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B3GNT4 Gene

protein-coding GIFtS: 50

GC12P121212

UDP-GlcNAc:betaGal beta-1,3-N-acetylglucosaminyltransferase 4
Symbol approved by the HUGO Gene Nomenclature Committee (HGNC) database

(According to ¹HGNC, ²Entrez Gene,
³UniProtKB/Swiss-Prot, ⁴UniProtKB/TrEMBL, ⁵OMIM, ⁶GeneLoc , and/or ⁷Ensembl, ⁸miRBase)
About This Section

Aliases
B3GN-T4 ^¹, ²
BGnT-4 ^³
Beta-1,3-N-acetylglucosaminyltransferase-4 ^³
Beta3Gn-T4 ^³
EC 2.4.1.- ^³
beta3Gn-T4 ^¹, ²

Descriptions
UDP-GlcNAc:betaGal beta-1,3-N-acetylglucosaminyltransferase 4 ^²
beta-1,3-N-acetylglucosaminyltransferase bGn-T4 ^²

External Ids
HGNC: 15683¹
Entrez Gene: 79369²
UniProtKB: Q9C0J1³
Ensembl: ENSG00000176383⁷

Search outside databases for aliases for B3GNT4 gene

Previous GC identifers: GC12M121471 GC12M122667 GC12P122496 GC12P121125

(According to Entrez Gene, Wikipedia's Gene Wiki,
UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL)
About This Section

EntrezGene summary for B3GNT4:

This gene encodes a member of the beta-1,3-N-acetylglucosaminyltransferase protein family. The

encoded enzyme is involved in the biosynthesis of poly-N-acetyllactosamine chains and prefers

lacto-N-neotetraose as a substrate. It is a type II transmembrane protein. [provided by RefSeq]

UniProtKB/Swiss-Prot: B3GN4_HUMAN, Q9C0J1

Function: Has a beta-1,3-N-acetylglucosaminyltransferase activity for type 2 oligosaccharides

(According to GeneLoc and/or HGNC, and/or
Entrez Gene (NCBI build 36),
and/or miRBase,
Genomic Views according to UCSC and Ensembl, Transcription factor binding sites according to SABiosciences)
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Genomic View:

UCSC Golden Path with GeneCards custom track

Transcription factor binding sites upstream to the B3GNT4 gene

Entrez Gene cytogenetic band: 12q24 Ensembl cytogenetic band: 12q24.31 HGNC cytogenetic band: 12q24

B3GNT4 Gene in genomic location: bands according to Ensembl, locations according to (and/or Entrez Gene and/or Ensembl if different)

GeneLoc gene densities for chromosome 12 GeneLoc Exon Structure

GeneLoc location for GC12P121212: (about GC identifiers)

Start:	121,254,043 bp from pter
End:	121,258,037 bp from pter
Size:	3,995 bases
Orientation:	plus strand

RefSeq DNA sequence:

NC_000012.10 NT_009755.18

(According to ¹UniProtKB, and/or Ensembl, Phosphorylation sites according to ²Phosphosite, RefSeq according to NCBI, PDB rendering according to OCA and/or Proteopedia, Recombinant Proteins from Invitrogen, Millipore, Sigma-Aldrich, R&D Systems, Enzo Life Sciences, Abnova, OriGene and/or, Abcam,
Biochemical Assays by Invitrogen, Millipore, R&D Systems, Cell Signaling Technology, and/or Enzo Life Sciences, Ontologies according to Gene Ontology Consortium 01 Apr 2009 and Entrez Gene, Antibodies by Invitrogen, Millipore, Sigma-Aldrich, R&D Systems, Cell Signaling Technology, Abcam, Abnova, and/or Novus Biologicals)
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UniProtKB/Swiss-Prot: B3GN4_HUMAN, Q9C0J1 (See protein sequence)

Recommended Name: UDP-GlcNAc:betaGal beta-1,3-N-acetylglucosaminyltransferase 4

Size: 378 amino acids; 42310 Da

Subcellular location: Golgi apparatus membrane; Single-pass type II membrane protein (By

similarity)

Secondary accessions: Q8N5W4 Q8N934 Q8ND21 Q8WWR5 Q8WY02 Q96QH5

Alternative splicing: 2 isoforms: Q9C0J1-1 Q9C0J1-2

REFSEQ proteins: NP_110392.1

ENSEMBL proteins:

