products
Genomic Locations for DNAH14 Gene
- chr1:224,896,262-225,399,294
- (GRCh38/hg38)
- Size:
- 503,033 bases
- Orientation:
- Plus strand
- chr1:225,083,964-225,586,996
- (GRCh37/hg19)
Genomic View for DNAH14 Gene
- Cytogenetic band:
-
- 1q42.12 by Ensembl
- 1q42.12 by Entrez Gene
- 1q42.12 by HGNC


RefSeq DNA sequence for DNAH14 Gene
Proteins for DNAH14 Gene
-
Protein details for DNAH14 Gene (UniProtKB/Swiss-Prot)
- Protein Symbol:
- Q0VDD8-DYH14_HUMAN
- Recommended name:
- Dynein heavy chain 14, axonemal
- Protein Accession:
- Q0VDD8
- A6NG62
- A6NNL2
- Q0VDD9
- Q4VXC7
- Q4VXG4
- Q4VXG5
- Q5VU33
- Q5VU34
Protein attributes for DNAH14 Gene
- Size:
- 3507 amino acids
- Molecular mass:
- 399895 Da
- Quaternary structure:
-
- Consists of at least two heavy chains and a number of intermediate and light chains.
- SequenceCaution:
-
- Sequence=AAI19717.1; Type=Erroneous initiation; Evidence={ECO:0000305}; Sequence=AAI19718.1; Type=Erroneous initiation; Evidence={ECO:0000305}; Sequence=CAI95150.2; Type=Erroneous gene model prediction; Evidence={ECO:0000305}; Sequence=CAI95641.1; Type=Erroneous gene model prediction; Evidence={ECO:0000305}; Sequence=CAI95642.2; Type=Erroneous gene model prediction; Evidence={ECO:0000305}; Sequence=EAW69735.1; Type=Erroneous gene model prediction; Evidence={ECO:0000305};
Protein Expression for DNAH14 Gene
Post-translational modifications for DNAH14 Gene
Other Protein References for DNAH14 Gene
- ENSEMBL proteins:
- REFSEQ proteins:
-
- NP_001138626.1
- NP_001336840.1
- NP_001336841.1
- NP_001364.1
- NP_659426.2
- XP_006711798.1
- XP_011542359.1
- XP_011542360.1
- XP_011542361.1
- XP_011542363.1
- XP_011542364.1
- XP_011542365.1
- XP_011542366.1
- XP_011542367.1
- XP_011542368.1
- XP_011542369.1
- XP_011542371.1
- XP_011542372.1
- XP_011542373.1
- XP_011542374.1
- XP_011542375.1
- XP_011542376.1
- XP_011542378.1
- XP_011542381.1
- XP_011542383.1
- XP_016855784.1
- XP_016855785.1
- XP_016855786.1
- XP_016855787.1
- XP_016855788.1
- XP_016855789.1
Antibody Products
- Novus Biologicals Antibodies for DNAH14
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Protein Products
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- Search GeneTex for Proteins for DNAH14
Assay Products
No data available for DME Specific Peptides for DNAH14 Gene
Domains & Families for DNAH14 Gene
Gene Families for DNAH14 Gene
- HGNC:
- Human Protein Atlas (HPA):
-
- Predicted intracellular proteins
- Predicted secreted proteins
Protein Domains for DNAH14 Gene
- InterPro:
- ProtoNet:
Suggested Antigen Peptide Sequences for DNAH14 Gene
Graphical View of Domain Structure for InterPro Entry
Q0VDD8UniProtKB/Swiss-Prot:
DYH14_HUMAN :- Dynein heavy chains probably consist of an N-terminal stem (which binds cargo and interacts with other dynein components), and the head or motor domain. The motor contains six tandemly-linked AAA domains in the head, which form a ring. A stalk-like structure (formed by two of the coiled coil domains) protrudes between AAA 4 and AAA 5 and terminates in a microtubule-binding site. A seventh domain may also contribute to this ring; it is not clear whether the N-terminus or the C-terminus forms this extra domain. There are four well-conserved and two non-conserved ATPase sites, one per AAA domain. Probably only one of these (within AAA 1) actually hydrolyzes ATP, the others may serve a regulatory function (By similarity).
