Aliases for CAMK2N2 Gene
Aliases for CAMK2N2 Gene
External Ids for CAMK2N2 Gene
- HGNC: 24197
- Entrez Gene: 94032
- Ensembl: ENSG00000163888
- OMIM: 608721
- UniProtKB: Q96S95
Previous GeneCards Identifiers for CAMK2N2 Gene
- GC03M185461
- GC03M183977
- GC03M181384
Summaries for CAMK2N2 Gene
-
This gene encodes a protein that is highly similar to the rat CaM-KII inhibitory protein, an inhibitor of calcium/calmodulin-dependent protein kinase II (CAMKII). CAMKII regulates numerous physiological functions, including neuronal synaptic plasticity through the phosphorylation of alpha-amino-3-hydroxy-5-methyl-4-isoxazolepropionic acid-type glutamate (AMPA) receptors. Studies of the similar protein in rat suggest that this protein may function as a negative regulator of CaM-KII and may act to inhibit the phosphorylation of AMPA receptors. [provided by RefSeq, Jul 2008]
GeneCards Summary for CAMK2N2 Gene
CAMK2N2 (Calcium/Calmodulin Dependent Protein Kinase II Inhibitor 2) is a Protein Coding gene. GO annotations related to this gene include protein kinase binding and calcium-dependent protein kinase inhibitor activity. An important paralog of this gene is CAMK2N1.
UniProtKB/Swiss-Prot for CAMK2N2 Gene
-
Potent and specific cellular inhibitor of CaM-kinase II (CAMK2). Traps Ca(2+)/calmodulin on CAMK2. May play an important role in the regulation of cell growth when overexpressed in colon adenocarcinoma LoVo cells. Traps Ca(2+)/calmodulin on CAMK2.
No data available for CIViC summary , Tocris Summary , Gene Wiki entry , PharmGKB "VIP" Summary , fRNAdb sequence ontologies and piRNA Summary for CAMK2N2 Gene
Genomics for CAMK2N2 Gene
Regulatory Elements for CAMK2N2 Gene
Enhancers for CAMK2N2 Gene
- Elite enhancer and/or Elite enhancer-gene association
Download GeneHancer data dump
Enhancers around CAMK2N2 on UCSC Golden Path with GeneCards custom track
- Transcription factor binding sites by QIAGEN in the CAMK2N2 gene promoter:
Regulatory Element Products
Genomic Location for CAMK2N2 Gene
- Chromosome:
- 3
- Start:
- 184,259,213 bp from pter
- End:
- 184,261,463 bp from pter
- Size:
- 2,251 bases
- Orientation:
- Minus strand
Genomic View for CAMK2N2 Gene
Genes around CAMK2N2 on UCSC Golden Path with GeneCards custom track
- Cytogenetic band:
-
- 3q27.1 by Ensembl
- 3q27.1 by Entrez Gene
- 3q27.1 by HGNC
CAMK2N2 Gene in genomic location: bands according to
Ensembl, locations according to GeneLoc
(and/or Entrez Gene and/or Ensembl if different)
Genomic Neighborhood
• Exon Structure
• Gene Density
RefSeq DNA sequence for CAMK2N2 Gene
Proteins for CAMK2N2 Gene
-
Protein details for CAMK2N2 Gene (UniProtKB/Swiss-Prot)
- Protein Symbol:
- Q96S95-CK2N2_HUMAN
- Recommended name:
- Calcium/calmodulin-dependent protein kinase II inhibitor 2
- Protein Accession:
- Q96S95
Protein attributes for CAMK2N2 Gene
- Size:
- 79 amino acids
- Molecular mass:
- 8658 Da
- Quaternary structure:
-
- Interacts with CAMK2A and CAMK2B in the presence of Ca(2+)/calmodulin or after autophosphorylation.
