CTDNEP1 Gene
protein-coding GIFtS: 46
GCID: GC17M007147
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CTD nuclear envelope phosphatase 1(Previous name: dullard homolog (Xenopus laevis) )
(Previous symbol: DULLARD)
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Aliases
for CTDNEP1 gene
(According to
1HGNC,
2Entrez Gene,
3UniProtKB/Swiss-Prot,
4UniProtKB/TrEMBL, 5OMIM, 6GeneLoc,
7Ensembl,
8DME,
9miRBase,
and/or 10fRNAdb)
About This Section
|
| Aliases |
|---|
| CTD Nuclear Envelope Phosphatase 11 2 | | Dullard Homolog (Xenopus Laevis)1 | | DULLARD1 2 3 | | C-Terminal Domain Nuclear Envelope Phosphatase 12 | | HSA0119161 2 | | Dullard Homolog2 | | NET561 2 | | EC 3.1.3.163 | | Serine/Threonine-Protein Phosphatase Dullard2 3 | | |
Export aliases for CTDNEP1 gene to outside databases |
Summaries
for CTDNEP1 gene(According to Entrez Gene,
Tocris Bioscience,
Wikipedia's
Gene Wiki,
PharmGKB,
UniProtKB/Swiss-Prot,
and/or
UniProtKB/TrEMBL)
About This Section
|
UniProtKB/Swiss-Prot: CNEP1_HUMAN, O95476Function: Serine/threonine protein phosphatase forming with CNEP1R1 an active phosphatase complex that dephosphorylatesand may activate LPIN1 and LPIN2. LPIN1 and LPIN2 are phosphatidate phosphatases that catalyze the conversion ofphosphatidic acid to diacylglycerol and control the metabolism of fatty acids at differents levels. May indirectlymodulate the lipid composition of nuclear and/or endoplasmic reticulum membranes and be required for proper nuclearmembrane morphology and/or dynamics. May also indirectly regulate the production of lipid droplets andtriacylglycerol. May antagonize BMP signaling
|
Genomic Views
for CTDNEP1 gene
(According to
GeneLoc and/or
HGNC, and/or
Entrez Gene (NCBI build 37),
and/or miRBase,
Genomic Views according to
UCSC (hg19) and
Ensembl (release 69),
Regulatory elements and Epigenetics data according to
QIAGEN,
SABiosciences, and/or
SwitchGear Genomics)
About This Section
| RefSeq DNA sequence:- NC_000017.10 NC_018928.1 NT_010718.16
Regulatory elements:
 Search SABiosciences Regulatory transcription factor binding sites for CTDNEP1
Other transcription factors
Search SABiosciences Chromatin IP Primers for CTDNEP1
Epigenetics:
| | QIAGEN PyroMark CpG Assay predesigned Pyrosequencing DNA Methylation assays in human, mouse, rat CTDNEP1 |
Genomic Location:
Genomic View: UCSC Golden Path with GeneCards custom track
Entrez Gene cytogenetic band: 17p13 Ensembl cytogenetic band: 17p13.1 HGNC cytogenetic band: 17p13CTDNEP1 Gene in genomic location: bands according to Ensembl, locations according to
 (and/or Entrez Gene and/or Ensembl if different)
GeneLoc information about chromosome 17 GeneLoc Exon Structure GeneLoc location for GC17M007147: view genomic region
(about GC identifiers)
Start:
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7,146,906 bp from pter |
End:
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7,155,810 bp from pter |
Size:
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8,905 bases |
Orientation:
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minus strand |
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Proteins
for CTDNEP1 gene
(According to