- Belongs to the dynein heavy chain family.
- Domain:
-
- Dynein heavy chains probably consist of an N-terminal stem (which binds cargo and interacts with other dynein components), and the head or motor domain. The motor contains six tandemly-linked AAA domains in the head, which form a ring. A stalk-like structure (formed by two of the coiled coil domains) protrudes between AAA 4 and AAA 5 and terminates in a microtubule-binding site. A seventh domain may also contribute to this ring; it is not clear whether the N-terminus or the C-terminus forms this extra domain. There are four well-conserved and two non-conserved ATPase sites, one per AAA domain. Probably only one of these (within AAA 1) actually hydrolyzes ATP, the others may serve a regulatory function (By similarity).
- Family:
-
- Belongs to the dynein heavy chain family.
Function for DNAH14 Gene
Molecular function for DNAH14 Gene
- UniProtKB/Swiss-Prot Function:
- Force generating protein of respiratory cilia. Produces force towards the minus ends of microtubules. Dynein has ATPase activity; the force-producing power stroke is thought to occur on release of ADP. Involved in sperm motility; implicated in sperm flagellar assembly (By similarity).
Phenotypes From GWAS Catalog for DNAH14 Gene
GO ID | Qualified GO term | Evidence | PubMed IDs |
---|---|---|---|
GO:0003774 | motor activity | IEA | -- |
GO:0003777 | microtubule motor activity | IEA | -- |
GO:0005524 | ATP binding | IEA | -- |
GO:0008569 | ATP-dependent microtubule motor activity, minus-end-directed | IBA | -- |
GO:0016887 | ATPase activity | IEA | -- |
Phenotypes for DNAH14 Gene
- GenomeRNAi human phenotypes for DNAH14:
Animal Model Products
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-
-
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CRISPR Products
-
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- Overview of 43 available DNAH14 gene related products
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- Overview of 43 available DNAH14 gene related products
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ViGene Biosciences ready-to-package AAV shRNAs for DNAH14 gene
Clone Products
-
GenScript: Next-day shipping of latest version cDNA ORF clones for DNAH14 in any vector
- XM_011544071.1
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- NM_144989.2
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- NM_001373.1
- XM_011544068.1
- Vector BioLabs ready-to-use adenovirus/AAV for human, mouse, rat
- VectorBuilder custom plasmid, inducible vectors for DNAH14
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-
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- Mammalian expression: PiggyBac
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- Mammalian conditional (Cre-Lox): plasmid and PiggyBac
- Mammalian shRNA knockdown: lentiviral, adenoviral, AAV, and PiggyBac
- CRISPR: plasmid gRNA, lentiviral gRNA, and donor plasmid
- Bacterial expression: pET, pBAD, and pCS
- Yeast expression
- Applied Biological Materials Clones for DNAH14
-
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Cell Line Products
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-
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-
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No data available for Enzyme Numbers (IUBMB) , Human Phenotype Ontology , Animal Models , miRNA , Transcription Factor Targets and HOMER Transcription for DNAH14 Gene
Localization for DNAH14 Gene
Subcellular locations from UniProtKB/Swiss-Prot for DNAH14 Gene
- Cytoplasm, cytoskeleton, cilium axoneme.
