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CEACAM4 Gene

protein-coding GIFtS: 49

GC19M046817

carcinoembryonic antigen-related cell adhesion molecule 4
Symbol approved by the HUGO Gene Nomenclature Committee (HGNC) database
(Previous symbol: CGM7)

(According to ¹HGNC, ²Entrez Gene,
³UniProtKB/Swiss-Prot, ⁴UniProtKB/TrEMBL, ⁵OMIM, ⁶GeneLoc , and/or ⁷Ensembl, ⁸miRBase)
About This Section

Aliases
CGM7 ^², ³
CGM7_HUMAN ^²
NCA ^²

Descriptions
Carcinoembryonic antigen CGM7 ^³
Non-specific cross-reacting antigen W236 ^³
Nonspecific cross-reacting antigen (NCA) ^²
carcinoembryonic antigen gene family member 7 ^²
carcinoembryonic antigen-related cell adhesion molecule 4 ^²
nonspecific cross-reacting antigen W236 ^²

External Ids
HGNC: 1816¹
Entrez Gene: 1089²
UniProtKB: O75871³
Ensembl: ENSG00000105352⁷

Search outside databases for aliases for CEACAM4 gene

Previous GC identifers: GC19M042764 GC19M042517 GC19M046801

(According to Entrez Gene, Wikipedia's Gene Wiki,
UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL)
About This Section

(According to GeneLoc and/or HGNC, and/or
Entrez Gene (NCBI build 36),
and/or miRBase,
Genomic Views according to UCSC and Ensembl, Transcription factor binding sites according to SABiosciences)
About This Section

Genomic View:

UCSC Golden Path with GeneCards custom track

Transcription factor binding sites upstream to the CEACAM4 gene

Entrez Gene cytogenetic band: 19q13.2 Ensembl cytogenetic band: 19q13.2 HGNC cytogenetic band: 19q13.2

CEACAM4 Gene in genomic location: bands according to Ensembl, locations according to (and/or Entrez Gene and/or Ensembl if different)

GeneLoc gene densities for chromosome 19 GeneLoc Exon Structure

GeneLoc location for GC19M046817: (about GC identifiers)

Start:	46,817,184 bp from pter
End:	46,825,282 bp from pter
Size:	8,099 bases
Orientation:	minus strand

RefSeq DNA sequence:

NC_000019.8 NT_011109.15

(According to ¹UniProtKB, and/or Ensembl, Phosphorylation sites according to ²Phosphosite, RefSeq according to NCBI, PDB rendering according to OCA and/or Proteopedia, Recombinant Proteins from Invitrogen, Millipore, Sigma-Aldrich, R&D Systems, Enzo Life Sciences, Abnova, OriGene and/or, Abcam,
Biochemical Assays by Invitrogen, Millipore, R&D Systems, Cell Signaling Technology, and/or Enzo Life Sciences, Ontologies according to Gene Ontology Consortium 01 Apr 2009 and Entrez Gene, Antibodies by Invitrogen, Millipore, Sigma-Aldrich, R&D Systems, Cell Signaling Technology, Abcam, Abnova, and/or Novus Biologicals)
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UniProtKB/Swiss-Prot: CEAM4_HUMAN, O75871 (See protein sequence)

Recommended Name: Carcinoembryonic antigen-related cell adhesion molecule 4 precursor

Size: 244 amino acids; 25909 Da

Subcellular location: Membrane; Single-pass type I membrane protein

Secondary accessions: Q03715 Q7LDZ7

Post-translational modifications:

N-glycosylated¹

REFSEQ proteins: NP_001808.2

ENSEMBL proteins:

ENSP00000221954

Human Recombinant Proteins

	Browse Drug Discovery Central at Invitrogen for human recombinant proteins
	Browse Purified and Recombinant Proteins at Millipore
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	Browse R&D Systems for human recombinant proteins
	Browse recombinant and purified proteins available from Enzo Life Sciences
	Recombinant Proteins from Abcam (pan CEACAM)
	Browse Abnova for Human Recombinant Proteins

Browse Origene for full length recombinant human proteins expressed in human HEK293 cells