1UniProtKB,
HORDE,
neXtProt,
Ensembl,
and/or Reactome,
Modification sites according to 2PhosphoSitePlus,
Specific Peptides from DME,
Protein expression images according to data from SPIRE MOPED and PaxDb,
RefSeq according to NCBI,
PDB rendering according to OCA and/or Proteopedia,
Recombinant Proteins
from
EMD Millipore,
R&D Systems,
GenScript,
Enzo Life Sciences,
OriGene,
Novus Biologicals,
Sino Biological,
ProSpec, and/or
Uscn,
Biochemical Assays by
EMD Millipore,
R&D Systems,
OriGene,
GenScript,
Cell Signaling Technology,
Enzo Life Sciences, and/or
Uscn,
Ontologies according to Gene
Ontology Consortium 01 Mar 2013 and
Entrez Gene,
Antibodies by
EMD Millipore,
R&D Systems,
GenScript,
Cell Signaling Technology,
OriGene,
Novus Biologicals,
Thermo Fisher Scientific,
Abcam, and/or
Uscn)
About This Section
|
UniProtKB/Swiss-Prot: CNEP1_HUMAN, O95476 (See
protein sequence)Recommended Name: CTD nuclear envelope phosphatase 1 Size: 244 amino acids; 28377 Da
Subunit: Interacts with CNEP1R1; the complex dephosphorylates LPIN1 and LPIN2
Subcellular location: Endoplasmic reticulum membrane; Single-pass membrane protein. Nucleus membrane; Single-passmembrane protein
Secondary accessions: D3DTN7 Q96GQ9Explore the universe of human proteins at neXtProt for CTDNEP1: NX_O95476
Post-translational modifications:
View modification sites using PhosphoSitePlus2View neXtProt modification sites for NX_O95476
CTDNEP1 Protein expression data from MOPED and PaxDb: About this image 
REFSEQ proteins (2 alternative transcripts):
NP_001137247.1 NP_056158.2
ENSEMBL proteins: ENSP00000460527 ENSP00000460683 ENSP00000461749 ENSP00000458758 ENSP00000321732 ENSP00000459866 ENSP00000458763 ENSP00000461122 ENSP00000461567 ENSP00000458408
Human Recombinant Protein Products:
Gene Ontology (GO): 5/6 cellular component terms (GO ID links to tree view) (see all 6): About this table
CTDNEP1 for ontologies About GeneDecksing
CTDNEP1 Antibody Products:
Assay Products for CTDNEP1: |
Protein
Domains /
Families
for CTDNEP1 gene(According to InterPro, ProtoNet,
UniProtKB, and/or BLOCKS,
Sets of similar genes according to GeneDecks)
About This Section
|
CTDNEP1 for domains About GeneDecksing
3 InterPro domains/families:Graphical View of Domain Structure for InterPro Entry O95476ProtoNet protein and cluster: O95476 1 Blocks protein family: IPB004274 NLI interacting factor
UniProtKB/Swiss-Prot: CNEP1_HUMAN, O95476Similarity: Belongs to the dullard familySimilarity: Contains 1 FCP1 homology domain |
Function
for CTDNEP1 gene
(According to 1UniProtKB,
Genatlas,
LifeMap Discovery™,
IUBMB, and/or
2DME,
Human phenotypes from GenomeRNAi,
Animal models from MGI Mar 06 2013,
bound targets from SABiosciences,
miRNA Gene Targets from miRTarBase
shRNA from
OriGene,
RNAi from
EMD Millipore,
siRNAs from
OriGene,
QIAGEN,
microRNA from QIAGEN,
Gene Editing from DNA2.0,
Clones from EMD Millipore,
OriGene,
SwitchGear Genomics,
GenScript,
Sino Biological,
DNA2.0,
and Vector BioLabs,
Cell Lines from GenScript,
LifeMap BioReagents,
In Situ Hybridization Assays from Advanced Cell Diagnostics,
Ontologies according to Gene Ontology Consortium 01 Mar 2013 via
Entrez Gene.)