Protein Expression for CAMK2N2 Gene
Post-translational modifications for CAMK2N2 Gene
Other Protein References for CAMK2N2 Gene
- ENSEMBL proteins:
- REFSEQ proteins:
Antibody Products
- Novus Biologicals Antibodies for CAMK2N2
- Invitrogen Antibodies for CAMK2N2
- antibodies-online Antibodies for CAMK2N2: See all 17
- GeneTex CAMK2N2 antibody for CAMK2N2
Protein Products
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OriGene Purified Proteins for CAMK2N2
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- Novus Biologicals proteins for CAMK2N2
- ProSpec Recombinant Proteins for CAMK2N2
- antibodies-online Proteins for CAMK2N2: See all 8
- Search antibodies-online for peptides
- Search GeneTex for Proteins for CAMK2N2
-
Abcam proteins for CAMK2N2
Assay Products
- antibodies-online Kits for CAMK2N2: See all 4
No data available for DME Specific Peptides for CAMK2N2 Gene
Domains & Families for CAMK2N2 Gene
Suggested Antigen Peptide Sequences for CAMK2N2 Gene
- GenScript: Design optimal peptide antigens:
Graphical View of Domain Structure for InterPro Entry
Q96S95UniProtKB/Swiss-Prot:
CK2N2_HUMAN :- The inhibitory domain blocks CAMK2 binding to GRIN2B.
- Belongs to the CAMK2N family.
- Domain:
-
- The inhibitory domain blocks CAMK2 binding to GRIN2B.
- Family:
-
- Belongs to the CAMK2N family.
No data available for Gene Families for CAMK2N2 Gene
Function for CAMK2N2 Gene
Molecular function for CAMK2N2 Gene
- UniProtKB/Swiss-Prot Function:
- Potent and specific cellular inhibitor of CaM-kinase II (CAMK2). Traps Ca(2+)/calmodulin on CAMK2. May play an important role in the regulation of cell growth when overexpressed in colon adenocarcinoma LoVo cells. Traps Ca(2+)/calmodulin on CAMK2.
| GO ID | Qualified GO term | Evidence | PubMed IDs |
|---|---|---|---|
| GO:0008427 | calcium-dependent protein kinase inhibitor activity | IEA | -- |
| GO:0019901 | protein kinase binding | IEA | -- |
Phenotypes for CAMK2N2 Gene
- GenomeRNAi human phenotypes for CAMK2N2:
Animal Model Products
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-
-
ViGene Biosciences lentiviral particle packaged cDNA for CAMK2N2 gene
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CRISPR Products
-
OriGene CRISPR knockouts for CAMK2N2
-
Santa Cruz Biotechnology (SCBT) CRISPR for CAMK2N2
- GenScript: Design CRISPR guide RNA sequences for CAMK2N2
miRNA for CAMK2N2 Gene
- miRTarBase miRNAs that target CAMK2N2
-
- hsa-mir-20a-5p (MIRT441022)
- hsa-mir-106b-5p (MIRT441023)
- hsa-mir-4497 (MIRT714620)
- hsa-mir-937-3p (MIRT714621)
- hsa-mir-3621 (MIRT714622)
- hsa-mir-23b-5p (MIRT724036)
- hsa-mir-23a-5p (MIRT724037)
- hsa-mir-519d-3p (MIRT728051)
- hsa-mir-93-5p (MIRT728052)
- hsa-mir-17-5p (MIRT728053)
- hsa-mir-20b-5p (MIRT728054)
miRNA Products
- Search ViGene Biosciences for CAMK2N2
Inhibitory RNA Products
- Origene shRNA, siRNA, and RNAi products in human, mouse, rat for CAMK2N2
- Browse OriGene Inhibitory RNA Products For CAMK2N2
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Clone Products
-
OriGene ORF clones in human for CAMK2N2
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- Vector BioLabs ready-to-use adenovirus/AAV for human, mouse, rat
- VectorBuilder custom plasmid, inducible vectors for CAMK2N2
- VectorBuilder custom lentivirus, adenovirus, AAV vector/virus packaging for CAMK2N2
-
VectorBuilder Other custom vectors
- Mammalian expression: PiggyBac
- Mammalian Tet-on expression: plasmid
- Mammalian conditional (Cre-Lox): plasmid and PiggyBac
- Mammalian shRNA knockdown: lentiviral, adenoviral, AAV, and PiggyBac
- CRISPR: plasmid gRNA, lentiviral gRNA, and donor plasmid
- Bacterial expression: pET, pBAD, and pCS
- Yeast expression
Cell Line Products
-
Horizon Cell Lines for CAMK2N2
-
ViGene Biosciences adenoviral particle packaged cDNA for CAMK2N2 gene
-
ViGene Biosciences lentiviral particle packaged cDNA for CAMK2N2 gene
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Flow Cytometry Products
No data available for Enzyme Numbers (IUBMB) , Human Phenotype Ontology , Animal Models , Transcription Factor Targets and HOMER Transcription for CAMK2N2 Gene
Localization for CAMK2N2 Gene
Subcellular locations from UniProtKB/Swiss-Prot for CAMK2N2 Gene
- Nucleus. Cytoplasm, cytosol. Note=Excluded from nucleus when coexpressed with activated CAMK2. {ECO:0000250}.