- Centrosome (2)
GO ID | Qualified GO term | Evidence | PubMed IDs |
---|---|---|---|
GO:0005737 | cytoplasm | IEA | -- |
GO:0005856 | cytoskeleton | IEA | -- |
GO:0005858 | axonemal dynein complex | IEA | -- |
GO:0005874 | microtubule | IEA | -- |
GO:0005929 | cilium | IBA | -- |
Pathways & Interactions for DNAH14 Gene
SuperPathway | Contained pathways | ||
---|---|---|---|
1 | Respiratory electron transport, ATP synthesis by chemiosmotic coupling, and heat production by uncoupling proteins. |
Pathways by source for DNAH14 Gene
1 KEGG pathway for DNAH14 Gene
Interacting Proteins for DNAH14 Gene
GO ID | Qualified GO term | Evidence | PubMed IDs |
---|---|---|---|
GO:0001539 | cilium or flagellum-dependent cell motility | IEA | -- |
GO:0007018 | microtubule-based movement | IBA | -- |
GO:0060271 | cilium assembly | IBA | -- |
No data available for SIGNOR curated interactions for DNAH14 Gene
Transcripts for DNAH14 Gene
mRNA/cDNA for DNAH14 Gene
- (31) REFSEQ mRNAs :
- (6) Additional mRNA sequences :
- (28) Selected AceView cDNA sequences:
- (19) Ensembl transcripts including schematic representations, and UCSC links where relevant :
Unigene Clusters for DNAH14 Gene
CRISPR Products
-
OriGene CRISPR knockouts for DNAH14
- genomics-online: gRNA clones - Search results for 43 available DNAH14 gene related products
- Overview of 43 available DNAH14 gene related products
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-
Vectors and viruses for KO, Activation, Repression, and more
-
Santa Cruz Biotechnology (SCBT) CRISPR for DNAH14
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miRNA Products
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Inhibitory RNA Products
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- Overview of 43 available DNAH14 gene related products
-
ViGene Biosciences ready-to-package AAV shRNAs for DNAH14 gene
Clone Products
-
GenScript: Next-day shipping of latest version cDNA ORF clones for DNAH14 in any vector
- XM_011544071.1
- NM_001145154.1
- XM_011544074.1
- XM_011544059.1
- XM_011544077.1
- XM_011544061.1
- XM_011544080.1
- XM_011544064.1
- XM_011544083.1
- XM_011544067.1
- XM_011544070.1
- XM_011544073.1
- XM_011544058.1
- XM_011544076.1
- XM_011544060.1
- XM_011544079.1
- XM_011544063.1
- XM_011544082.1
- XM_011544066.1
- XM_011544069.1
- XM_011544072.1
- XM_011544057.1
- NM_144989.2
- XM_011544075.1
- XM_006711735.2
- XM_011544078.1
- XM_011544062.1
- XM_011544081.1
- XM_011544065.1
- NM_001373.1
- XM_011544068.1
- Vector BioLabs ready-to-use adenovirus/AAV for human, mouse, rat
- VectorBuilder custom plasmid, inducible vectors for DNAH14
- VectorBuilder custom lentivirus, adenovirus, AAV vector/virus packaging for DNAH14
-
VectorBuilder Other custom vectors
- Mammalian expression: PiggyBac
- Mammalian Tet-on expression: plasmid
- Mammalian conditional (Cre-Lox): plasmid and PiggyBac
- Mammalian shRNA knockdown: lentiviral, adenoviral, AAV, and PiggyBac
- CRISPR: plasmid gRNA, lentiviral gRNA, and donor plasmid
- Bacterial expression: pET, pBAD, and pCS
- Yeast expression
- Applied Biological Materials Clones for DNAH14
-
Vectors and viruses for ORF, Lenti, Retro, Adenovirus, AAV, and more
Expression for DNAH14 Gene
mRNA differential expression in normal tissues according to GTEx for DNAH14 Gene
Integrated Proteomics: protein expression in normal tissues and cell lines from ProteomicsDB, PaxDb, and MOPED for DNAH14 Gene
NURSA nuclear receptor signaling pathways regulating expression of DNAH14 Gene:
DNAH14SOURCE GeneReport for Unigene cluster for DNAH14 Gene:
Hs.133977Primer Products
-
OriGene qPCR primer pairs for DNAH14
-
OriGene qPCR primer pairs and template standards for DNAH14
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- Overview of 43 available DNAH14 gene related products
No data available for mRNA expression in embryonic tissues and stem cells from LifeMap Discovery , Protein tissue co-expression partners , mRNA Expression by UniProt/SwissProt , Evidence on tissue expression from TISSUES and Phenotype-based relationships between genes and organs from Gene ORGANizer for DNAH14 Gene
Orthologs for DNAH14 Gene
This gene was present in the common ancestor of chordates.