ENSP00000319636

Human Recombinant Proteins

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	Browse Human Recombinant Proteins at Sigma-Aldrich
	Browse R&D Systems for human recombinant proteins
	Browse recombinant and purified proteins available from Enzo Life Sciences
	Recombinant Proteins from Abcam (B3GNT4)
	Human Recombinant Proteins from Abnova (B3GNT4)

Browse Origene for full length recombinant human proteins expressed in human HEK293 cells

4 Gene Ontology (GO) cellular component terms (links to tree view):

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0000139	Golgi membrane	IEA	--
GO:0005794	Golgi apparatus	IEA	--
GO:0016020	membrane	IEA	--
GO:0016021	integral to membrane	IEA	--

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Antibodies for B3GNT4:

	Browse Antibodies Central at Invitrogen
	Browse Millipore's Extensive Line of Mono- and Polyclonal Antibodies
	Browse Antibodies at Sigma-Aldrich
	Browse R&D Systems for Antibodies
	Antibodies from Abcam (B3GNT4), each with their Abpromise^SM
	Browse Abnova for Monoclonal and Polyclonal Antibodies
	Search Novus for antibodies for B3GNT4

Assays for B3GNT4:

	Browse Invitrogen for biochemical assays
	Browse Kits and Assays available from Millipore
	Browse R&D Systems for biochemical assays
	Browse biochemical assays available from Enzo Life Sciences

(According to InterPro, ProtoNet, UniProtKB, and/or BLOCKS, Sets of similar genes according to GeneDecks)
About This Section

Graphical View of Domain Structure for InterPro Entry Q9C0J1

ProtoNet protein and cluster: Q9C0J1

1 Blocks protein family: IPB002659 Glycosyl transferase

UniProtKB/Swiss-Prot: B3GN4_HUMAN, Q9C0J1

Similarity: Belongs to the glycosyltransferase 31 family

(According to MGI Jun 06 2009, UniProtKB, IUBMB,and/or Genatlas,
shRNA from OriGene, Sigma-Aldrich, RNAi from Sigma-Aldrich,
RNAi Products, Clones, and Q-PCR Products from Invitrogen, Millipore, OriGene, and/or Abnova, siRNAs from Applied Biosystems, SYBR primers from OriGene, Cell-based Assays from Millipore, Ontologies according to Gene Ontology Consortium 01 Apr 2009 via Entrez Gene.)
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Inhib. RNA:		Invitrogen RNAi Products for gene knock-down (B3GNT4)
		Browse for Gene Knock-down Tools from Millipore
		Browse Abnova for Chimera RNAi Products

OriGene 29mer shRNA kit in GFP-retroviral vector: NM_030765

Applied Biosystems Silencer® siRNAs for B3GNT4

Sigma-Aldrich siRNA for B3GNT4

Sigma-Aldrich shRNA for B3GNT4

Explore Sigma-Aldrich super-pooled esiRNAs

Clones:		Browse Clone Ranger at Invitrogen for clones
		Browse Clones for the Expression of Recombinant Proteins Available from Millipore

OriGene GFP tagged cDNA clone in CMV expression vector: NM_030765
Myc/DDK tagged cDNA clone in CMV expression vector: NM_030765
untagged cDNA clone in CMV expression vector: NM_030765

Primers:

Browse Quantitative PCR Central at Invitrogen for Q-PCR LUX™ Primers

OriGene genome-wide validated SYBR primer pairs: NM_030765

UniProtKB/Swiss-Prot: B3GN4_HUMAN, Q9C0J1

Function: Has a beta-1,3-N-acetylglucosaminyltransferase activity for type 2 oligosaccharides

Enzyme Number (IUBMB): EC 2.4.1.-

2 Gene Ontology (GO) molecular function terms (links to tree view):

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0008378	galactosyltransferase activity	IEA	--
GO:0016757	transferase activity, transferring glycosyl groups	IEA	--

