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Category   Symbol Source: HGNC EntrezGene Ensembl GeneCards RNA genes CroW21
GIFtS

CYTL1 Gene

protein-coding   GIFtS: 40
GCID: GC04M005016

cytokine-like 1
 Explore 3 diseases affiliated with
CYTL1 via
MalaCards
our new
 Human Malady Compendium 
Biological research products
for CYTL1    

Aliases
for CYTL1 gene

(According to 1HGNC, 2Entrez Gene,
3UniProtKB/Swiss-Prot, 4UniProtKB/TrEMBL, 5OMIM, 6GeneLoc, 7Ensembl, 8DME, 9miRBase, and/or 10fRNAdb)
About This Section
Aliases
Cytokine-Like 11 2
C4orf41 2 3
C171 2
Cytokine-Like Protein 12
Cytokine-Like Protein C172
Protein C173

External Ids:    HGNC: 244351   Entrez Gene: 543602   Ensembl: ENSG000001708917   OMIM: 6079305   UniProtKB: Q9NRR13   

Export aliases for CYTL1 gene to outside databases

Previous GC identifer: GC04M005135


Summaries
for CYTL1 gene

(According to Entrez Gene, Tocris Bioscience, Wikipedia's Gene Wiki, PharmGKB,
UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL)
About This Section
Entrez Gene summary for CYTL1:
C17 is a cytokine-like protein specifically expressed in bone marrow and cord blood mononuclear cells that bear the
CD34 (MIM 142230) surface marker (Liu et al., 2000 (PubMed 10857752)).(supplied by OMIM, Mar 2008)




Genomic Views
for CYTL1 gene

(According to GeneLoc and/or HGNC, and/or
Entrez Gene (NCBI build 37),
and/or miRBase,
Genomic Views according to UCSC (hg19) and Ensembl (release 69), Regulatory elements and Epigenetics data according to QIAGEN, SABiosciences, and/or SwitchGear Genomics)
About This Section
RefSeq DNA sequence:
NC_000004.11  NC_018915.1  NT_006051.18  
Regulatory elements:
   SABiosciences Regulatory transcription factor binding sites in the CYTL1 gene promoter:
         GR   p53   GR-beta   NRSF form 1   IRF-1   NRSF form 2   HNF-4alpha1   AREB6   Sox9   GR-alpha   
         Other transcription factors

SwitchGear Promoter luciferase reporter plasmid: CYTL1 promoter sequence
   Search SABiosciences Chromatin IP Primers for CYTL1

Epigenetics:
QIAGEN PyroMark CpG Assay predesigned Pyrosequencing DNA Methylation assays in human, mouse, rat CYTL1


Genomic Location:
Genomic View: UCSC Golden Path with GeneCards custom track

Entrez Gene cytogenetic band: 4p16-p15   Ensembl cytogenetic band:  4p16.2   HGNC cytogenetic band: 4p16-p15

CYTL1 Gene in genomic location: bands according to Ensembl, locations according to (and/or Entrez Gene and/or Ensembl if different)
CYTL1 gene location

GeneLoc information about chromosome 4         GeneLoc Exon Structure

GeneLoc location for GC04M005016:  view genomic region     (about GC identifiers)

Start:
 5,016,313 bp from pter      End:
 5,021,199 bp from pter
Size:
 4,887 bases      Orientation:
 minus strand

Proteins
for CYTL1 gene

(According to 1UniProtKB, HORDE, neXtProt, Ensembl, and/or Reactome, Modification sites according to 2PhosphoSitePlus, Specific Peptides from DME, Protein expression images according to data from SPIRE MOPED and PaxDb, RefSeq according to NCBI, PDB rendering according to OCA and/or Proteopedia, Recombinant Proteins from EMD Millipore, R&D Systems, GenScript, Enzo Life Sciences, OriGene, Novus Biologicals, Sino Biological, ProSpec, and/or Uscn,
Biochemical Assays by EMD Millipore, R&D Systems, OriGene, GenScript, Cell Signaling Technology, Enzo Life Sciences, and/or Uscn, Ontologies according to Gene Ontology Consortium 01 Mar 2013 and Entrez Gene, Antibodies by EMD Millipore, R&D Systems, GenScript, Cell Signaling Technology, OriGene, Novus Biologicals, Thermo Fisher Scientific, Abcam, and/or Uscn)
About This Section
UniProtKB/Swiss-Prot: CYTL1_HUMAN, Q9NRR1 (See protein sequence)
Recommended Name: Cytokine-like protein 1 precursor  
Size: 136 amino acids; 15577 Da
Subcellular location: Secreted

Explore the universe of human proteins at neXtProt for CYTL1: NX_Q9NRR1

Post-translational modifications:

  • View modification sites using PhosphoSitePlus2
  • View neXtProt modification sites for NX_Q9NRR1

    • CYTL1 Protein expression data from MOPED and PaxDb:    About this image 
    Estimated protein expression log10 (pmol).