2 Gene Ontology (GO) cellular component terms (links to tree view):

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0016020	membrane	IEA	--
GO:0016021	integral to membrane	IEA	--

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Antibodies for CEACAM4:

	Browse Antibodies Central at Invitrogen
	Browse Millipore's Extensive Line of Mono- and Polyclonal Antibodies
	Sigma-Aldrich Antibodies for CEACAM4
	Browse R&D Systems for Antibodies
	Antibodies from Abcam (pan CEACAM), each with their Abpromise^SM
	Browse Abnova for Monoclonal and Polyclonal Antibodies
	Search Novus for antibodies for CEACAM4

Assays for CEACAM4:

	Browse Invitrogen for biochemical assays
	Browse Kits and Assays available from Millipore
	Browse R&D Systems for biochemical assays
	Browse biochemical assays available from Enzo Life Sciences

(According to InterPro, ProtoNet, UniProtKB, and/or BLOCKS, Sets of similar genes according to GeneDecks)
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Graphical View of Domain Structure for InterPro Entry O75871

ProtoNet protein and cluster: O75871

1 Blocks protein family: IPB013106 Immunoglobulin V-set

UniProtKB/Swiss-Prot: CEAM4_HUMAN, O75871

Similarity: Belongs to the immunoglobulin superfamily. CEA family

Similarity: Contains 1 Ig-like V-type (immunoglobulin-like) domain

(According to MGI Jun 06 2009, UniProtKB, IUBMB,and/or Genatlas,
shRNA from OriGene, Sigma-Aldrich, RNAi from Sigma-Aldrich,
RNAi Products, Clones, and Q-PCR Products from Invitrogen, Millipore, OriGene, and/or Abnova, siRNAs from Applied Biosystems, SYBR primers from OriGene, Cell-based Assays from Millipore, Ontologies according to Gene Ontology Consortium 01 Apr 2009 via Entrez Gene.)
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Inhib. RNA:		Invitrogen RNAi Products for gene knock-down (CEACAM4)
		Browse for Gene Knock-down Tools from Millipore
		Browse Abnova for Chimera RNAi Products

OriGene 29mer shRNA kit in GFP-retroviral vector: NM_001817

Applied Biosystems Silencer® siRNAs for CEACAM4

Sigma-Aldrich siRNA and siRNA Panels for CEACAM4

Sigma-Aldrich shRNA for CEACAM4

Explore Sigma-Aldrich super-pooled esiRNAs

Clones:		Browse Clone Ranger at Invitrogen for clones
		Browse Clones for the Expression of Recombinant Proteins Available from Millipore

OriGene GFP tagged cDNA clone in CMV expression vector: NM_001817
Myc/DDK tagged cDNA clone in CMV expression vector: NM_001817
untagged cDNA clone in CMV expression vector: NM_001817

Primers:

Browse Quantitative PCR Central at Invitrogen for Q-PCR LUX™ Primers

OriGene genome-wide validated SYBR primer pairs: NM_001817

Genatlas biochemistry entry for CEACAM4:

carcinoembryonic antigen-related cell adhesion molecule 4 (see CEA@)

(Pathways according to Invitrogen (maps by GeneGo), Millipore, Cell Signaling Technology, Sigma-Aldrich, KEGG and/or UniProtKB,
Sets of similar genes according to GeneDecks, Proteins Network according to SABiosciences, Interactions according to ¹UniProtKB, ²MINT, and/or ³STRING, with links to IntAct and Ensembl, Ontologies according to Gene Ontology Consortium 01 Apr 2009 via Entrez Gene.)
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1 Interacting protein for CEACAM4 (ENSP00000221954³) via UniProtKB, MINT, and/or STRING

Interactant		Interaction Details
GeneCard	External ID(s)	Interaction Details
APOH	ENSP00000205948³	STRING (score=.916)

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(Chemical Compounds according to UniProtKB, Enzo Life Sciences, Sigma-Aldrich, Tocris Bioscience, and/or Novoseek and Drugs according to Enzo Life Sciences and/or PharmGKB)
About This Section

Browse drugs & compounds from Enzo Life Sciences

Browse Small Molecules at Sigma-Aldrich

Browse Tocris compounds for CEACAM4
1 Novoseek chemical compound relationship for CEACAM4 gene