About this table

(Pathways according to Invitrogen (maps by GeneGo), Millipore, Cell Signaling Technology, Sigma-Aldrich, KEGG and/or UniProtKB,
Sets of similar genes according to GeneDecks, Proteins Network according to SABiosciences, Interactions according to ¹UniProtKB, ²MINT, and/or ³STRING, with links to IntAct and Ensembl, Ontologies according to Gene Ontology Consortium 01 Apr 2009 via Entrez Gene.)
About This Section

UniProtKB/Swiss-Prot: B3GN4_HUMAN, Q9C0J1

Pathway: Protein modification; protein glycosylation

5/16 Interacting proteins for B3GNT4 (Q9C0J1² ENSP00000319636³) via UniProtKB, MINT, and/or STRING (see all 16 )

Interactant		Interaction Details
GeneCard	External ID(s)	Interaction Details
GOLM1	Q8NBJ4²	MINT-64978
B3GNT1	ENSP00000305595³	STRING (score=.911)
B4GALT1	ENSP00000369055³	STRING (score=.899)
B4GALT2	ENSP00000361399³	STRING (score=.899)
B4GALT3	ENSP00000356977³	STRING (score=.899)

About this table

1 Gene Ontology (GO) biological process term (links to tree view):

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0006486	protein amino acid glycosylation	IEA	--

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(Chemical Compounds according to UniProtKB, Enzo Life Sciences, Sigma-Aldrich, Tocris Bioscience, and/or Novoseek and Drugs according to Enzo Life Sciences and/or PharmGKB)
About This Section

Browse drugs & compounds from Enzo Life Sciences

Browse Small Molecules at Sigma-Aldrich

Browse Tocris compounds for B3GNT4

(GenBank/EMBL/DDBJ Accessions according to
Unigene (Build 219 Homo sapiens; Jun 2 2009) or GenBank,
RefSeq according to Entrez Gene,
DOTS (version 10), and/or AceView,
non coding RNAs according to RNAdb,
ESTs according to GeneTide,
exon structure from GeneLoc, alternative splicing isoforms according to ASD and/or ECgene,
RNAi Products from Invitrogen, Millipore, and/or Abnova,
siRNAs from Applied Biosystems, Sigma-Aldrich,
shRNA from Sigma-Aldrich, OriGene,
Tagged/untagged cDNA clones from OriGene,
Expression Assays from Applied Biosystems)
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Inhib. RNA:		Invitrogen RNAi Products for gene knock-down (B3GNT4)
		Browse for Gene Knock-down Tools from Millipore
		Browse Abnova for Chimera RNAi Products

OriGene 29mer shRNA kit in GFP-retroviral vector: NM_030765

Sigma-Aldrich siRNA for B3GNT4

Sigma-Aldrich shRNA for B3GNT4

Explore Sigma-Aldrich super-pooled esiRNAs

Applied Biosystems Silencer® siRNAs:

NM_030765

REFSEQ mRNAs for B3GNT4 gene:

NM_030765.2

Applied Biosystems TaqMan ® Gene Expression Assays:

NM_030765

              OriGene GFP tagged cDNA clone in CMV expression vector: NM_030765
                                 Myc/DDK tagged cDNA clone in CMV expression vector: NM_030765
                                 untagged cDNA clone in CMV expression vector: NM_030765

Additional cDNA sequence:

AB049586.1 AJ130848.1 AJ278962.1 AL834452.1 AY358940.1 BC031399.1 CR591685.1

3 DOTS entries:

DT.431320 DT.100031197 DT.95297085

24/39 AceView cDNA sequences (see all 39 ):

R41690 CR591685 BX108933 NM_030765 AL522684 BC030247 CA416426 BX090781

AL834452 AA133381 AA704041 AI910059 BC031399 AA936705 AY358940 AJ130848

AB049586 AK095746 BI916321 AJ278962 AI961337 BM820715 AI690104 AL522683

highest scoring ESTs for B3GNT4:

AA133340 AA133381 AB049586 AI267992 AI628729 AI690104 AI961337 AJ130848 AJ278962 AL522683

Unigene Cluster for B3GNT4:

UDP-GlcNAc:betaGal beta-1,3-N-acetylglucosaminyltransferase 4

Hs.363315 [show with all ESTs]