Organism | Taxonomy | Gene | Similarity | Type | Details |
---|---|---|---|---|---|
chimpanzee (Pan troglodytes) |
Mammalia | DNAH14 33 34 |
|
||
cow (Bos Taurus) |
Mammalia | DNAH14 33 |
|
||
dog (Canis familiaris) |
Mammalia | DNAH14 33 |
|
||
mouse (Mus musculus) |
Mammalia | Dnah14 33 16 |
|
||
platypus (Ornithorhynchus anatinus) |
Mammalia | DNAH14 34 |
|
OneToOne | |
oppossum (Monodelphis domestica) |
Mammalia | DNAH14 34 |
|
OneToOne | |
chicken (Gallus gallus) |
Aves | DNAH14 34 |
|
OneToOne | |
tropical clawed frog (Silurana tropicalis) |
Amphibia | dnah14 33 |
|
- Species where no ortholog for DNAH14 was found in the sources mined by GeneCards:
-
- A. gosspyii yeast (Ashbya gossypii)
- Actinobacteria (Mycobacterium tuberculosis)
- African clawed frog (Xenopus laevis)
- African malaria mosquito (Anopheles gambiae)
- Alicante grape (Vitis vinifera)
- alpha proteobacteria (Wolbachia pipientis)
- amoeba (Dictyostelium discoideum)
- Archea (Pyrococcus horikoshii)
- baker's yeast (Saccharomyces cerevisiae)
- barley (Hordeum vulgare)
- beta proteobacteria (Neisseria meningitidis)
- bread mold (Neurospora crassa)
- Chromalveolata (Phytophthora infestans)
- common water flea (Daphnia pulex)
- corn (Zea mays)
- E. coli (Escherichia coli)
- filamentous fungi (Aspergillus nidulans)
- Firmicute bacteria (Streptococcus pneumoniae)
- fission yeast (Schizosaccharomyces pombe)
- fruit fly (Drosophila melanogaster)
- green algae (Chlamydomonas reinhardtii)
- honey bee (Apis mellifera)
- K. lactis yeast (Kluyveromyces lactis)
- lizard (Anolis carolinensis)
- loblloly pine (Pinus taeda)
- malaria parasite (Plasmodium falciparum)
- medicago trunc (Medicago Truncatula)
- moss (Physcomitrella patens)
- orangutan (Pongo pygmaeus)
- pig (Sus scrofa)
- rainbow trout (Oncorhynchus mykiss)
- rat (Rattus norvegicus)
- rice (Oryza sativa)
- rice blast fungus (Magnaporthe grisea)
- schistosome parasite (Schistosoma mansoni)
- sea anemone (Nematostella vectensis)
- sea squirt (Ciona intestinalis)
- sea squirt (Ciona savignyi)
- sea urchin (Strongylocentrotus purpuratus)
- sorghum (Sorghum bicolor)
- soybean (Glycine max)
- stem rust fungus (Puccinia graminis)
- sugarcane (Saccharum officinarum)
- thale cress (Arabidopsis thaliana)
- tomato (Lycopersicon esculentum)
- toxoplasmosis (Toxoplasma gondii)
- Trichoplax (Trichoplax adhaerens)
- wheat (Triticum aestivum)
- worm (Caenorhabditis elegans)
- zebrafish (Danio rerio)
Paralogs for DNAH14 Gene
Paralogs for DNAH14 Gene
(16) SIMAP similar genes for DNAH14 Gene using alignment to 11 proteins:
Variants for DNAH14 Gene
SNP ID | Clin | Chr 01 pos | Variation | AA Info | Type |
---|---|---|---|---|---|
rs786205668 | benign, Non-immune hydrops fetalis | 225,100,772(+) | T/A | coding_sequence_variant, genic_downstream_transcript_variant, genic_upstream_transcript_variant, missense_variant, non_coding_transcript_variant | |