Compound	Score	Articles	PubMed IDs for Articles with Shared Sentences (# sentences)
perchloric acid	41.70	1	8395690 (1)

About this table

(GenBank/EMBL/DDBJ Accessions according to
Unigene (Build 219 Homo sapiens; Jun 2 2009) or GenBank,
RefSeq according to Entrez Gene,
DOTS (version 10), and/or AceView,
non coding RNAs according to RNAdb,
ESTs according to GeneTide,
exon structure from GeneLoc, alternative splicing isoforms according to ASD and/or ECgene,
RNAi Products from Invitrogen, Millipore, and/or Abnova,
siRNAs from Applied Biosystems, Sigma-Aldrich,
shRNA from Sigma-Aldrich, OriGene,
Tagged/untagged cDNA clones from OriGene,
Expression Assays from Applied Biosystems)
About This Section

Inhib. RNA:		Invitrogen RNAi Products for gene knock-down (CEACAM4)
		Browse for Gene Knock-down Tools from Millipore
		Browse Abnova for Chimera RNAi Products

OriGene 29mer shRNA kit in GFP-retroviral vector: NM_001817

Sigma-Aldrich siRNA and siRNA Panels for CEACAM4

Sigma-Aldrich shRNA for CEACAM4

Explore Sigma-Aldrich super-pooled esiRNAs

Applied Biosystems Silencer® siRNAs:

NM_001817

REFSEQ mRNAs for CEACAM4 gene:

NM_001817.2

Applied Biosystems TaqMan ® Gene Expression Assays:

NM_001817

              OriGene GFP tagged cDNA clone in CMV expression vector: NM_001817
                                 Myc/DDK tagged cDNA clone in CMV expression vector: NM_001817
                                 untagged cDNA clone in CMV expression vector: NM_001817

Additional cDNA sequence:

AK298701.1 D90276.1

1 DOTS entry:

DT.75161402

10 AceView cDNA sequences:

BM727358 BV202329 BX644557 BM681735 BX644579 AI721225 NM_001817 AI242105

D90276 BF819374

highest scoring ESTs for CEACAM4:

D90276 AI242105 AI721225 BF819374 BM681735 BM727358 BX644557 BX644579 NM_001817 AV737313

Unigene Cluster for CEACAM4:

Carcinoembryonic antigen-related cell adhesion molecule 4

Hs.12 [show with all ESTs]

Unigene Representative Sequence: NM_001817

GeneLoc Exon Structure

2 Alternative Splicing Database (ASD) splice patterns (SP) for CEACAM4

ExUns:	1	^	2	^	3	^	4	^	5	^	6	^	7
SP1:			-
SP2:

About this scheme

ECgene alternative splicing isoforms for CEACAM4

1 Ensembl transcript including schematic representation:

ENST00000221954

(Experimental results according to ¹GeneNote and GNF BioGPS,
probe sets-to-genes annotations according to ²GeneAnnot , ³GeneTide , Sets of similar genes according to GeneDecks, Electronic Northern calculations according to data from UniGene (Build 219 Homo sapiens), SAGE tags according to CGAP, plus additional links to SOURCE, and/or GNF BioGPS, and/or EXPOLDB, and/or UniProtKB,
Expression Assays from Applied Biosystems )
About This Section

CEACAM4 expression in normal and diseased human tissues

Applied Biosystems TaqMan ® Gene Expression Assays for CEACAM4

¹ / ² / ³

3 probe-sets matching CEACAM4 gene

Affymetrix probe-set	Array	# genes	Sensitivity	Specificity	Correlation	Length	Gb_Accession	Consensus	Uniqueness	Score	Rank
Affymetrix probe-set	Array	GeneAnnot data			GeneNote data		GeneTide data
32490_at²^{, 3}	U95-A	3	1.00	0.91	1.00	1.00	AC005955	0.20	1.00	0.72	1
207205_at²^{, 3}	U133-A	2	1.00	0.95	--	--	NM_001817	0.60	1.00	0.82	1
207205_at²	U133Plus2	2	1.00	0.95	--	--	--	--	--	--	--

GeneDecks CEACAM4 for binary patterns associated with the probe-sets selected above