Unigene Representative Sequence: AY358940

GeneLoc Exon Structure

1 Ensembl transcript including schematic representation:

ENST00000324189

(Experimental results according to ¹GeneNote and GNF BioGPS,
probe sets-to-genes annotations according to ²GeneAnnot , ³GeneTide , Sets of similar genes according to GeneDecks, Electronic Northern calculations according to data from UniGene (Build 219 Homo sapiens), SAGE tags according to CGAP, plus additional links to SOURCE, and/or GNF BioGPS, and/or EXPOLDB, and/or UniProtKB,
Expression Assays from Applied Biosystems )
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B3GNT4 expression in normal and diseased human tissues

Applied Biosystems TaqMan ® Gene Expression Assays for B3GNT4

¹ / ² / ³

3 probe-sets matching B3GNT4 gene

Affymetrix probe-set	Array	# genes	Sensitivity	Specificity	Correlation	Length	Gb_Accession	Consensus	Uniqueness	Score	Rank
Affymetrix probe-set	Array	GeneAnnot data			GeneNote data		GeneTide data
72668_s_at²^{, 3}	U95-D	1	1.00	1.00	1.00	1.00	AI348773	0.40	1.00	0.76	1
221240_s_at²^{, 3}	U133-A	1	1.00	1.00	--	--	NM_030765	0.60	1.00	0.82	1
221240_s_at²	U133Plus2	1	1.00	1.00	--	--	--	--	--	--	--

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Data from

(Publications) and GNF BioGPS
About these images
About these images

CGAP SAGE TAG: TGGGCACTGG

SOURCE GeneReport for Unigene cluster: Hs.363315
UniProtKB/Swiss-Prot: B3GN4_HUMAN, Q9C0J1
Tissue specificity: Mainly expressed in brain tissues such as whole brain, hippocampus, amygdala,
cerebellum and caudate nucleus. Also expressed in colon, esophagus and kidney

(Orthologs according to ^1,2HomoloGene (²older version, for species not in ¹newer version), ³euGenes, ⁴SGD and/or ⁵MGI Jun 06 2009, with possible further links to Flybase and/or WormBase, Gene Trees according to Ensembl)
About This Section

Orthologs for B3GNT4 gene from 5/6 species (see all 6 )

Organism	Gene	Locus	Description	Human Similarity	NCBI accessions
dog (Canis familiaris)	B3GNT4¹	--	UDP-GlcNAc:betaGal beta-1,3-N-acetylglucosaminyltransferase more	84.8(n) 87.09(a)	486255 XM_543380.2 XP_543380.2
chimpanzee (Pan troglodytes)	B3GNT4¹	--	UDP-GlcNAc:betaGal beta-1,3-N-acetylglucosaminyltransferase more	98.77(n) 98.15(a)	744166 XM_001166798.1 XP_001166798.1
cow (Bos taurus)	B3GNT4¹	--	UDP-GlcNAc:betaGal beta-1,3-N-acetylglucosaminyltransferase more	85.32(n) 85.51(a)	534619 XM_868278.2 XP_873371.2
rat (Rattus norvegicus)	B3gnt4¹	--	UDP-GlcNAc:betaGal beta-1,3-N-acetylglucosaminyltransferase more	81.89(n) 80.35(a)	288752 XM_222292.2 XP_222292.1
mouse (Mus musculus)	B3gnt4^{1, 5}	5⁵	UDP-GlcNAc:betaGal beta-1,3-N-acetylglucosaminyltransferase more^{1, 5}	80.06(n)¹ 78.9(a)¹	231727¹ NM_198611.1¹ NP_941013.1¹ AY037786⁵ BC114987⁵ (see all 6)

About this table Species with no ortholog for B3GNT4

ENSEMBL Gene Tree for B3GNT4

(Paralogs according to ¹HomoloGene
and ²Ensembl, Pseudogenes according to ³Pseudogene.org)
About This Section

Paralogs for B3GNT4 gene

B3GNT3² B3GNT7²

(According to the ¹NCBI SNP Database, ²Ensembl, ³PupaSUITE, and UniProtKB, Linkage Disequilibrium by HapMap, Genotyping Reagents from Applied Biosystems)
About This Section

10/13 NCBI SNPs in B3GNT4 are shown (see all 13 )
(Click for Applied Biosystems TaqMan ® Genotyping Assay) (see all 31)