About This Section
|
Function Summary:
UniProtKB/Swiss-Prot: CNEP1_HUMAN, O95476Function: Serine/threonine protein phosphatase forming with CNEP1R1 an active phosphatase complex that dephosphorylatesand may activate LPIN1 and LPIN2. LPIN1 and LPIN2 are phosphatidate phosphatases that catalyze the conversion ofphosphatidic acid to diacylglycerol and control the metabolism of fatty acids at differents levels. May indirectlymodulate the lipid composition of nuclear and/or endoplasmic reticulum membranes and be required for proper nuclearmembrane morphology and/or dynamics. May also indirectly regulate the production of lipid droplets andtriacylglycerol. May antagonize BMP signalingCatalytic activity: A phosphoprotein + H(2)O = a protein + phosphateBiophysicochemical properties: Kinetic parameters: KM=18 mM for p-nitrophenylphosphate; pH dependence: Optimum pH is5.5;Enzyme Number (IUBMB): EC 3.1.3.161
Clone
Products: |  | Browse Clones for the Expression of Recombinant Proteins Available from EMD Millipore | |  | OriGene Myc/DDK tagged cDNA clones in CMV expression vector in human, mouse, rat for CTDNEP1 (see all 3)
OriGene untagged cDNA clones in CMV expression vector in human, mouse, rat for CTDNEP1 (see all 2)
OriGene custom cloning services – gene synthesis, subcloning, mutagenesis,
variant library, vector shuttling 
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In Situ Assay
Products: |
 | Advanced Cell Diagnostics RNAscope RNA in situ hybridization assays for CTDNEP1 |
Gene Ontology (GO): 3 molecular function terms (GO ID links to tree view): About this table
CTDNEP1 for ontologies About GeneDecksing
2 GenomeRNAi human phenotypes for CTDNEP1:
|
Pathways & Interactions
for CTDNEP1 gene
(Pathways according to
EMD Millipore,
R&D Systems,
Cell Signaling Technology,
KEGG,
PharmGKB,
BioSystems,
Reactome,
Tocris Bioscience,
GeneGo (Thomson Reuters),
QIAGEN,
and/or UniProtKB,
Sets of similar genes according to GeneDecks,
Interaction Networks according to
SABiosciences,
and/or STRING,
Interactions according to 1UniProtKB,
2MINT,
3I2D, and/or
4STRING,
with links to IntAct and
Ensembl,
Ontologies according to Gene Ontology Consortium 01 Mar 2013 via
Entrez Gene).
About This Section
|
Interactions:
Search SABiosciences Gene Network CentralTM Interacting Genes and Proteins Networks for CTDNEP1
STRING Interaction
Network Preview (showing 5 interactants - click image to see 25)
5/46 Interacting proteins for CTDNEP1 (O954761, 3 ENSP000003217324) via UniProtKB, MINT, STRING, and/or I2D (see all 46)About this table
Gene Ontology (GO): 4 biological process terms (GO ID links to tree view): About this table | GO ID | Qualified GO term | Evidence | PubMed IDs |
|---|
| GO:0006470 | protein dephosphorylation |
IDA | -- | | GO:0006998 | nuclear envelope organization |
IDA | 17420445 | | GO:0010867 | positive regulation of triglyceride biosynthetic process |
IGI | -- | | GO:0034504 | protein localization to nucleus |
IDA | -- |
CTDNEP1 for ontologies About GeneDecksing
|
Drugs & Compounds
for CTDNEP1 gene(Chemical Compounds according to UniProtKB, Enzo Life Sciences,
EMD Millipore, Tocris Bioscience
HMDB,
BitterDB, and/or
Novoseek, and Drugs according to
DrugBank,
Enzo Life Sciences, and/or
PharmGKB, with drugs/clinical trials/news
search links to CenterWatch)
About This Section
|
 Browse Tocris compounds for CTDNEP1
Search CenterWatch for drugs/clinical trials and news about CTDNEP1 / CNEP1
|
Transcripts
for CTDNEP1 gene(Secondary structures according to
fRNAdb,
GenBank/EMBL/DDBJ Accessions according to
Unigene
(Build 235 Homo sapiens; Mar 10 2013) or GenBank,
RefSeq according to Entrez Gene,
DOTS (version 10), and/or
AceView,
transcript ids from Ensembl
with links to UCSC,
exon structure from GeneLoc,
alternative splicing isoforms according to ASD and/or
ECgene,
RNAi Products from
EMD Millipore,
siRNAs from
OriGene,
QIAGEN,