| Compartment | Confidence |
|---|---|
| cytoskeleton | 5 |
| nucleus | 5 |
| cytosol | 5 |
| GO ID | Qualified GO term | Evidence | PubMed IDs |
|---|---|---|---|
| GO:0005634 | nucleus | IDA | -- |
| GO:0005737 | cytoplasm | IEA | -- |
| GO:0005813 | centrosome | IDA | -- |
| GO:0005829 | cytosol | IEA | -- |
Pathways & Interactions for CAMK2N2 Gene
No Data Available
Interacting Proteins for CAMK2N2 Gene
| GO ID | Qualified GO term | Evidence | PubMed IDs |
|---|---|---|---|
| GO:0006469 | negative regulation of protein kinase activity | IEA | -- |
No data available for Pathways by source and SIGNOR curated interactions for CAMK2N2 Gene
Transcripts for CAMK2N2 Gene
mRNA/cDNA for CAMK2N2 Gene
- (1) REFSEQ mRNAs :
- (3) Additional mRNA sequences :
- (6) Selected AceView cDNA sequences:
- (1) Ensembl transcripts including schematic representations, and UCSC links where relevant :
Unigene Clusters for CAMK2N2 Gene
CRISPR Products
-
OriGene CRISPR knockouts for CAMK2N2
-
Santa Cruz Biotechnology (SCBT) CRISPR for CAMK2N2
- GenScript: Design CRISPR guide RNA sequences for CAMK2N2
miRNA Products
- Search ViGene Biosciences for CAMK2N2
Inhibitory RNA Products
- Origene shRNA, siRNA, and RNAi products in human, mouse, rat for CAMK2N2
- Browse OriGene Inhibitory RNA Products For CAMK2N2
- Search ViGene Biosciences for CAMK2N2
Clone Products
-
OriGene ORF clones in human for CAMK2N2
- Custom cloning services - gene synthesis, subcloning, mutagenesis, variant library, vector shuttling
- Vector BioLabs ready-to-use adenovirus/AAV for human, mouse, rat
- VectorBuilder custom plasmid, inducible vectors for CAMK2N2
- VectorBuilder custom lentivirus, adenovirus, AAV vector/virus packaging for CAMK2N2
-
VectorBuilder Other custom vectors
- Mammalian expression: PiggyBac
- Mammalian Tet-on expression: plasmid
- Mammalian conditional (Cre-Lox): plasmid and PiggyBac
- Mammalian shRNA knockdown: lentiviral, adenoviral, AAV, and PiggyBac
- CRISPR: plasmid gRNA, lentiviral gRNA, and donor plasmid
- Bacterial expression: pET, pBAD, and pCS
- Yeast expression
Flow Cytometry Products
Expression for CAMK2N2 Gene
mRNA differential expression in normal tissues according to GTEx for CAMK2N2 Gene
This gene is overexpressed in Brain - Hypothalamus (x6.3), Brain - Cerebellum (x5.3), Brain - Cerebellar Hemisphere (x5.0), and Brain - Amygdala (x4.7).
This gene is overexpressed in Lung (69.0).
Integrated Proteomics: protein expression in normal tissues and cell lines from ProteomicsDB, PaxDb, and MOPED for CAMK2N2 Gene
- Elite partner
NURSA nuclear receptor signaling pathways regulating expression of CAMK2N2 Gene:
CAMK2N2SOURCE GeneReport for Unigene cluster for CAMK2N2 Gene:
Hs.585003mRNA Expression by UniProt/SwissProt for CAMK2N2 Gene:
Q96S95-CK2N2_HUMAN
Tissue specificity:
Expressed in cell lines including hemopoietic cell lines and some tumor cell lines. Highly Expressed in stimulated dendritic cell (DC) and weakly expressed in unstimulated mature and immature DC. Highly expressed in kidney, liver, in cell lines HeLaS3, lymphoblastic leukemia MOLT-4, and Burkitts lymphoma Raji. Moderately expressed in heart, skeletal muscle, placenta, and chronic myelogenous leukemia K-562 cells. Weakly expressed in small intestine, colorectal adenocarcinoma SW480, and lung carcinoma A-549 cells.