AB	SNP ID	Valid	Chr 12 pos	Sequence	Recs	AA Chg	Type	More	Recs	Allele freq	Pop	Total sample	More
AB	Genomic Data				Transcription Data				Allele Frequencies
				--	--	--		--	--				--
	rs1001178^1,2	C,F,H	121256998(-)	`GCTAGA/TCACTG`	1	S/T	mis¹ ese³		6		MN EU EA WA NA	634
	rs7136356^1,2	A,C,F	121255134(+)	`CCCAGC/GCTTCC`	1	P/A	mis¹ ese³		3		NA	226
	rs35203505^1,2	F	121257011(+)	`CTCTCT/CGTCCC`	1	P/L	mis¹		1		NA	68
	rs12827843^1,2	F	121255114(+)	`GCTGCC/GGTTCA`	1	--	ut5¹ ese³		4		NA EA	362
--	rs879132^1,2	A	121254733(-)	`CTCGGG/ATCCCC`	1	--	int¹		1		MN	184
	rs12828049^1,2	F	121255197(+)	`TCTTTC/TACCGC`	1	--	int¹		2		NA	182
--	rs59453864^1,2	--	121257494(+)	`CATCCA/GCCAAG`	1	R/H	mis¹		0	--	--	--	--
--	rs61302482^1,2	--	121255563(+)	`CTAGCC/TGGGCT`	1	--	int¹		0	--	--	--	--
--	rs7304287^1,2	--	121255448(+)	`tcgctG/Tgaacc`	1	--	int¹		0	--	--	--	--
--	rs73419785^1,2	--	121256474(+)	`CCAAGC/TTAAAC`	1	--	int¹		0	--	--	--	--

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HapMap Linkage Disequilibrium images for B3GNT4 (up to first 250kb)

(in which this Gene is Involved, According to OMIM, UniProtKB, PharmGKB, Genatlas, GeneTests, Blood group antigen gene mutations by BGMUT, HGMD, GAD, HuGE Navigator, BCGD, and/or TGDB.)
About This Section

OMIM: 605864

(Possibly Related Articles in Doctor's Guide)
About This Section

(in PubMed. Associations of this gene to articles via ¹Novoseek, ²HGNC, ³Entrez Gene, ⁴UniProtKB/Swiss-Prot, ⁵UniProtKB/TrEMBL, ⁶GAD, and/or ⁷PharmGKB)
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7 PubMed articles for B3GNT4 gene:

Identification and characterization of three novel beta 1,3-N-acetylglucosaminyltransferases structurally related to the beta 1,3-galactosyltransferase family. (PubMed id 11042166)^{2, 3, 4} Shiraishi N....Sasaki K. (2001)
Complete sequencing and characterization of 21,243 full-length human cDNAs. (PubMed id 14702039)^{3, 4} Ota T....Sugano S. (2004)
The secreted protein discovery initiative (SPDI), a large-scale effort to identify novel human secreted and transmembrane proteins: a bioinformatics assessment. (PubMed id 12975309)^{3, 4} Clark H.F....Gray A. (2003)
Identification and characterization of large galactosyltransferase gene families: galactosyltransferases for all functions. (PubMed id 10580128)^{3, 4} Amado M.... Clausen H. (1999)
The full-ORF clone resource of the German cDNA Consortium. (PubMed id 17974005)⁴ Bechtel S....Schupp I. (2007)
The status, quality, and expansion of the NIH full-length cDNA project: the Mammalian Gene Collection (MGC). (PubMed id 15489334)⁴ Gerhard D.S....Malek J. (2004)
Generation and initial analysis of more than 15,000 full-length human and mouse cDNA sequences. (PubMed id 12477932)³ Strausberg R.L....Marra M.A. (2002)

(in PubMed, OMIM, and NCBI Bookshelf)
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		AND	OR
Aliases
Free Text

Query String		PubMed OMIM NCBI Bookshelf
	(Note: In FireFox, select the above section and copy using Ctrl-C)

(According to Entrez Gene, HGNC, AceView, euGenes, Ensembl, miRBase, ECgene, and/or H-InvDB)
About This Section

Entrez Gene: 79369	HGNC: 15683	AceView: B3GNT4	Ensembl:ENSG00000176383	euGenes: HUgn79369
ECgene: B3GNT4	H-InvDB: B3GNT4