shRNA from
OriGene,
microRNA from QIAGEN,
Tagged/untagged cDNA clones from
OriGene,
SwitchGear Genomics,
GenScript,
DNA2.0,
Vector BioLabs,
Primers from
OriGene,
SABiosciences, and/or
QIAGEN
)
About This Section
|
REFSEQ mRNAs for CTDNEP1 gene (2 alternative transcripts): NM_001143775.1 NM_015343.4 Unigene Cluster for CTDNEP1: CTD nuclear envelope phosphatase 1 Hs.741244 [show with all ESTs]Unigene Representative Sequence: NM_01534312 Ensembl transcripts including schematic representations, and UCSC links where relevant: ENST00000570828 ENST00000574322(uc002gfd.2 uc002gfe.2) ENST00000573600
ENST00000574205 ENST00000318988 ENST00000572043 ENST00000570484 ENST00000576613
ENST00000575783 ENST00000570385 ENST00000570380 ENST00000571409
Clone
Products: | | OriGene Myc/DDK tagged cDNA clones in CMV expression vector in human, mouse, rat for CTDNEP1 (see all 3)
OriGene untagged cDNA clones in CMV expression vector in human, mouse, rat for CTDNEP1 (see all 2)
OriGene custom cloning services – gene synthesis, subcloning, mutagenesis,
variant library, vector shuttling
| | | GenScript: all cDNA clones in your preferred vector (see all 2): CTDNEP1 (NM_001143775) | | | DNA2.0 Custom Codon Optimized Gene
Synthesis Service for CTDNEP1 | | | Vector BioLabs ready-to-use adenovirus/AAV for human, mouse, rat CTDNEP1 |
Additional cDNA sequence: AJ011916.1 AY364239.1 BC009295.2 24 DOTS entries: DT.99937435 DT.95163850 DT.99957873 DT.450749 DT.120968467 DT.97815358 DT.100822729 DT.100808943 DT.99968689 DT.120968797 DT.100808957 DT.120968539 DT.97860299 DT.120968540 DT.100808950 DT.100808954 DT.120968636 DT.91959307 DT.95370686 DT.100734388 DT.100796871 DT.120862573 DT.120968764 DT.87079522 24/672 AceView cDNA sequences (see all 672): BU179461 AA489517 BE615020 CD675104 BU149493 AI459882 CR611348 BQ888567 CK430552 AJ346040 AI363755 BM851680 AA298491 T32915 BM309960 AA477128 AI383729 CB269137 AA427910 BX384546 BM999373 BG912922 CR606307 AA903542
GeneLoc Exon Structure
|
Expression
for CTDNEP1 gene
(RNA expression data according to
H-InvDB,
NONCODE,
miRBase, and
RNAdb,
Expression images according to data from
BioGPS,
Illumina Human BodyMap, and
CGAP
SAGE,
Sets of similar genes according to GeneDecks,
in vivo and in vitro expression data from LifeMap Discovery™,
plus additional links to
Genevestigator, and/or
SOURCE, and/or
BioGPS, and/or
UniProtKB,
PCR Arrays from
SABiosciences,
Primers from
OriGene,
SABiosciences, and/or
QIAGEN,
In Situ Hybridization Assays from Advanced Cell Diagnostics)
About This Section
| CTDNEP1 expression in normal human tissues (normalized intensities)
See probesets specificity/sensitivity at GeneAnnot
About this imageBioGPS
CGAP TAG: CATTATAACT
About this image
See CTDNEP1 Protein Expression from SPIRE MOPED and PaxDB
Genevestigator expression for CTDNEP1
SOURCE GeneReport for Unigene cluster: Hs.741244
UniProtKB/Swiss-Prot: CNEP1_HUMAN, O95476Tissue specificity: Muscle specific with lower expression in other metabolic tissues
SABiosciences Custom PCR Arrays for CTDNEP1
Primer
Products: | | OriGene genome-wide validated SYBR primer pairs in human, mouse, rat for CTDNEP1
Browse OriGene validated miRNA SYBR primer pairs
| | | SABiosciences RT2 qPCR Primer Assay in human, mouse / rat CTDNEP1 | | | QIAGEN QuantiTect SYBR Green Assays in human, mouse, rat CTDNEP1 | | | QIAGEN QuantiFast Probe-based Assays in human, mouse, rat CTDNEP1 | In Situ
Assay Products: |
| Advanced Cell Diagnostics RNAscope RNA in situ hybridization assays for CTDNEP1 |
Orthologs
for CTDNEP1 gene
(Orthologs according to
1,2HomoloGene (2older version, for species not in 1newer version),
3euGenes,
4SGD
,
5MGI Mar 06 2013,
with possible further links to
Flybase
and/or
WormBase,
and/or
6Ensembl pan taxonomic compara ,
Gene Trees according to Ensembl and
TreeFam)
About This Section
|
This gene was present in the common ancestor of animals.