Evidence on tissue expression from TISSUES for CAMK2N2 Gene
- Nervous system(4.4)
- Blood(4.2)
- Lymph node(4.2)
- Spleen(4.2)
- Stomach(2.2)
Primer Products
-
OriGene qPCR primer pairs for CAMK2N2
No data available for mRNA expression in embryonic tissues and stem cells from LifeMap Discovery and Phenotype-based relationships between genes and organs from Gene ORGANizer for CAMK2N2 Gene
Orthologs for CAMK2N2 Gene
This gene was present in the common ancestor of chordates.
| Organism | Taxonomy | Gene | Similarity | Type | Details |
|---|---|---|---|---|---|
| chimpanzee (Pan troglodytes) |
Mammalia | CAMK2N2 34 35 |
|
||
| oppossum (Monodelphis domestica) |
Mammalia | CAMK2N2 35 |
|
OneToOne | |
| cow (Bos Taurus) |
Mammalia | CAMK2N2 34 35 |
|
||
| dog (Canis familiaris) |
Mammalia | CAMK2N2 34 35 |
|
||
| rat (Rattus norvegicus) |
Mammalia | Camk2n2 34 |
|
||
| mouse (Mus musculus) |
Mammalia | Camk2n2 34 16 35 |
|
||
| chicken (Gallus gallus) |
Aves | LOC101747380 34 |
|
||
| lizard (Anolis carolinensis) |
Reptilia | CAMK2N2 35 |
|
OneToOne | |
| tropical clawed frog (Silurana tropicalis) |
Amphibia | camk2n2 34 |
|
||
| zebrafish (Danio rerio) |
Actinopterygii | camk2n2 35 |
|
OneToMany | |
| camk2n1a 34 35 |
|
- Species where no ortholog for CAMK2N2 was found in the sources mined by GeneCards:
-
- A. gosspyii yeast (Ashbya gossypii)
- Actinobacteria (Mycobacterium tuberculosis)
- African clawed frog (Xenopus laevis)
- African malaria mosquito (Anopheles gambiae)
- Alicante grape (Vitis vinifera)
- alpha proteobacteria (Wolbachia pipientis)
- amoeba (Dictyostelium discoideum)
- Archea (Pyrococcus horikoshii)
- baker's yeast (Saccharomyces cerevisiae)
- barley (Hordeum vulgare)
- beta proteobacteria (Neisseria meningitidis)
- bread mold (Neurospora crassa)
- Chromalveolata (Phytophthora infestans)
- common water flea (Daphnia pulex)
- corn (Zea mays)
- E. coli (Escherichia coli)
- filamentous fungi (Aspergillus nidulans)
- Firmicute bacteria (Streptococcus pneumoniae)
- fission yeast (Schizosaccharomyces pombe)
- fruit fly (Drosophila melanogaster)
- green algae (Chlamydomonas reinhardtii)
- honey bee (Apis mellifera)
- K. lactis yeast (Kluyveromyces lactis)
- loblloly pine (Pinus taeda)
- malaria parasite (Plasmodium falciparum)
- medicago trunc (Medicago Truncatula)
- moss (Physcomitrella patens)
- orangutan (Pongo pygmaeus)
- pig (Sus scrofa)
- platypus (Ornithorhynchus anatinus)
- rainbow trout (Oncorhynchus mykiss)
- rice (Oryza sativa)
- rice blast fungus (Magnaporthe grisea)
- schistosome parasite (Schistosoma mansoni)
- sea anemone (Nematostella vectensis)
- sea squirt (Ciona intestinalis)
- sea squirt (Ciona savignyi)
- sea urchin (Strongylocentrotus purpuratus)
- sorghum (Sorghum bicolor)
- soybean (Glycine max)
- stem rust fungus (Puccinia graminis)
- sugarcane (Saccharum officinarum)
- thale cress (Arabidopsis thaliana)
- tomato (Lycopersicon esculentum)
- toxoplasmosis (Toxoplasma gondii)
- Trichoplax (Trichoplax adhaerens)
- wheat (Triticum aestivum)
- worm (Caenorhabditis elegans)
Paralogs for CAMK2N2 Gene