    REFSEQ proteins: NP_061129.1  
    ENSEMBL proteins: 
     ENSP00000303550   ENSP00000425397   ENSP00000421743  

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    Uscn Proteins for CYTL1

    Gene Ontology (GO): 2 cellular component terms (GO ID links to tree view):    About this table

    GO IDQualified GO termEvidencePubMed IDs
    GO:0005615extracellular space TAS10857752
    GO:0005625soluble fraction ----


    CYTL1 for ontologies           About GeneDecksing



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    Protein Domains /
    Families
    for CYTL1 gene

    (According to InterPro, ProtoNet, UniProtKB, and/or BLOCKS, Sets of similar genes according to GeneDecks)
    About This Section

    ProtoNet protein and cluster: Q9NRR1


    Function
    for CYTL1 gene

    (According to 1UniProtKB, Genatlas, LifeMap Discovery™, IUBMB, and/or 2DME, Human phenotypes from GenomeRNAi, Animal models from MGI Mar 06 2013,
    bound targets from SABiosciences, miRNA Gene Targets from miRTarBase shRNA from OriGene, RNAi from EMD Millipore, siRNAs from OriGene, QIAGEN, microRNA from QIAGEN, Gene Editing from DNA2.0, Clones from EMD Millipore, OriGene, SwitchGear Genomics, GenScript, Sino Biological, DNA2.0, and Vector BioLabs, Cell Lines from GenScript, LifeMap BioReagents, In Situ Hybridization Assays from Advanced Cell Diagnostics, Ontologies according to Gene Ontology Consortium 01 Mar 2013 via Entrez Gene.)
    About This Section
    miRNA
    Products:            		 OriGene 3'-UTR Clone: CYTL1
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    1 QIAGEN miScript miRNA Assays for microRNA that regulate CYTL1:
    hsa-miR-657
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    Inhib. RNA
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    In Situ Assay
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    Gene Ontology (GO): 1 molecular function term (GO ID links to tree view):    About this table

    GO IDQualified GO termEvidencePubMed IDs
    GO:0005102receptor binding TAS10857752


    CYTL1 for ontologies           About GeneDecksing


    Animal Models:
         Mouse knock-out Cytl1tm1Chun for CYTL1
         5 MGI mutant phenotypes (inferred from 1 allele) (MGI details for Cytl1):
     cardiovascular system  growth/size  immune system  respiratory system  skeleton 

    CYTL1 for phenotypes           About GeneDecksing


    Pathways & Interactions
    for CYTL1 gene

    (Pathways according to EMD Millipore, R&D Systems, Cell Signaling Technology, KEGG, PharmGKB, BioSystems, Reactome, Tocris Bioscience, GeneGo (Thomson Reuters), QIAGEN, and/or UniProtKB, Sets of similar genes according to GeneDecks, Interaction Networks according to SABiosciences, and/or STRING, Interactions according to 1UniProtKB, 2MINT, 3I2D, and/or 4STRING, with links to IntAct and Ensembl, Ontologies according to Gene Ontology Consortium 01 Mar 2013 via Entrez Gene).
    About This Section



    Interactions:

        Search SABiosciences Gene Network CentralTM Interacting Genes and Proteins Networks for CYTL1

    Gene Ontology (GO): 5/6 biological process terms (GO ID links to tree view) (see all 6):    About this table

    GO IDQualified GO termEvidencePubMed IDs
    GO:0002062chondrocyte differentiation IEA--
    GO:0007165signal transduction TAS10857752
    GO:0045944positive regulation of transcription from RNA polymerase II promoter IEA--
    GO:0048839inner ear development IEA--
    GO:0050650chondroitin sulfate proteoglycan biosynthetic process IEA--


    CYTL1 for ontologies           About GeneDecksing



    Drugs & Compounds
    for CYTL1 gene

    (Chemical Compounds according to UniProtKB, Enzo Life Sciences, EMD Millipore, Tocris Bioscience HMDB, BitterDB, and/or Novoseek, and Drugs according to DrugBank, Enzo Life Sciences, and/or PharmGKB, with drugs/clinical trials/news search links to CenterWatch)
    About This Section
    Browse Small Molecules at EMD Millipore
    Browse drugs & compounds from Enzo Life Sciences