Orthologs for CTDNEP1 gene from 4/23 species (see all 23) About this table
ENSEMBL Gene Tree for CTDNEP1 (if available)
TreeFam Gene Tree for CTDNEP1 (if available) |
Paralogs
for CTDNEP1 gene(Paralogs according to
1HomoloGene,
2Ensembl, and 3SIMAP, Pseudogenes according to 4Pseudogene.org Build 68)
About This Section
| Paralogs for CTDNEP1 gene
3 SIMAP similar genes for CTDNEP1 using alignment to 7 protein entries: CNEP1_HUMAN (see all proteins):CTDSPL CTDSP2 CTDSP1
CTDNEP1 for paralogs About GeneDecksing
2 Pseudogenes.org Pseudogenes for CTDNEP1
PGOHUM00000238135 PGOHUM00000235650
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Genomic Variants
for CTDNEP1 gene(SNPs/Variants according to the
1NCBI SNP Database,
2Ensembl,
3PupaSUITE,
UniProtKB, and
DNA2.0,
Linkage Disequilibrium by HapMap,
Structural Variations(CNVs/InDels/Inversions) from the Database of Genomic Variants, Mutations from the Human Gene
Mutation Database (HGMD) and the Locus Specific Mutation
Databases (LSDB), Blood group antigen gene mutations by BGMUT,
Resequencing Primers from QIAGEN,
Cancer Mutation PCR Arrays and Assays and Copy Number PCR Arrays from SABiosciences)
About This Section
|
| Genomic Data | Transcription Related Data | Allele Frequencies | | SNP ID | Valid | Clinical
significance | Chr 17 pos | Sequence | # | AA
Chg | Type | More | # | Allele
freq | Pop | Total
sample | More |
|---|
HapMap Linkage Disequilibrium report for CTDNEP1 (7146906 - 7155810 bp)
Structural Variations (Copy Number Variations, Insertions/Deletions, Inversions)
Database of Genomic Variants (DGV) variations for CTDNEP1: -- | SABiosciences Cancer Mutation PCR Assays |
| | QIAGEN SeqTarget long-range PCR primers in human, mouse, rat for resequencing CTDNEP1 |
|
Disorders
/ Diseases
for CTDNEP1 gene
(in which this Gene is Involved, According to MalaCards,
OMIM, UniProtKB,
the University of Copenhagen DISEASES
database,
Genatlas, GeneTests,
GAD,
HuGE Navigator,
and/or TGDB.)
About This Section
|
CTDNEP1 for disorders About GeneDecksing
OMIM gene information: 610684
OMIM disorders: --
1  disease for CTDNEP1: About MalaCardsmalaria
Export disorders for CTDNEP1 gene to outside databases
|
Publications
for CTDNEP1 gene (in
PubMed.