rs560792252 | likely-benign, not specified | 225,272,064(+) | T/C | coding_sequence_variant, genic_downstream_transcript_variant, intron_variant, non_coding_transcript_variant, synonymous_variant | |
rs375791690 | uncertain-significance, not specified | 224,967,434(+) | C/T | coding_sequence_variant, genic_upstream_transcript_variant, missense_variant, non_coding_transcript_variant | |
rs10495237 | benign, not specified | 225,318,592(+) | A/G | coding_sequence_variant, genic_downstream_transcript_variant, missense_variant, non_coding_transcript_variant | |
rs10915816 | benign, not specified | 225,374,683(+) | C/T | genic_downstream_transcript_variant, intron_variant |
Additional Variant Information for DNAH14 Gene
No data available for Polymorphic Variants from UniProtKB/Swiss-Prot for DNAH14 Gene
Disorders for DNAH14 Gene
Disorder | Aliases | PubMed IDs |
---|---|---|
communicating hydrocephalus |
|
|
hydrocephalus, normal-pressure |
|
|
Additional Disease Information for DNAH14
No data available for UniProtKB/Swiss-Prot and Genatlas for DNAH14 Gene
Publications for DNAH14 Gene
- Multiple mouse chromosomal loci for dynein-based motility. (PMID: 8812413) Vaughan KT … Vallee RB (Genomics 1996) 2 3 58
- Identification of novel mutations in endometrial cancer patients by whole-exome sequencing. (PMID: 28339086) Chang YS … Chang JG (International journal of oncology 2017) 3 58
- Architecture of the human interactome defines protein communities and disease networks. (PMID: 28514442) Huttlin EL … Harper JW (Nature 2017) 3 58
- Identification of embryonic lethal genes in humans by autozygosity mapping and exome sequencing in consanguineous families. (PMID: 26036949) Shamseldin HE … Alkuraya FS (Genome biology 2015) 3 58
- The BioPlex Network: A Systematic Exploration of the Human Interactome. (PMID: 26186194) Huttlin EL … Gygi SP (Cell 2015) 3 58
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- Mammalian expression: PiggyBac
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- Mammalian conditional (Cre-Lox): plasmid and PiggyBac
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- CRISPR: plasmid gRNA, lentiviral gRNA, and donor plasmid
- Bacterial expression: pET, pBAD, and pCS
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- Search GeneTex for Proteins for DNAH14
- ViGene Biosciences adenoviral particle packaged cDNA for DNAH14 gene
- ViGene Biosciences lentiviral particle packaged cDNA for DNAH14 gene
- ViGene Biosciences ready-to-package AAV shRNAs for DNAH14 gene
- Search ViGene Biosciences for DNAH14
- Horizon Cell Lines for DNAH14
- genomics-online: cdna clones - Search results for 43 available DNAH14 gene related products
- Overview of 43 available DNAH14 gene related products
- orf clones - Search results for 43 available DNAH14 gene related products
- Overview of 43 available DNAH14 gene related products
- genomics-online: gRNA clones - Search results for 43 available DNAH14 gene related products