    Browse Tocris compounds for CYTL1
    Search CenterWatch for drugs/clinical trials and news about CYTL1 

    Transcripts
    for CYTL1 gene

    (Secondary structures according to fRNAdb,
    GenBank/EMBL/DDBJ Accessions according to
    Unigene (Build 235 Homo sapiens; Mar 10 2013) or GenBank,
    RefSeq according to Entrez Gene,
    DOTS (version 10), and/or AceView, transcript ids from Ensembl with links to UCSC,
    exon structure from GeneLoc, alternative splicing isoforms according to ASD and/or ECgene,
    RNAi Products from EMD Millipore,
    siRNAs from OriGene, QIAGEN, shRNA from OriGene, microRNA from QIAGEN,
    Tagged/untagged cDNA clones from OriGene, SwitchGear Genomics, GenScript, DNA2.0, Vector BioLabs, Primers from OriGene, SABiosciences, and/or QIAGEN )
    About This Section
    REFSEQ mRNAs for CYTL1 gene: 
    NM_018659.2  

    Unigene Cluster for CYTL1:

    Cytokine-like 1
    Hs.13872  [show with all ESTs]
    Unigene Representative Sequence: BC031391
    3 Ensembl transcripts including schematic representations, and UCSC links where relevant:
    ENST00000307746(uc003gig.3) ENST00000506508 ENST00000509419

    miRNA
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    1 QIAGEN miScript miRNA Assays for microRNA that regulate CYTL1:
    hsa-miR-657
    SwitchGear 3'UTR luciferase reporter plasmid: CYTL1 3' UTR sequence
    Inhib. RNA
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    OriGene 29mer shRNA kits in GFP-retroviral vector in human, mouse, rat for CYTL1 (see all 7)
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    Clone
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    GenScript: all cDNA clones in your preferred vector: CYTL1 (NM_018659)
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    Vector BioLabs ready-to-use adenovirus/AAV for human, mouse, rat CYTL1 
    Primer
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      QIAGEN QuantiFast Probe-based Assays in human, mouse, rat CYTL1

    Additional cDNA sequence: 

    AF193766.1 AF274956.1 AY359101.1 BC031391.1 

    2 DOTS entries:

    DT.411938  DT.95349397 

    24/83 AceView cDNA sequences (see all 83):

    CR602076 CR620065 BQ045035 AA233071 CD679224 BG897954 BX107819 BF477408 
    BI764244 BQ773859 AL550225 AI127309 CB054753 NM_018659 AI742654 BQ182477 
    BQ183907 BQ183827 AA448648 AF193766 BQ183891 BU624889 CD677388 AA701618 

    GeneLoc Exon Structure

    2 Alternative Splicing Database (ASD) splice patterns (SP) for CYTL1    About this scheme

    ExUns: 1a · 1b ^ 2 ^ 3 ^ 4a · 4b
    SP1:                                    
    SP2:                    -               


    ECgene alternative splicing isoforms for CYTL1

    Expression
    for CYTL1 gene

    (RNA expression data according to H-InvDB, NONCODE, miRBase, and RNAdb, Expression images according to data from BioGPS, Illumina Human BodyMap, and CGAP SAGE, Sets of similar genes according to GeneDecks, in vivo and in vitro expression data from LifeMap Discovery™, plus additional links to Genevestigator, and/or SOURCE, and/or BioGPS, and/or UniProtKB,
    PCR Arrays from SABiosciences, Primers from OriGene, SABiosciences, and/or QIAGEN, In Situ Hybridization Assays from Advanced Cell Diagnostics)
    About This Section

    CYTL1 expression in normal human tissues (normalized intensities)
    See probesets specificity/sensitivity at GeneAnnot
    About this imageBioGPS
    CGAP TAG: --

    Microarray
    RNAseq (Illumina Body Map)
    (100×FPKM)½
    SAGE (Serial Analysis of Gene Expression)
    About this image