About GeneDecksing

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Data from

(Publications) and GNF BioGPS
About these images
About these images

CGAP SAGE TAG: --

SOURCE GeneReport for Unigene cluster: Hs.12
Expression variation in blood from EXPOLDB for CEACAM4
UniProtKB/Swiss-Prot: CEAM4_HUMAN, O75871
Tissue specificity: Granulocyte

(Orthologs according to ^1,2HomoloGene (²older version, for species not in ¹newer version), ³euGenes, ⁴SGD and/or ⁵MGI Jun 06 2009, with possible further links to Flybase and/or WormBase, Gene Trees according to Ensembl)
About This Section

Orthologs for CEACAM4 gene from 2 species

Organism	Gene	Locus	Description	Human Similarity	NCBI accessions
chimpanzee (Pan troglodytes)	CEACAM4¹	--	carcinoembryonic antigen-related cell adhesion molecule more	96.84(n) 95.06(a)	456061 XM_512688.2 XP_512688.2
rat (Rattus norvegicus)	Ceacam10¹	--	carcinoembryonic antigen-related cell adhesion molecule more	63.87(n) 48.09(a)	287009 NM_173339.1 NP_775461.1

About this table Species with no ortholog for CEACAM4

ENSEMBL Gene Tree for CEACAM4

(Paralogs according to ¹HomoloGene
and ²Ensembl, Pseudogenes according to ³Pseudogene.org)
About This Section

Paralogs for CEACAM4 gene

CEACAM21² CEACAM3²

(According to the ¹NCBI SNP Database, ²Ensembl, ³PupaSUITE, and UniProtKB, Linkage Disequilibrium by HapMap, Genotyping Reagents from Applied Biosystems)
About This Section

10/83 NCBI SNPs in CEACAM4 are shown (see all 83 )
(Click for Applied Biosystems TaqMan ® Genotyping Assay) (see all 41)

AB	SNP ID	Valid	Chr 19 pos	Sequence	Recs	AA Chg	Type	More	Recs	Allele freq	Pop	Total sample	More
AB	Genomic Data				Transcription Data				Allele Frequencies
				--	--	--		--	--				--
	rs3848568^1,2	A,C,F,H	46824033(-)	`GCACAA/GGGGGA`	1	K/R	mis¹ ese³ trp³		15		EA NA MN EU WA	1226
	rs2052080^1,2	A,C,F,H	46827188(+)	`taaggA/Taatgc`	1	--	ng3¹ tfbs³		11		EU EA WA NA	966
	rs1122493^1,2	C,F,H	46825493(-)	`TGCAGG/AGGTGA`	1	--	ng3¹		8		NA EA EU WA	762
	rs1126454^1,2	C,F,H	46824154(-)	`TCTGGC/GACCCG`	1	H/D	mis¹ ese³		9		EA EU WA NA	2072
--	rs1041994^1,2	A,C,F	46817500(-)	`TCCCCG/AAAGCT`	1	--	ut3¹ ese³		1		MN	184
	rs3745933^1,2	F,H	46817308(-)	`CGGGGG/TCTTGC`	1	--	ut3¹ ese³		5		EA EU WA	1886
--	rs8113662^1,2	A,C	46817479(+)	`TTCCCC/TGTCCC`	1	--	ut3¹ ese³		0	--	--	--	--
	rs1041992^1,2	C,F,H,O	46824113(-)	`CTGCCA/GTCCAG`	1	P/P	syn¹ ese³		9		EA NA EU WA	2050
	rs929503^1,2	A,C,F,H,O	46822818(+)	`GTTCTG/AAGGTC`	1	--	int¹		10		NA EA EU WA	760
	rs929502^1,2	A,C,F,H,O	46822616(+)	`ACGAAA/GCCATT`	1	--	int¹		28		NA EA NS EU WA	3592

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HapMap Linkage Disequilibrium images for CEACAM4 (up to first 250kb)

(in which this Gene is Involved, According to OMIM, UniProtKB, Novoseek, PharmGKB, Genatlas, GeneTests, Blood group antigen gene mutations by BGMUT, HGMD, GAD, HuGE Navigator, BCGD, and/or TGDB.)
About This Section