- Overview of 43 available DNAH14 gene related products
- genomics-online: primer clones - Search results for 43 available DNAH14 gene related products
- Overview of 43 available DNAH14 gene related products
- genomics-online: shRNA clones - Search results for 43 available DNAH14 gene related products
- Overview of 43 available DNAH14 gene related products
Sources for DNAH14 Gene
- (1) GeneCards
- (2) HGNC
- (3) EntrezGene
- (4) Swiss-Prot
- (5) Ensembl
- (6) OMIM
- (7) GeneLoc
- (8) Gene Wiki
- (9) UCSC
- (10) PhosphoSitePlus
- (11) GO
- (12) TrEMBL
- (13) InterPro
- (14) ProtoNet
- (15) Blocks
- (16) MGI
- (17) IUBMB
- (18) KEGG
- (19) MINT
- (20) STRING
- (21) IntAct
- (22) Novoseek
- (23) PharmGKB
- (24) DrugBank
- (25) HMDB
- (26) UniGene
- (27) AceView
- (28) ASD
- (29) ECgene
- (30) GeneAnnot
- (31) CGAP SAGE
- (32) SOURCE
- (33) HomoloGene
- (34) PanEnsembl
- (35) euGenes
- (36) SGD
- (37) FlyBase
- (38) WormBase
- (39) Pseudogene
- (40) DGV
- (41) dbSNP
- (42) GenAtlas
- (43) HGMD
- (44) GAD
- (45) BGMUT
- (46) HuGE
- (47) Atlas
- (48) Cell Signaling Technology
- (49) GenBank
- (50) H-invDB
- (51) HORDE
- (52) HUGE
- (53) IMGT
- (54) Leiden
- (55) miRBase
- (56) DME
- (57) OriGene
- (58) PubMed
- (59) R&D Systems
- (60) TGDB
- (61) Tocris
- (62) Abcam
- (63) Novus Biologicals
- (64) ProSpec
- (65) Sino Biological
- (66) GenScript
- (67) Qiagen
- (68) Cloud-Clone Corp.
- (69) OCA
- (70) Proteopedia
- (71) MOPED
- (72) neXtProt
- (73) Reactome
- (74) GeneGo (Thomson Reuters)
- (75) fRNAdb
- (76) DISEASES
- (77) SIMAP
- (78) GenomeRNAi
- (79) LifeMap
- (80) miRTarBase
- (81) MalaCards
- (82) Invitrogen
- (83) BitterDB
- (84) Vector BioLabs
- (85) ESI-BIO
- (86) RefSeq
- (87) BioSystems
- (88) MaxQB
- (89) IUPHAR
- (90) BioGPS
- (91) Illumina
- (92) COMPARTMENTS
- (93) HOMER
- (94) PaxDb
- (95) ApexBio
- (96) Addgene
- (97) antibodies-online
- (98) CYP
- (99) NONCODE
- (100) SwitchGear Genomics
- (101) TreeFam
- (102) PathCards
- (103) GeneReviews
- (104) GeneTex
- (105) Taconic Biosciences
- (106) GTEx
- (107) ProteomicsDB
- (108) SCBT
- (109) DGIdb
- (110) ClinicalTrials
- (111) FDA Approved Drugs
- (112) RVIS
- (113) SIGNOR
- (114) diseasecard
- (115) NIH Rare Diseases
- (116) Orphanet
- (117) UMLS
- (118) GTR
- (119) Disease Ontology
- (120) Genetics Home Reference
- (121) MeSH
- (122) MedlinePlus
- (123) CDC
- (124) NINDS
- (125) NCBI Bookshelf
- (126) ClinVar
- (127) Gene Damage Index
- (128) ViGene Biosciences
- (129) HPO
- (130) UDN
- (131) VISTA
- (132) FANTOM5
- (133) ENCODE
- (134) ProSci
- (135) Horizon
- (136) NURSA
- (137) IID
- (138) Cyagen
- (139) VectorBuilder
- (140) SNPedia
- (141) BRCA Exchange
- (142) St John's Lab
- (143) CIViC
- (144) ProteoGenix
- (145) dbSUPER
- (146) TISSUES
- (147) Gene ORGANizer
- (148) abm
- (149) CrownBio
- (150) Human Protein Atlas
- (151) GWAS Catalog
- (152) Monarch Initiative
- (153) DataMed
- (154) HumanCyc
- (155) genomics-online
- (156) UCNEbase
- (157) EPDnew