    CYTL1 expression in embryonic tissues and stem cells
    Expression by the Database of Embryonic development, Stem cell research, and Regenerative medicine    About this table
    10/11 LifeMap In Vivo Development Anatomical Compartments/Cells (see all 11) 
    Tissue Anatomical Compartment CellCategory (developmental path)
    LimbZeugopodMesenchymal Condensate CellsBone, Cartilage
    LungTracheal CartilageChondrocytesLung
    BoneAutopod Long BoneBone
    BoneEndochondral Facial BonesBone
    BoneStylopod Long BoneBone
    BoneThoracic RibBone
    BoneZeugopod Epiphyseal EndBone
    BoneZeugopod Long BoneBone
    CartilageMeckel's CartilageCartilage
    LimbAutopodLimb
    Expression: Positive    Negative     Selective marker
    Experimental details: Curated     Microarrays     In-situ hybridization
    Stem Cell Differentiation: 8 LifeMap Cells 
    NameCategory
    PureStem™ progenitor T42 (Embryonic Progenitor Cell)
    PureStem™ mesenchymal progenitor 7SMOO32 (Embryonic Progenitor Cell)Adipose, Bone, Cartilage
    PureStem™ mesenchymal progenitor 7PEND24 (Embryonic Progenitor Cell)Cartilage, Neural Crest
    PureStem™ mesenchymal progenitor E15 (Embryonic Progenitor Cell)Adipose, Bone, Cartilage
    PureStem™ progenitor E69 (Embryonic Progenitor Cell)
    PureStem™ progenitor EN2 (Embryonic Progenitor Cell)
    PureStem™ progenitor EN7 (Embryonic Progenitor Cell)
    PureStem™ progenitor Z1 (Embryonic Progenitor Cell)

    See CYTL1 Protein Expression from SPIRE MOPED and PaxDB
    Genevestigator expression for CYTL1

    SOURCE GeneReport for Unigene cluster: Hs.13872

    UniProtKB/Swiss-Prot: CYTL1_HUMAN, Q9NRR1
    Tissue specificity: Specifically expressed in CD34+ hematopoietic cells

        SABiosciences Custom PCR Arrays for CYTL1
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    In Situ
    Assay Products:     
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    Orthologs
    for CYTL1 gene

    (Orthologs according to 1,2HomoloGene (2older version, for species not in 1newer version), 3euGenes, 4SGD , 5MGI Mar 06 2013, with possible further links to Flybase and/or WormBase, and/or 6Ensembl pan taxonomic compara , Gene Trees according to Ensembl and TreeFam)
    About This Section
    This gene was present in the common ancestor of chordates.

    Orthologs for CYTL1 gene from 3/9 species (see all 9)    About this table
    Organism Taxonomic
    classification    		 Gene Description Human
    Similarity    		 Orthology
    Type    		 Details
    mouse
    (Mus musculus)    		 Mammalia Cytl11 , 5 cytokine-like 11, 5 73.93(n)1
    71.43(a)1    		   5 (20.21 cM)5
    2311621  NM_001081106.11  NP_001074575.11 
     377355195 
    chicken
    (Gallus gallus)    		 Aves CYTL11 cytokine-like 1 67.24(n)
    66.38(a)    		   422849  XM_420795.2  XP_420795.1 
    zebrafish
    (Danio rerio)    		 Actinopterygii CABZ01024686.16
    		 --
    		 23(a)
    		 1 ↔ 1
    		 21(41275648-41281649)


    ENSEMBL Gene Tree for CYTL1 (if available)
    TreeFam Gene Tree for CYTL1 (if available) 

    Paralogs
    for CYTL1 gene

    (Paralogs according to 1HomoloGene,
    2Ensembl, and 3SIMAP, Pseudogenes according to 4Pseudogene.org Build 68)
    About This Section
    		  --

    Genomic Variants
    for CYTL1 gene

    (SNPs/Variants according to the 1NCBI SNP Database, 2Ensembl, 3PupaSUITE, UniProtKB, and DNA2.0, Linkage Disequilibrium by HapMap, Structural Variations(CNVs/InDels/Inversions) from the Database of Genomic Variants, Mutations from the Human Gene Mutation Database (HGMD) and the Locus Specific Mutation Databases (LSDB), Blood group antigen gene mutations by BGMUT, Resequencing Primers from QIAGEN, Cancer Mutation PCR Arrays and Assays and Copy Number PCR Arrays from SABiosciences)
    About This Section
    SABiosciences Cancer Mutation PCR Assays
    QIAGEN SeqTarget long-range PCR primers in human, mouse, rat for resequencing CYTL1
    DNA2.0 Custom Variant and Variant Library Synthesis for CYTL1

    Disorders / Diseases
    for CYTL1 gene

    (in which this Gene is Involved, According to MalaCards, OMIM, UniProtKB, the University of Copenhagen DISEASES database, Genatlas, GeneTests, GAD, HuGE Navigator, and/or TGDB.)
    About This Section
    CYTL1 for disorders           About GeneDecksing