(1) SIMAP similar genes for CAMK2N2 Gene using alignment to 1 proteins:
Variants for CAMK2N2 Gene
| SNP ID | Clin | Chr 03 pos | Sequence Context | AA Info | Type |
|---|---|---|---|---|---|
| rs1002374443 | -- | 184,263,370(+) | GCAAT(A/G)CCAGC | intron-variant, upstream-variant-2KB | |
| rs1002762361 | -- | 184,260,473(+) | CTCCC(A/G)GTGCG | intron-variant | |
| rs1002792509 | -- | 184,260,144(+) | CCGCC(C/G)GAGCC | intron-variant, utr-variant-3-prime | |
| rs1002941412 | -- | 184,260,323(+) | TGCCC(A/G)TGGCC | intron-variant | |
| rs1002992139 | -- | 184,259,496(+) | TGTCG(G/T)GGGCT | intron-variant, utr-variant-3-prime |
Residual Variation Intolerance Score:
49.5% of all genes are more intolerant (likely to be disease-causing)
Gene Damage Index Score:
0.13;
2.90% of all genes are more intolerant (likely to be disease-causing)
Relevant External Links for CAMK2N2 Gene
- SNPedia medical, phenotypic, and genealogical associations of SNPs for
- CAMK2N2
No data available for Polymorphic Variants from UniProtKB/Swiss-Prot for CAMK2N2 Gene
Disorders for CAMK2N2 Gene
Relevant External Links for CAMK2N2
- Atlas of Genetics and Cytogenetics in Oncology and Haematology:
- CAMK2N2
No disorders were found for CAMK2N2 Gene.
No data available for MalaCards , UniProtKB/Swiss-Prot and Genatlas for CAMK2N2 Gene
Publications for CAMK2N2 Gene
- Molecular cloning and characterization of a novel calcium/calmodulin- dependent protein kinase II inhibitor from human dendritic cells. (PMID: 11444830) Zhang J. … Cao X. (Biochem. Biophys. Res. Commun. 2001) 2 3 4 22 64
- Endogenous human CaMKII inhibitory protein suppresses tumor growth by inducing cell cycle arrest and apoptosis through down-regulation of the phosphatidylinositide 3-kinase/Akt/HDM2 pathway. (PMID: 19586909) Ma S. … Cao X. (J. Biol. Chem. 2009) 3 22 64
- Characterization of a calmodulin kinase II inhibitor protein in brain. (PMID: 9724800) Chang B.H. … Soderling T.R. (Proc. Natl. Acad. Sci. U.S.A. 1998) 2 3 64
- Downregulation of extracellular signal-regulated kinase 1/2 activity by calmodulin KII modulates p21Cip1 levels and survival of immortalized lymphocytes from Alzheimer's disease patients. (PMID: 23153928) Esteras N. … MartA-n-Requero A. (Neurobiol. Aging 2013) 3 64
- Genome wide array analysis indicates that an amyotrophic lateral sclerosis mutation of FUS causes an early increase of CAMK2N2 in vitro. (PMID: 23545117) Convertini P. … Stamm S. (Biochim. Biophys. Acta 2013) 3 64
Products for CAMK2N2 Gene
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- Novus Biologicals Antibodies for CAMK2N2
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- antibodies-online Antibodies for CAMK2N2: See all 17
- antibodies-online Kits for CAMK2N2: See all 4
- antibodies-online Proteins for CAMK2N2: See all 8
- Search antibodies-online for peptides
- GeneTex CAMK2N2 antibody for CAMK2N2
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- ViGene Biosciences adenoviral particle packaged cDNA for CAMK2N2 gene
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- Horizon Cell Lines for CAMK2N2