Associations of this gene to articles via
1Entrez Gene,
2UniProtKB/Swiss-Prot,
3HGNC,
4GAD,
5PharmGKB,
6HMDB,
7DrugBank,
8UniProtKB/TrEMBL,
9 Novoseek, and/or
10fRNAdb)
About This Section
|
PubMed articles for CTDNEP1 gene, integrated from 9 sources (see all 13):
(articles sorted by number of sources associating them with CTDNEP1) | |  | Utopia: connect your pdf to the dynamic
world of online information |
- Molecular cloning and characterization of dullard: a novel gene required for neural development. (PubMed id 12083771)1, 2, 3 Satow R.... Asashima M. (2002)
- Nuclear envelope phosphatase-regulatory subunit 1 (formerly TMEM188) is the metazoan SPO7 ortholog and functions in the lipin activation pathway. (PubMed id 22134922)1, 2 Han S.... Goodman J.M. (2011)
- A conserved phosphatase cascade that regulates nuclear membrane biogenesis. (PubMed id 17420445)1, 2 Kim Y....Dixon J.E. (2007)
- Dullard promotes degradation and dephosphorylation of BMP receptors and is required for neural induction. (PubMed id 17141153)1, 3 Satow R....Asashima M. (2006)
- Homo sapiens dullard protein phosphatase shows a pref erence for the insulin-dependent phosphorylation site of lipin1. (PubMed id 21413788)1 Wu R....Allen K.N. (2011)
- Mass spectrometric analysis of lysine ubiquitylation r eveals promiscuity at site level. (PubMed id 21139048)1 Danielsen J.M....Nielsen M.L. (2011)
- A quantitative atlas of mitotic phosphorylation. (PubMed id 18669648)2 Dephoure N.... Gygi S.P. (2008)
- NovelFam3000 -- uncharacterized human protein domains conserved across model organisms. (PubMed id 16533400)2 Kemmer D.... Wasserman W.W. (2006)
- Determinants for dephosphorylation of the RNA polymerase II C- terminal domain by Scp1. (PubMed id 17157258)1 Zhang Y.... Noel J.P. (2006)
- DNA sequence of human chromosome 17 and analysis of rearrangement in the human lineage. (PubMed id 16625196)2 Zody M.C.... Nusbaum C. (2006)
|
External Searches for CTDNEP1 gene
(in PubMed,
OMIM, and NCBI Bookshelf)
About This Section
|
|
Genome Databases showing CTDNEP1 gene
(According to
Entrez Gene,
HGNC,
AceView,
euGenes,
Ensembl,
miRBase,
ECgene,
Kegg,
and/or
H-InvDB)
About This Section
|
|
Other Databases showing CTDNEP1 gene
(According to HUGE)
About This Section
| -- |
Specialized Databases showing CTDNEP1 gene(According to PharmGKB,
ATLAS, HORDE, IMGT, LEIDEN, UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL,
Wikipedia and/or GeneReviews via UniProtKB/Swiss-Prot)
About This Section
|
| Name | Description |
| PharmGKB entry for CTDNEP1 | Pharmacogenomics, SNPs, Pathways |
|
| | |
About This Section
| Patent Information for CTDNEP1 gene:
Search GeneIP for patents involving CTDNEP1
GeneCards and IP:
Japan Patent Office Licenses GeneCards European Patent Office Licenses GeneCards Improving the IP Search
|
Products
for CTDNEP1 gene(Antibodies, recombinant proteins, and assays from EMD Millipore, R&D Systems, OriGene, QIAGEN, GenScript, Cell Signaling Technology, SABiosciences, Novus Biologicals, Sino Biological, Enzo Life Sciences, Abcam, ProSpec, Uscn, Thermo Fisher Scientific, Gene Editing from DNA2.0, Clones from EMD Millipore, OriGene, GenScript, Sino Biological, DNA2.0, SwitchGear Genomics, Vector BioLabs, Cell lines from GenScript and LifeMap BioReagents, PCR Arrays from SABiosciences, Drugs and/or compounds from EMD Millipore, Tocris Bioscience, and/or
Enzo Life Sciences),
In Situ Hybridization Assays from
Advanced Cell Diagnostics
About This Section
|
 | |
 | |
 |
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| | | QIAGEN Custom miScript Target Protector blocks miRNA-binding site of in human, mouse, rat CTDNEP1 | | | QIAGEN SeqTarget long-range PCR primers in human, mouse, rat for resequencing CTDNEP1 | | | QIAGEN PyroMark CpG Assay predesigned Pyrosequencing DNA Methylation assays in human, mouse, rat CTDNEP1 | | | QIAGEN FlexiTube/FlexiPlate siRNA for gene silencing in human, mouse, rat CTDNEP1 | | | QIAGEN QuantiFast Probe-based Assays in human, mouse, rat CTDNEP1 | | | QIAGEN QuantiTect SYBR Green Assays in human, mouse, rat CTDNEP1 |
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| | | | Search Tocris compounds for CTDNEP1 |
| |  |  |  |  | | | | |
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 | | | CTDNEP1 Proteins, Antibodies, CLIAs, and ELISAs |
| | | | | Advanced Cell Diagnostics RNAscope RNA in situ hybridization assays for CTDNEP1 |
|  |  |  | | | | Search ThermoFisher Antibodies for CTDNEP1 |
| | | Vector BioLabs ready-to-use adenovirus/AAV for human, mouse, rat CTDNEP1 |
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