    OMIM gene information: 607930    OMIM disorders: --

    3 diseases for CYTL1:    About MalaCards
    multiple sclerosis    osteoarthritis    arthritis


    Export disorders for CYTL1 gene to outside databases

    Publications
    for CYTL1 gene

    (in PubMed. Associations of this gene to articles via 1Entrez Gene, 2UniProtKB/Swiss-Prot, 3HGNC, 4GAD, 5PharmGKB, 6HMDB, 7DrugBank, 8UniProtKB/TrEMBL, 9 Novoseek, and/or 10fRNAdb)
    About This Section

    PubMed articles for CYTL1 gene, integrated from 9 sources (see all 14):
    (articles sorted by number of sources associating them with CYTL1)    		    Utopia: connect your pdf to the dynamic
    world of online information

    1. Molecular cloning and chromosomal mapping of a candidate cytokine gene selectively expressed in human CD34+ cells. (PubMed id 10857752)1, 2, 3 Liu X.... Cheng L. (2000)
    2. The status, quality, and expansion of the NIH full-length cDNA project: the Mammalian Gene Collection (MGC). (PubMed id 15489334)1, 2 Gerhard D.S....Malek J. (2004)
    3. Signal peptide prediction based on analysis of experimentally verified cleavage sites. (PubMed id 15340161)1, 2 Zhang Z. and Henzel W.J. (2004)
    4. The secreted protein discovery initiative (SPDI), a large-scale effort to identify novel human secreted and transmembrane proteins: a bioinformatics assessment. (PubMed id 12975309)1, 2 Clark H.F.... Gray A.M. (2003)
    5. ANKRD7 and CYTL1 are novel risk genes for alcohol drin king behavior. (PubMed id 22613542)1 Chen X.D....Deng H.W. (2012)
    6. Genome-wide analysis of DNA methylation and the gene e xpression change in lung cancer. (PubMed id 22011669)1 Kwon Y.J....Park J.H. (2012)
    7. Identification of CCR2-binding features in Cytl1 by a CCL2-like chemokine model. (PubMed id 21322034)1 Tomczak A. and Pisabarro M.T. (2011)
    8. [Study on the mechanism of C17orf62 induced cell death ]. (PubMed id 21503106)1 Deng X....Shi T.P. (2011)
    9. C17 prevents inflammatory arthritis and associated jo int destruction in mice. (PubMed id 21799806)1 Chao C....Pflanz S. (2011)
    10. Generation and initial analysis of more than 15,000 full-length human and mouse cDNA sequences. (PubMed id 12477932)1 Strausberg R.L....Marra M.A. (2002)

    External Searches for CYTL1 gene

    (in PubMed, OMIM, and NCBI Bookshelf)
    About This Section
     ANDOR
    Aliases
    Free Text  
      Query String
    		PubMed
    OMIM
    NCBI Bookshelf
      (Note: In FireFox, select the above section and copy using Ctrl-C)

    Genome Databases showing CYTL1 gene

    (According to Entrez Gene, HGNC, AceView, euGenes, Ensembl, miRBase, ECgene, Kegg, and/or H-InvDB)
    About This Section
    Entrez Gene: 54360 HGNC: 24435 AceView: C17 Ensembl:ENSG00000170891 euGenes: HUgn54360
    ECgene: CYTL1 H-InvDB: CYTL1

    Other Databases showing CYTL1 gene

    (According to HUGE)
    About This Section
    		   --

    Specialized Databases showing CYTL1 gene

    (According to PharmGKB, ATLAS, HORDE, IMGT, LEIDEN, UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL,
    Wikipedia and/or GeneReviews via UniProtKB/Swiss-Prot)
    About This Section
    NameDescription
    PharmGKB entry for CYTL1 Pharmacogenomics, SNPs, Pathways

    Intellectual Property
    for CYTL1 gene

    (Patent information from GeneIP,
    Licensable technologies from WIS Yeda, Salk, Tufts,
    IP news from LifeMap Sciences, Inc.)
    About This Section
    		Patent Information for CYTL1 gene:
    Search GeneIP for patents involving CYTL1

    GeneCards and IP:
    Japan Patent Office Licenses GeneCards     European Patent Office Licenses GeneCards     Improving the IP Search



    Products
    for CYTL1 gene

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    Advanced Cell Diagnostics
    About This Section

     
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