- Cyagen custom Knockout/knockin (KOKI) mouse models for CAMK2N2
- VectorBuilder custom plasmid, inducible vectors for CAMK2N2
- VectorBuilder custom lentivirus, adenovirus, AAV vector/virus packaging for CAMK2N2
- VectorBuilder Other custom vectors
- Mammalian expression: PiggyBac
- Mammalian Tet-on expression: plasmid
- Mammalian conditional (Cre-Lox): plasmid and PiggyBac
- Mammalian shRNA knockdown: lentiviral, adenoviral, AAV, and PiggyBac
- CRISPR: plasmid gRNA, lentiviral gRNA, and donor plasmid
- Bacterial expression: pET, pBAD, and pCS
- Yeast expression
Sources for CAMK2N2 Gene
- (1) GeneCards
- (2) HGNC
- (3) EntrezGene
- (4) Swiss-Prot
- (5) Ensembl
- (6) OMIM
- (7) GeneLoc
- (8) Gene Wiki
- (9) UCSC
- (10) PhosphoSitePlus
- (11) GO
- (12) TrEMBL
- (13) InterPro
- (14) ProtoNet
- (15) Blocks
- (16) MGI
- (17) IUBMB
- (18) KEGG
- (19) MINT
- (20) STRING
- (21) IntAct
- (22) Novoseek
- (23) PharmGKB
- (24) DrugBank
- (25) HMDB
- (26) UniGene
- (27) AceView
- (28) RNAdb
- (29) ASD
- (30) ECgene
- (31) GeneAnnot
- (32) CGAP SAGE
- (33) SOURCE
- (34) HomoloGene
- (35) PanEnsembl
- (36) euGenes
- (37) SGD
- (38) FlyBase
- (39) WormBase
- (40) Pseudogene
- (41) DGV
- (42) dbSNP
- (43) GenAtlas
- (44) GeneTests
- (45) HGMD
- (46) GAD
- (47) LSDB
- (48) BGMUT
- (49) HuGE
- (50) eBioscience
- (51) Atlas
- (52) Cell Signaling Technology
- (53) GenBank
- (54) H-invDB
- (55) HORDE
- (56) HUGE
- (57) IMGT
- (58) Leiden
- (59) MILLIPORE
- (60) miRBase
- (61) DME
- (62) NCBI
- (63) OriGene
- (64) PubMed
- (65) R&D Systems
- (66) TGDB
- (67) Tocris
- (68) Abcam
- (69) Novus
- (70) ProSpec
- (71) Sino Biological
- (72) GenScript
- (73) Qiagen
- (74) Cloud-Clone Corp.
- (75) Enzo Life Sciences
- (76) OCA
- (77) Proteopedia
- (78) MOPED
- (79) SPIRE
- (80) neXtProt
- (81) Reactome
- (82) GeneGo (Thomson Reuters)
- (83) fRNAdb
- (84) DISEASES
- (85) SIMAP
- (86) GenomeRNAi
- (87) LifeMap
- (88) miRTarBase
- (89) MalaCards
- (90) Invitrogen
- (91) BitterDB
- (92) Vector BioLabs
- (93) ESI-BIO
- (94) RefSeq
- (95) BioSystems
- (96) MaxQB
- (97) IUPHAR
- (98) BioGPS
- (99) Illumina
- (100) COMPARTMENTS
- (101) HOMER
- (102) PaxDb
- (103) ApexBio
- (104) Addgene
- (105) antibodies-online
- (106) CYP
- (107) NONCODE
- (108) SwitchGear Genomics
- (109) TreeFam
- (110) PathCards
- (111) GeneReviews
- (112) GeneTex
- (113) Taconic Biosciences
- (114) GTEx
- (115) ProteomicsDB
- (116) SCBT
- (117) DGIdb
- (118) ClinicalTrials
- (119) FDA Approved Drugs
- (120) RVIS
- (121) SIGNOR
- (122) diseasecard
- (123) NIH Rare Diseases
- (124) Orphanet
- (125) UMLS
- (126) GTR
- (127) Disease Ontology
- (128) Genetics Home Reference
- (129) MeSH
- (130) MedlinePlus
- (131) CDC
- (132) NINDS
- (133) NCBI Bookshelf
- (134) ClinVar
- (135) Gene Damage Index
- (136) ViGene Biosciences
- (137) HPO
- (138) UDN
- (139) VISTA
- (140) FANTOM5
- (141) ENCODE
- (142) ProSci
- (143) Horizon
- (144) NURSA
- (145) IID
- (146) Cyagen
- (147) VectorBuilder
- (148) SNPedia
- (149) BRCA Exchange
- (150) St John's Lab
- (151) CIViC
- (152) ProteoGenix
- (153) dbSUPER
- (154) TISSUES
- (155) Gene ORGANizer