10/12 Novoseek disease relationships for CEACAM4 gene (see all 12 )

Disease	Score	Articles	PubMed IDs for Articles with Shared Sentences (# sentences)
colon carcinoma	62.86	1	9196893 (1)
colorectal carcinoma	50.83	5	9041239 (2), 9250164 (1), 1962152 (1), 2125040 (1)
tumors	32.64	5	10910050 (1), 9196893 (1), 9756415 (1), 2050678 (1) (see all 5)
adenocarcinoma	31.33	1	7690349 (1)
adenoma	30.04	2	2125040 (2)
myeloid leukemia chronic	25.75	3	2332039 (1), 2011210 (1)
carcinoma	24.55	2	2341397 (1), 2125040 (1)
colon cancer	23.42	1	9085168 (1)
endometrial carcinoma	22.08	1	7690349 (1)
colorectal cancer	4.31	1	9085168 (1)

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(Possibly Related Articles in Doctor's Guide)
About This Section

(in PubMed. Associations of this gene to articles via ¹Novoseek, ²HGNC, ³Entrez Gene, ⁴UniProtKB/Swiss-Prot, ⁵UniProtKB/TrEMBL, ⁶GAD, and/or ⁷PharmGKB)
About This Section

10/75 PubMed articles for CEACAM4 gene (see all 75 ):

Molecular cloning of nonspecific cross-reacting antigens in human granulocytes. (PubMed id 2050678)^{1, 2, 3, 4} Kuroki M....Matsuoka Y. (1991)
The DNA sequence and biology of human chromosome 19. (PubMed id 15057824)⁴ Grimwood J.... Lucas S.M. (2004)
Shotgun sequencing of the human transcriptome with ORF expressed sequence tags. (PubMed id 10737800)³ Dias Neto E....Simpson A.J.G. (2000)
Sequence and glycosylation site identity of two distinct glycoforms of nonspecific cross-reacting antigen as demonstrated by sequence analysis and fast atom bombardment mass spectrometry. (PubMed id 2341397)¹ Hefta S.A....Shively J.E. (1990)
Nonspecific cross-reacting antigen (NCA) expressed by human granulocytes: six species with different peptide sizes and membrane anchoring forms. (PubMed id 2154215)¹ Kuroki M....Matsuoka Y. (1990)
Prognostic significance of secondary cytogenetic changes and nonspecific cross-reacting antigen (NCA) in patients with Ph-positive chronic myeloid leukemia. (PubMed id 2332039)¹ Haus O....Kotlarek-Haus S. (1990)
[Nonspecific cross-reacting antigen (NCA)] (PubMed id 16149618)¹ Hayashi T. and Imai K. (2005)
A polymerase-chain-reaction assay for the specific identification of transcripts encoded by individual carcinoembryonic antigen (CEA)-gene-family members. (PubMed id 7690349)¹ Thompson J....Zimmermann W. (1993)
Identification of three new genes and estimation of the size of the carcinoembryonic antigen family. (PubMed id 1427854)¹ Khan W.N....Hammarstroem S. (1992)
Behavior in immunoblotting of dimeric and monomeric forms of nonspecific cross-reacting antigen (NCA-50) from normal lung. (PubMed id 1919023)¹ Krop-Watorek A....Lisowska E. (1991)

(in PubMed, OMIM, and NCBI Bookshelf)
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		AND	OR
Aliases
Free Text

Query String		PubMed OMIM NCBI Bookshelf
	(Note: In FireFox, select the above section and copy using Ctrl-C)

(According to Entrez Gene, HGNC, AceView, euGenes, Ensembl, miRBase, ECgene, and/or H-InvDB)
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Entrez Gene: 1089	HGNC: 1816	AceView: CEACAM4	Ensembl:ENSG00000105352	euGenes: HUgn1089
ECgene: CEACAM4	H-InvDB: CEACAM4

(According to HUGE)
About This Section

(According to ATLAS, HORDE, IMGT, MTDB, LEIDEN, UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL,
Wikipedia and/or GeneReviews via UniProtKB/Swiss-Prot)
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Name	Description
ATLAS Chromosomes in Cancer entry for CEACAM4	Genetics and Cytogenetics in Oncology and Haematology