(According to HUGE)
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(According to ATLAS, HORDE, IMGT, MTDB, LEIDEN, UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL,
Wikipedia and/or GeneReviews via UniProtKB/Swiss-Prot)
About This Section

Name	Description
GGDB	http://riodb.ibase.aist.go.jp/rcmg/ggdb/Homolog?cat=symbol&symbol=B3GNT4

Licensable Technologies for B3GNT4
(Available from WIS Yeda, Salk, Tufts)

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(Reagents available from Applied Biosystems, Antibodies and assays by Cell Signaling Technology, Abcam, Novus Biologicals,
Sigma-Aldrich, R&D Systems, Millipore, Abnova, and/or Invitrogen, Clones available from OriGene,and/or Invitrogen, Drugs and/or compounds by Sigma-Aldrich,
Enzo Life Sciences, and/or Tocris Bioscience)
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Products for B3GNT4:

TaqMan ® Gene Expression Assays

TaqMan ® Genotyping Assays

Free SNP selection tool

Invitrogen iPath Pathways	Invitrogen BLOCK-iT™ RNAi
Invitrogen Antibodies	Invitrogen Assays
Invitrogen Clones	Invitrogen Q-PCR Products
Invitrogen Human Recombinant Kinases	Invitrogen Custom Antibody and Peptide Service
Invitrogen Proteins / Assays / Screening Services	Search Invitrogen catalog for B3GNT4-related products

	Millipore Custom Antibody & Bulk Services
	Millipore Preclinical / Clinical Development Services
	Millipore Immunoassay Services
	Millipore Target Screening & Profiling Services

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B3GNT4 Gene

protein-coding GIFtS: 50

Aliases &
Descriptions
for B3GNT4

Summaries
for B3GNT4

Genomic Location
for B3GNT4

Proteins
for B3GNT4

Protein Domains/
Families
for B3GNT4

Gene Function
for B3GNT4

Pathways & Interactions
for B3GNT4

Drugs & Compounds
for B3GNT4

Transcripts
for B3GNT4

Expression
for B3GNT4

Orthologs
for B3GNT4

Paralogs
for B3GNT4

SNPs/Variants
for B3GNT4

Disorders & Mutations
for B3GNT4

Medical News
for B3GNT4

Publications
for B3GNT4

Search for B3GNT4

Genome Databases showing B3GNT4

Other Databases showing B3GNT4

Specialized Databases showing B3GNT4

Licensable Technologies
for B3GNT4

Services
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Gene Index: 3 5 A B C D E F G H I J K L M N O P Q R S T U V W X Y Z

B3GNT4 Gene

protein-coding GIFtS: 50

Aliases & Descriptionsfor B3GNT4

Summariesfor B3GNT4

Genomic Locationfor B3GNT4

Proteinsfor B3GNT4

Protein Domains/Familiesfor B3GNT4

Gene Functionfor B3GNT4

Pathways & Interactionsfor B3GNT4

Drugs & Compoundsfor B3GNT4

Transcriptsfor B3GNT4

Expression for B3GNT4

Orthologsfor B3GNT4

Paralogsfor B3GNT4

SNPs/Variantsfor B3GNT4

Disorders & Mutationsfor B3GNT4

Medical Newsfor B3GNT4

Publicationsfor B3GNT4

Search for B3GNT4

Genome Databases showing B3GNT4

Other Databases showing B3GNT4

Specialized Databases showing B3GNT4

Licensable Technologiesfor B3GNT4

Servicesfor B3GNT4

Gene Index: 3 5 A B C D E F G H I J K L M N O P Q R S T U V W X Y Z

Aliases &
Descriptions
for B3GNT4

Summaries
for B3GNT4

Genomic Location
for B3GNT4

Proteins
for B3GNT4

Protein Domains/
Families
for B3GNT4

Gene Function
for B3GNT4

Pathways & Interactions
for B3GNT4

Drugs & Compounds
for B3GNT4

Transcripts
for B3GNT4

Expression
for B3GNT4

Orthologs
for B3GNT4

Paralogs
for B3GNT4

SNPs/Variants
for B3GNT4

Disorders & Mutations
for B3GNT4

Medical News
for B3GNT4

Publications
for B3GNT4

Licensable Technologies
for B3GNT4

Services
for B3GNT4