Licensable Technologies for CEACAM4
(Available from WIS Yeda, Salk, Tufts)

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(Reagents available from Applied Biosystems, Antibodies and assays by Cell Signaling Technology, Abcam, Novus Biologicals,
Sigma-Aldrich, R&D Systems, Millipore, Abnova, and/or Invitrogen, Clones available from OriGene,and/or Invitrogen, Drugs and/or compounds by Sigma-Aldrich,
Enzo Life Sciences, and/or Tocris Bioscience)
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Products for CEACAM4:

TaqMan ® Gene Expression Assays

TaqMan ® Genotyping Assays

Free SNP selection tool

Invitrogen iPath Pathways	Invitrogen BLOCK-iT™ RNAi
Invitrogen Antibodies	Invitrogen Assays
Invitrogen Clones	Invitrogen Q-PCR Products
Invitrogen Human Recombinant Kinases	Invitrogen Custom Antibody and Peptide Service
Invitrogen Proteins / Assays / Screening Services	Search Invitrogen catalog for CEACAM4-related products

	Millipore Custom Antibody & Bulk Services
	Millipore Preclinical / Clinical Development Services
	Millipore Immunoassay Services
	Millipore Target Screening & Profiling Services

	Predesigned and custom siRNAs for CEACAM4		Antibodies for CEACAM4
	Explore super-pooled esiRNAs		Browse proteins at Sigma-Aldrich
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CEACAM4 Gene

protein-coding GIFtS: 49

Aliases &
Descriptions
for CEACAM4

Summaries
for CEACAM4

Genomic Location
for CEACAM4

Proteins
for CEACAM4

Protein Domains/
Families
for CEACAM4

Gene Function
for CEACAM4

Pathways & Interactions
for CEACAM4

Drugs & Compounds
for CEACAM4

Transcripts
for CEACAM4

Expression
for CEACAM4

Orthologs
for CEACAM4

Paralogs
for CEACAM4

SNPs/Variants
for CEACAM4

Disorders & Mutations
for CEACAM4

Medical News
for CEACAM4

Publications
for CEACAM4

Search for CEACAM4

Genome Databases showing CEACAM4

Other Databases showing CEACAM4

Specialized Databases showing CEACAM4

Licensable Technologies
for CEACAM4

Services
for CEACAM4

Gene Index: 3 5 A B C D E F G H I J K L M N O P Q R S T U V W X Y Z

CEACAM4 Gene

protein-coding GIFtS: 49

Aliases & Descriptionsfor CEACAM4

Summariesfor CEACAM4

Genomic Locationfor CEACAM4

Proteinsfor CEACAM4

Protein Domains/Familiesfor CEACAM4

Gene Functionfor CEACAM4

Pathways & Interactionsfor CEACAM4

Drugs & Compoundsfor CEACAM4

Transcriptsfor CEACAM4

Expression for CEACAM4

Orthologsfor CEACAM4

Paralogsfor CEACAM4

SNPs/Variantsfor CEACAM4

Disorders & Mutationsfor CEACAM4

Medical Newsfor CEACAM4

Publicationsfor CEACAM4

Search for CEACAM4

Genome Databases showing CEACAM4

Other Databases showing CEACAM4

Specialized Databases showing CEACAM4

Licensable Technologiesfor CEACAM4

Servicesfor CEACAM4

Gene Index: 3 5 A B C D E F G H I J K L M N O P Q R S T U V W X Y Z

Aliases &
Descriptions
for CEACAM4

Summaries
for CEACAM4

Genomic Location
for CEACAM4

Proteins
for CEACAM4

Protein Domains/
Families
for CEACAM4

Gene Function
for CEACAM4

Pathways & Interactions
for CEACAM4

Drugs & Compounds
for CEACAM4

Transcripts
for CEACAM4

Expression
for CEACAM4

Orthologs
for CEACAM4

Paralogs
for CEACAM4

SNPs/Variants
for CEACAM4

Disorders & Mutations
for CEACAM4

Medical News
for CEACAM4

Publications
for CEACAM4

Licensable Technologies
for CEACAM4

Services
for CEACAM4